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Data last updated
Mon 1 Dec 2025

Bivalvia | ... | Unionidae | Ambleminae | Quadrulini | Quadrulina | Quadrula quadrula

nominal species Unio rugosus Barnes, 1823

= Quadrula quadrula

Taxonomic Opinions

+ Quadrula quadrula (Rafinesque, 1820) — Lopes-Lima et al., 2019

+ Quadrula fragosa (Conrad, 1835) — Watters et al., 2009

+ Quadrula quadrula (Rafinesque, 1820) — Williams et al., 2008

+ Quadrula quadrula (Rafinesque, 1820) — Parmalee & Bogan, 1998

+ Quadrula quadrula (Rafinesque, 1820) — R.I. Johnson, 1980

+ Quadrula quadrula (Rafinesque, 1820) — Burch, 1975

+ Quadrula (s.s.) quadrula quadrula (Rafinesque, 1820) — Haas, 1969

+ Quadrula (s.s.) gigantea [sic] (Raf.) — Haas, 1930

+ Quadrula (s.s.) quadrula (Rafinesque, 1820) — F.C. Baker, 1928

+ Quadrula (s.s.) quadrula (Rafinesque, 1820) — Frierson, 1927

+ Quadrula quadrula (Rafinesque, 1820) — Ortmann & Walker, 1922

+ — Simpson, 1914

+ Unio metanevrus Rafinesque — Call, 1895

+ Margaron (Unio) metanevrus (Raf.) — Lea, 1870

+ Unio metanever (Rafinesque, 1820) — Reeve, 1864

+ Unio metaneurus [sic] Raf. — Conrad, 1853

= Quadrula rugosus — Agassiz, 1852

+ Margaron (Unio) metanevrus (Raf.) — Lea, 1852

+ Unio metanever [sic] Lea — Hanley, 1843

+ Unio quadrulus (Rafinesque, 1820) — Conrad, 1840

= Unio (Theliderma) rugosus — Swainson, 1840

+ Margarita (Unio) metanever Lea — Lea, 1838

+ Margarita (Unio) metanever (Rafinesque, 1820) — Lea, 1836

+ Unio quadrulus Raf. — Ferussac, 1835

+ Unio metanever [sic] Rafin. — Lea, 1829

= Unio rugosus — Hildreth, 1828

= Unio rugosus — Barnes, 1823