Bivalvia | ... | Unionoidea | Unionidae | Ambleminae | Quadrulini | Quadrulina | Quadrula

species Quadrula quadrula (Rafinesque, 1820): specimens

13 available nominal species synonyms (25 total synonyms)

Recent Taxonomic Opinions

Quadrula quadrula — Graf & Cummings (2007), Zhou et al. (2007), Tiemann et al. (2007), Williams et al. (2008), Chapman et al. (2008), ... Hou et al. (2025), Chen et al. (2025), Klymus et al. (2025), Campbell & Lydeard (2026), Barraza et al. (2026)
Quadrula apiculata — Graf & Cummings (2007), Williams et al. (2008), Bogan (2010), Cummings & Graf (2010), ITIS (2012), ... Marshall et al. (2018), Pfeiffer et al. (2019), Pfeiffer et al. (2019), Inoue et al. (2020), Hauck et al. (2023)
Quadrula rumphiana — Graf & Cummings (2007), Williams et al. (2008), Bogan (2010), Cummings & Graf (2010), IUCN (2012), ... Haag (2012), Williams et al. (2017), Inoue et al. (2018), Burlakova et al. (2019), Hauck et al. (2023)
Quadrula fragosa — Watters et al. (2009)
Quadrula quadrula quadrula — ITIS (2012)
“Quadrula quadrula” ** — Reis et al. (2013), Amaro et al. (2017), Gasparini et al. (2020)
“Quadrula quadrula” ** — Reis et al. (2013)
Q. quadrula — Sheng et al. (2014)
Quadrula quadrula rumphiana — Keogh et al. (2024)

Range

Widespread in the Interior, Nelson and Great Lakes basins, as well as the western Gulf Coastal Plain, North America. (North America: NA1 NA2 NA4)

Map of Georeferenced Specimens

specimens map of Quadrula quadrula

MUSSELp Specimen Records

Type Specimens

image of lot ANSP_20224
ANSP_20224
Salt River
neotype
Obliquaria quadrula Rafinesque, 1820

image of lot UMMZ_209203
UMMZ_209203
Fox River, nr. De Pere, Wisc.
type
Quadrula bullocki F.C. Baker, 1928

image of lot CM_61.6892
CM_61.6892
Lake Contrary, St. Joseph, Buchanan Co., Missouri
neotype
Quadrula contraryensis Utterback, 1915

image of lot USNM_84212
USNM_84212
Claiborne, Ala. R., Alabama
type
Unio asper Lea, 1831

image of lot USNM_84174
USNM_84174
Cincinnati, Ohio
paratype
Unio asperrimus Lea, 1831

image of lot USNM_84190
USNM_84190
Othcaloosa Cr., Ga.
figured type
Unio blandianus Lea, 1857

image of lot USNM_84169
USNM_84169
Texas
lectotype
Unio forsheyi Lea, 1859

image of lot USNM_84165
USNM_84165
Cincinnati, Ohio
paratype
Unio lacrymosus Lea, 1828

image of lot USNM_84189
USNM_84189
West Georgia
type
Unio rumphianus Lea, 1852

image of lot USNM_84172
USNM_84172
Leon Co., Texas
lectotype
Unio speciosus Lea, 1862

TYPE_LOCALITY_MP_016. Lake Ponchartrain, near N. Orleans — type locality Unio apiculatus Say, 1829

GenBank Vouchers

Georeferenced GenBank Vouchers

Mexico

FLMNH_438809_1 (SK20021). (COI: PQ038721, COI M-type: PQ142682, go to GenBank) Rio Grande (27.62609, -99.58903).

FLMNH_438809_1 (SK20030). (COI: PQ038730, go to GenBank) ditto.

FLMNH_438809_2 (SK20023). (COI: PQ038723, go to GenBank) ditto.

FLMNH_438809_3 (SK20024). (COI: PQ038724, COI M-type: PQ142684, go to GenBank) ditto.

FLMNH_438809_4 (SK20025). (COI: PQ038725, go to GenBank) ditto.

FLMNH_438809_5 (SK20026). (COI: PQ038726, go to GenBank) ditto.

FLMNH_438809_6 (SK20027). (COI: PQ038727, go to GenBank) ditto.

FLMNH_438809_7 (SK20022). (COI: PQ038722, COI M-type: PQ142683, go to GenBank) ditto.

FLMNH_438809_8 (SK20028). (COI: PQ038728, go to GenBank) ditto.

FLMNH_438809_9 (SK20029). (COI: PQ038729, COI M-type: PQ142685, go to GenBank) ditto.

USA

FLMNH_438891_1 (SK19058). (COI: PQ038654, COI M-type: PQ142668, ITS1, 5.8S: PQ143140, ND1: PQ057629, go to GenBank) Alabama River (31.6139, -87.55).

FLMNH_438891_1 (SK19068). (ITS1, 5.8S: PQ143149, ND1: PQ057639, go to GenBank) ditto.

FLMNH_438891_1 (SK19069). (COI: PQ038660, ITS1, 5.8S: PQ143150, ND1: PQ057640, go to GenBank) ditto.

FLMNH_438891_1 (SK19070). (ITS1: PQ143151, ND1: PQ057641, go to GenBank) ditto.

FLMNH_438891_2 (SK19059). (COI: PQ038655, ITS1, 5.8S: PQ143141, ND1: PQ057630, go to GenBank) ditto.

FLMNH_438891_3 (SK19060). (ITS1, 5.8S: PQ143142, ND1: PQ057631, go to GenBank) ditto.

FLMNH_438891_4 (SK19061). (ND1: PQ057632, go to GenBank) ditto.

FLMNH_438891_5 (SK19062). (ITS1, 5.8S: PQ143143, ND1: PQ057633, go to GenBank) ditto.

FLMNH_438891_6 (SK19063). (COI: PQ038656, ITS1, 5.8S: PQ143144, ND1: PQ057634, go to GenBank) ditto.

FLMNH_438891_7 (SK19064). (ITS1, 5.8S: PQ143145, ND1: PQ057635, go to GenBank) ditto.

FLMNH_438891_8 (SK19065). (COI: PQ038657, ITS1, 5.8S: PQ143146, ND1: PQ057636, go to GenBank) ditto.

FLMNH_438891_9 (SK19066). (COI: PQ038658, ITS1, 5.8S: PQ143147, ND1: PQ057637, go to GenBank) ditto.

Mp_4399_ (SK19067). (COI: PQ038659, ITS1, 5.8S: PQ143148, ND1: PQ057638, go to GenBank) ditto.

UA_1726 (QspeMob015). (COI: PQ038620, ND1: PQ057613, go to GenBank) Alabama River (32.31, -87.28).

JFBM_22857 (SK18144). (COI: PQ038624, COI M-type: PQ142658, ITS1: PQ143134, ND1: PQ057617, go to GenBank) Alabama River (32.392701, -87.004616).

ASUMZ_1234_1. (ND1: PQ057579, go to GenBank) Arkansas River (35.11874, -92.5532).

ASUMZ_1234_10. (ND1: PQ057580, go to GenBank) ditto.

ASUMZ_1234_2. (ND1: PQ057581, go to GenBank) ditto.

ASUMZ_1234_3. (COI: PQ038591, ITS1, 5.8S: PQ143111, ND1: PQ057582, go to GenBank) ditto.

ASUMZ_1234_4. (ND1: PQ057583, go to GenBank) ditto.

ASUMZ_1234_9. (ND1: PQ057584, go to GenBank) ditto.

ASUMZ_1460_7 (SK19054). (COI: PQ038652, ND1: PQ057628, go to GenBank) Bayou LaGrue (34.09942, -91.16562).

JFBM_23034_1 (SK19083). (COI: PQ038661, COI M-type: PQ142669, go to GenBank) Big Canoe Creek (33.848566, -86.250472).

JFBM_23034_2 (SK19084). (COI: PQ038662, go to GenBank) ditto.

JFBM_23034_3 (SK19085). (COI: PQ038663, COI M-type: PQ142670, go to GenBank) ditto.

FLMNH_438419_1 (SK19090). (COI: PQ038666, ITS1, 5.8S: PQ143154, ND1: PQ057643, go to GenBank) Black Warrior (32.555, -87.84409).

FLMNH_438419_2 (SK19091). (COI: PQ038667, ITS1, 5.8S: PQ143155, ND1: PQ057644, go to GenBank) ditto.

UAUC_1044. (ND1: AY158770, go to GenBank) Black Warrior R., Jefferson Co., Alabama.

JFBM_23111_1 (SK21001). (COI: PQ038734, ND1: PQ057679, go to GenBank) Black Warrior River (33.736553, -86.928699).

JFBM_23111_2 (SK21002). (COI: PQ038735, ND1: PQ057680, go to GenBank) ditto.

JFBM_23111_3 (SK21003). (COI: PQ038736, ND1: PQ057681, go to GenBank) ditto.

FLMNH_438826_1 (SK19086/20049). (COI: PQ038664, ITS1, 5.8S: PQ143152, ND1: PQ057642, go to GenBank) Bogue Chitto AL (32.31001, -87.27863).

FLMNH_438826_2 (SK19087/20050). (COI: PQ038665, ITS1, 5.8S: PQ143153, ND1: PQ217097, go to GenBank) ditto.

FLMNH_438826_3 (SK20002). (COI: PQ038716, go to GenBank) ditto.

JFBM_22858_1 (SK18155). (COI: PQ038625, COI M-type: PQ142659, ITS1, 5.8S: PQ143135, ND1: PQ057618, go to GenBank) Bogue Chitto AL (32.37014, -87.31969).

JFBM_22858_2 (SK18156). (COI: PQ038626, COI M-type: PQ142660, ND1: PQ057619, go to GenBank) ditto.

JFBM_22858_3 (SK18157). (COI: PQ038627, COI M-type: PQ142661, go to GenBank) ditto.

JFBM_22858_4 (SK18158). (COI: PQ038628, ITS1, 5.8S: PQ143136, go to GenBank) ditto.

FLMNH_439143 (QapiPrl126). (COI: PQ038601, ITS1, 5.8S: PQ143121, ND1: PQ057594, go to GenBank) Bogue Chitto LA (30.62254, -89.87655).

FLMNH_439143 (SK19099). (COI: PQ038675, ITS1, 5.8S: PQ143162, ND1: PQ057652, go to GenBank) ditto.

TXA&M_NRI_8427_1. (COI: PQ038586, ITS1: PQ143108, ND1: PQ057575, go to GenBank) Brazos River (29.785, -95.747).

TXA&M_NRI_8427_2. (COI: PQ038587, ITS1, 5.8S: PQ143109, ND1: PQ057576, go to GenBank) ditto.

TXA&M_NRI_8437_1. (COI: PQ038588, go to GenBank) Colorado River (30.90116, -99.91932).

TXA&M_NRI_8437_2. (COI: PQ038589, ND1: PQ057577, go to GenBank) ditto.

FLMNH_441183 (QapiCol065). (COI: PQ038593, ITS1, 5.8S: PQ143113, ND1: PQ057586, go to GenBank) Colorado River (31.4831003, -99.030595).

UAUC_331. (ND1: AY158777, go to GenBank) Coosawattee R., Gordon Co., Georgia.

Mp_2667_H1774. (COI: EF033268, COII: EF033288, go to GenBank) East Fork of the White River, downstream of US 50/SR 37 bridge, Lawrence County, Indiana.

Mp_2667_H1774g. (COI M-type: EF033308, COII M-type: EF033328, go to GenBank) ditto.

FLMNH_440990_2 (SK20007). (COI: PQ038717, ND1: PQ057673, go to GenBank) Galveston Bay (29.59798, -95.28663).

FLMNH_438818 (QapiGal016). (COI: PQ038594, ITS1, 5.8S: PQ143114, ND1: PQ057587, go to GenBank) Galveston Bay (29.789741, -95.623742).

FLMNH_438818 (QapiGal017). (COI: PQ038595, ITS1, 5.8S: PQ143115, ND1: PQ057588, go to GenBank) ditto.

JFBM_23178 (152). (COI: PQ038578, go to GenBank) Glover River (34.032063, -94.9393).

UMMZ_265699. (28S: DQ191417, COI: AF156511, go to GenBank) Grand River, MI.

JFBM_23040_1 (SK19109). (COI: PQ038682, go to GenBank) Iowa River (41.555722, -91.519682).

JFBM_23040_2 (SK19110). (COI: PQ038683, go to GenBank) ditto.

JFBM_23040_3 (SK19111). (COI: PQ038684, go to GenBank) ditto.

JFBM_23040_4 (SK19112). (COI: PQ038685, COI M-type: PQ142672, go to GenBank) ditto.

JFBM_23040_5 (SK19113). (COI: PQ038686, COI M-type: PQ142673, go to GenBank) ditto.

JFBM_23040_6 (SK19114). (COI: PQ038687, go to GenBank) ditto.

JFBM_23040_7 (SK19115). (COI: PQ038688, go to GenBank) ditto.

JFBM_23040_8 (SK19116). (COI: PQ038689, go to GenBank) ditto.

JFBM_23179_1 (178). (COI: PQ038579, go to GenBank) Kiamichi River (34.656876, -95.055331).

JFBM_23179_2 (179). (COI: PQ038580, go to GenBank) ditto.

ASUMZ_1230_7 (QapiRed139). (COI: PQ038602, ND1: PQ057595, go to GenBank) Little River (33.691899, -93.965148).

Mp_4399_ (SK21115). (COI: PQ038748, go to GenBank) Little River (33.936448, -94.829126).

Mp_4399_ (SK21119). (COI: PQ038749, go to GenBank) ditto.

Mp_4399_ (SK21120). (COI: PQ038750, go to GenBank) ditto.

Mp_4399_ (SK21123). (COI: PQ038751, go to GenBank) ditto.

Mp_4399_ (SK21125). (COI: PQ038752, go to GenBank) ditto.

Mp_4399_ (SK21126). (COI: PQ038753, go to GenBank) ditto.

Mp_4399_ (SK21127). (COI: PQ038754, go to GenBank) ditto.

TXA&M_NRI_8566_1 (SK19176). (COI: PQ038713, ND1: PQ057672, go to GenBank) Little River (33.9762, -94.9322).

TXA&M_NRI_8566_2 (SK19177). (COI: PQ038714, go to GenBank) ditto.

TXA&M_NRI_8566_3 (SK19178). (COI: PQ038715, go to GenBank) ditto.

JFBM_23044_1 (SK19121). (COI: PQ038694, COI M-type: PQ142677, go to GenBank) Mississippi River (42.056605, -90.14924).

JFBM_23044_2 (SK19122). (COI: PQ038695, ND1: PQ057660, go to GenBank) ditto.

JFBM_23044_3 (SK19123). (COI: PQ038696, COI M-type: PQ142678, ND1: PQ057661, go to GenBank) ditto.

JFBM_23044_4 (SK19124). (COI: PQ038697, ND1: PQ057662, go to GenBank) ditto.

JFBM_23044_5 (SK19125). (COI: PQ038698, COI M-type: PQ142679, ND1: PQ057663, go to GenBank) ditto.

JFBM_23042 (SK19118). (COI: PQ038691, ND1: PQ057659, go to GenBank) Mississippi River (43.603198, -91.230798).

JFBM_23041 (SK19117). (COI: PQ038690, COI M-type: PQ142674, ND1: PQ057658, go to GenBank) Mississippi River (43.634071, -91.241814).

JFBM_23043_1 (SK19119). (COI: PQ038692, COI M-type: PQ142675, go to GenBank) Mississippi River (43.664114, -91.235728).

JFBM_23043_2 (SK19120). (COI: PQ038693, COI M-type: PQ142676, go to GenBank) ditto.

JFBM_23045_1 (SK19126). (COI: PQ038699, ND1: PQ057664, go to GenBank) Mississippi River (45.030727, -93.282595).

JFBM_23045_2 (SK19127). (COI: PQ038700, ND1: PQ057665, go to GenBank) ditto.

JFBM_23045_3 (SK19128). (COI: PQ038701, COI M-type: PQ142680, ND1: PQ057666, go to GenBank) ditto.

UAUC_902. (ND1: AY158772, go to GenBank) Muskingum R., Washington Co., Ohio.

FLMNH_441004_1 (SK19092). (COI: PQ038668, ITS1, 5.8S: PQ143156, ND1: PQ057645, go to GenBank) Navidad River (28.91412, -96.56886).

FLMNH_441004_2 (SK19093). (COI: PQ038669, ITS1, 5.8S: PQ143157, ND1: PQ057646, go to GenBank) ditto.

FLMNH_438814 (QapiNav028). (COI: PQ038596, ITS1, 5.8S: PQ143116, ND1: PQ057589, go to GenBank) Navidad River (28.95756, -96.54243).

FLMNH_438814 (QapiNav029). (COI: PQ038597, ITS1, 5.8S: PQ143117, ND1: PQ057590, go to GenBank) ditto.

UAUC_2620. (ND1: AY158805, go to GenBank) Neches R., Tyler Co., Texas.

FLMNH_439196 (SK20035). (COI: PQ038731, ITS1, 5.8S: PQ143167, ND1: PQ057676, go to GenBank) Neches River (30.659012, -94.170894).

FLMNH_438804_1 (SK19097). (5.8S, ITS1: PQ143161, COI: PQ038673, ND1: PQ057650, go to GenBank) Nueces River (28.337458, -98.086636).

FLMNH_438801_1 (SK19096). (5.8S, ITS1: PQ143160, COI: PQ038672, ND1: PQ057649, go to GenBank) Nueces River (28.382964, -98.109617).

FLMNH_438561_1 (SK19094). (5.8S, ITS1: PQ143158, COI: PQ038670, COI M-type: PQ142671, ND1: PQ057647, go to GenBank) Nueces River (28.47405, -98.25166).

FLMNH_438561_2 (SK19095). (5.8S, ITS1: PQ143159, COI: PQ038671, ND1: PQ057648, go to GenBank) ditto.

FLMNH_438561_3 (QapiNue088). (5.8S, ITS1: PQ143118, COI: PQ038598, COI M-type: PQ142656, ND1: PQ057591, go to GenBank) ditto.

FLMNH_438561_4 (QapiNue089). (5.8S, ITS1: PQ143119, COI: PQ038599, ND1: PQ057592, go to GenBank) ditto.

FLMNH_438561_5 (QapiNue090). (COI: PQ038600, COI M-type: PQ142657, ITS1: PQ143120, ND1: PQ057593, go to GenBank) ditto.

UAUC_1695. (ND1: AY158774, go to GenBank) Ohio R.,Vanderburgh Co., Indiana.

FLMNH_439156 (QquaOhi016). (COI: PQ038614, ND1: PQ057607, go to GenBank) Ohio River (39.39123122, -81.42094424).

FLMNH_439156 (SK19100). (COI: PQ038676, ITS1, 5.8S: PQ143163, ND1: PQ057653, go to GenBank) ditto.

FLMNH_439156 (SK19101). (COI: PQ038677, ITS1, 5.8S: PQ143164, ND1: PQ057654, go to GenBank) ditto.

FLMNH_439156 (SK19102). (COI: PQ038678, ITS1, 5.8S: PQ143165, ND1: PQ057655, go to GenBank) ditto.

UAUC_435. (ND1: AY158776, go to GenBank) Oostanaula R., Gordon Co., Georgia.

JFBM_22865_1 (SK18179). (COI: PQ038637, COI M-type: PQ142664, ND1: PQ057623, go to GenBank) Osage River (38.231031, -92.558275).

JFBM_22865_2 (SK18180). (COI: PQ038638, ND1: PQ057624, go to GenBank) ditto.

JFBM_22865_3 (SK18181). (COI: PQ038639, ND1: PQ057625, go to GenBank) ditto.

JFBM_22865_4 (SK18182). (COI: PQ038640, COI M-type: PQ142665, ITS1, 5.8S: PQ143139, go to GenBank) ditto.

JFBM_22865_5 (SK18183). (COI: PQ038641, COI M-type: PQ142666, go to GenBank) ditto.

JFBM_22865_6 (SK18184). (COI: PQ038642, COI M-type: PQ142667, go to GenBank) ditto.

ASUMZ_QF5 (QfraOua021). (COI: PQ038613, ND1: PQ057606, go to GenBank) Ouachita River (32.95613, -92.07864).

ASUMZ_QF1 (QfraOua017). (COI: PQ038611, ND1: PQ057604, go to GenBank) Ouachita River (33.03438, -92.09618).

FLMNH_438884_1 (SK20036). (COI M-type: PQ142686, go to GenBank) Pascagoula River (30.63228, -88.652401).

FLMNH_438884_3 (SK20038). (COI: PQ038732, ITS1, 5.8S: PQ143168, ND1: PQ057677, go to GenBank) ditto.

FLMNH_438884_4 (SK20039). (COI: PQ038733, ITS1, 5.8S: PQ143169, ND1: PQ057678, go to GenBank) ditto.

UAUC_1045. (ND1: AY158790, go to GenBank) Red R., Powell Co., Kentucky.

TXA&M_NRI_8602_1 (SK19168). (COI: PQ038707, go to GenBank) Red River (33.654324, -96.048929).

TXA&M_NRI_8602_2 (SK19169). (COI: PQ038708, go to GenBank) ditto.

TXA&M_NRI_8602_3 (SK19170). (COI: PQ038709, go to GenBank) ditto.

TXA&M_NRI_8610_1 (SK19171). (COI: PQ038710, go to GenBank) Red River (33.65766, -96.05555).

TXA&M_NRI_8610_2 (SK19172). (COI: PQ038711, go to GenBank) ditto.

TXA&M_NRI_8584_1 (SK19165). (COI: PQ038704, go to GenBank) Red River (33.686069, -94.994325).

TXA&M_NRI_8584_2 (SK19166). (COI: PQ038705, ND1: PQ057668, go to GenBank) ditto.

TXA&M_NRI_8584_3 (SK19167). (COI: PQ038706, go to GenBank) ditto.

TXA&M_NRI_8656_1 (SK19173). (COI: PQ038712, ND1: PQ057669, go to GenBank) Red River (33.826894, -95.412156).

TXA&M_NRI_8656_2 (SK19174). (ND1: PQ057670, go to GenBank) ditto.

TXA&M_NRI_8656_3 (SK19175). (ND1: PQ057671, go to GenBank) ditto.

TXA&M_NRI_8122_1. (5.8S, ITS1: PQ143106, COI: PQ038581, ND1: PQ057572, go to GenBank) Red River (33.82702, -95.41176).

TXA&M_NRI_8122_2. (COI: PQ038582, go to GenBank) ditto.

FLMNH_438787 (QquaRed002). (5.8S, ITS1: PQ143130, COI: PQ038615, ND1: PQ057608, go to GenBank) Red River (33.854656, -95.341305).

TXA&M_NRI_8382_1. (COI: PQ038584, ND1: PQ057574, go to GenBank) Rio Grande (27.602238, -99.580507).

TXA&M_NRI_8382_2. (COI: PQ038585, go to GenBank) ditto.

FLMNH_438554_2 (QapiRGr081). (5.8S, ITS1: PQ143122, COI: PQ038603, ND1: PQ057596, go to GenBank) Rio Grande (27.63774, -99.62695).

FLMNH_438554_3 (QapiRGr082). (5.8S, ITS1: PQ143123, COI: PQ038604, ND1: PQ057597, go to GenBank) ditto.

FLMNH_438554_4 (QapiRGr083). (5.8S, ITS1: PQ143124, COI: PQ038605, ND1: PQ057598, go to GenBank) ditto.

FLMNH_438747 (QapiRGr092). (5.8S, ITS1: PQ143125, COI: PQ038606, ND1: PQ057599, go to GenBank) Rio Grande (27.64193, -99.60222).

FLMNH_441088 (QapiCol044). (COI: PQ038592, ITS1, 5.8S: PQ143112, ND1: PQ057585, go to GenBank) San Saba River (31.20738, -98.68761).

UAUC_722. (ND1: AY158775, go to GenBank) Sipsey R., Pickens Co., Alabama.

FLMNH_439169 (QrumTom005). (COI: PQ038617, ITS1, 5.8S: PQ143131, ND1: PQ057610, go to GenBank) Sipsey River (33.040556, -88.113611).

FLMNH_439169 (QrumTom008). (COI: PQ038618, ITS1, 5.8S: PQ143132, ND1: PQ057611, go to GenBank) ditto.

FLMNH_439169 (QrumTom011). (COI: PQ038619, ITS1, 5.8S: PQ143133, ND1: PQ057612, go to GenBank) ditto.

UA_1489 (QrumMob003). (COI: PQ038616, ND1: PQ057609, go to GenBank) Sipsey River (33.1025, -87.95).

JFBM_22862_1 (SK18164). (COI: PQ038629, ND1: PQ057620, go to GenBank) Sipsey River (33.331581, -87.761735).

JFBM_22862_2 (SK18165). (COI: PQ038630, COI M-type: PQ142662, ND1: PQ057621, go to GenBank) ditto.

JFBM_22862_3 (SK18166). (COI: PQ038631, COI M-type: PQ142663, go to GenBank) ditto.

JFBM_22862_4 (SK18167). (COI: PQ038632, ITS1: PQ143137, go to GenBank) ditto.

JFBM_22862_5 (SK18168). (COI: PQ038633, ITS1, 5.8S: PQ143138, go to GenBank) ditto.

UAUC_1698. (ND1: AY158773, go to GenBank) Spring R., Cherokee Co., Kansas.

Mp_4399_ (SK21111). (COI: PQ038746, go to GenBank) St. Croix River (45.395003, -92.664611).

Mp_4399_ (SK21112). (COI: PQ038747, go to GenBank) ditto.

FLMNH_438313_1 (SK20010/041). (5.8S, ITS1: PQ143166, COI: PQ038718, ND1: PQ057674, go to GenBank) Trinity River (30.627278, -95.008917).

FLMNH_438313_6 (SK20015/046). (COI: PQ038719, COI M-type: PQ142681, go to GenBank) ditto.

TXA&M_NRI_8470_2. (5.8S, ITS1: PQ143110, COI: PQ038590, ND1: PQ057578, go to GenBank) Trinity River (30.88456, -95.77792).

TXA&M_NRI_8262. (5.8S, ITS1: PQ143107, COI: PQ038583, ND1: PQ057573, go to GenBank) Trinity River (31.6632089, -95.79061759).

FLMNH_441182_2 (QapiTri064). (5.8S, ITS1: PQ143127, COI: PQ038608, ND1: PQ057601, go to GenBank) Trinity River (32.592543, -96.486377).

FLMNH_438952_2 (QapiTri113). (5.8S, ITS1: PQ143129, COI: PQ038610, ND1: PQ057603, go to GenBank) Trinity River (32.980521, -97.09841).

FLMNH_438951_1 (QapiTri107). (5.8S, ITS1: PQ143128, COI: PQ038609, ND1: PQ057602, go to GenBank) Trinity River (33.036203, -97.1759).

TXA&M_NRI_8378_2 (SK19163). (COI: PQ038702, go to GenBank) Trinity River (33.305606, -96.607848).

TXA&M_NRI_8378_3 (SK19164). (COI: PQ038703, ND1: PQ057667, go to GenBank) ditto.

UF_439156 / UA_3563. (28S: MK036133, COI: MH633643, ITS1, 5.8S: MH362613, ND1: MH633595, go to GenBank) USA, Ohio, Wood County, Ohio River; Buckley Island backchannel (39.3912312, -81.4209442).

UF_441088. (16S: MK001791, 28S: MK001825, COI: KT285648, ITS1, 5.8S: KT285692, SRA: SRX5278551, go to GenBank) USA, Texas, San Saba County, San Saba River at CR 104, San Saba River Golf Course, near San Saba, TX (31.20738, -98.68761).

UF_438554 (QapiRGr081). (COI: MH633638, ITS1, 5.8S: MH362562, ND1: MH633590, go to GenBank) USA, Texas, Webb County, Rio Grande (site 2), about 300 m upstream of Santa Isabel Creek (27.63774, -99.62695).

ASUMZ_QF4 (QfraOua020). (COI: PQ038680, ND1: PQ057657, go to GenBank) USA: Ouachita River (32.95613 N 92.07864 W).

UA_3609 (QquaOhi010). (ND1: PQ057651, go to GenBank) USA: Red River KY (37.84167 N 83.76 W).

GenBank Vouchers Not Yet Georeferenced

ANSP_397246. (28S: U82340, go to GenBank)

ASUMZ_QF3 (QfraOua019). (COI: PQ038612, ND1: PQ057605, go to GenBank)

BM_19581. (ND1: HM852936, go to GenBank)

FLMNH_441182_1 (QapiTri063). (5.8S, ITS1: PQ143126, COI: PQ038607, ND1: PQ057600, go to GenBank)

JBFMC_10006. (transcriptome shotgun assembly: GKMJ00000000, go to GenBank)

JBFMC_10006_5. (COI: OK562100, go to GenBank)

Mp_3901_BIV1923. (COI: MK503267, go to GenBank)

UA_20703. (COI: HM230409, go to GenBank)

UA20703. (ND1: HM230421, go to GenBank)