Bivalvia | ... | Unionidae | Ambleminae | Quadrulini | Quadrulina | Quadrula fragosa

nominal species Unio fragosus Conrad, 1835

Unio fragosus Conrad, 1835 = Quadrula fragosa

Conrad, 1835. Monogr. Fam. Unionidae 1(1): 12, pl. 6, fig. 2.
Conrad, 1853. Proc. Acad. Nat. Sci. 6: 249.

Taxonomic Opinions

= Quadrula fragosa (Conrad, 1836) — Campbell & Lydeard, 2026: 289.

= Quadrula fragosa (Conrad, 1835) — FMCS, 2025: [6].

= Quadrula fragosa — Keogh et al., 2024: 5, Table S1.

= Quadrula fragosa (Conrad, 1835) — Neemuchwala et al., 2023: 2, Table 1.

= Quadrula fragosa (Conrad, 1835) — Hopper et al., 2023.

= Quadrula fragosa (Conrad, 1835) — Graf & Cummings, 2021: 20.

= Quadrula fragosa (Conrad, 1835) — FMCS, 2021: [6].

= Quadrula fragosa (Conrad, 1835) — Lopes-Lima et al., 2019: [18], Table 7.

= Quadrula fragosa (Conrad, 1835) — Williams et al., 2017: 43.

= Quadrula fragosa (Conrad, 1836) — Roe & Boyer, 2015: 52.

= Quadrula fragosa (Conrad, 1835) — Paterson et al., 2015: 1124.

= Quadrula fragosa (Conrad, 1835) — ITIS, 2012: [web].

= Quadrula fragosa — Haag, 2012: 69.

= Quadrula fragosa (Conrad, 1835) — McMurray et al., 2012: 75.

= Quadrula fragosa Conrad, 1836 — Bogan, 2010: [web].

= Quadrula fragosa — Cummings & Graf, 2010: 376.

= Quadrula fragosa — Almany et al., 2009: 1464.

= Quadrula fragosa (Conrad, 1835) — Watters et al., 2009: 269, figs. 177-178.

= Quadrula fragosa (Conrad, 1836) — Graf & Cummings, 2007: 308.

= Quadrula fragosa (Conrad, 1835) — Tiemann et al., 2007: 120.

= Quadrula fragosa (Conrad, 1835) — Cicerello & Schuster, 2003: 46, text fig..

= Quadrula fragosa — Sietman, 2003: 52, text fig..

= Quadrula fragosa (Conrad) — Lydeard et al., 1999: 130.

= Quadrula fragosa (Conrad, 1836) — Parmalee & Bogan, 1998: 212, pl. 103.

= Quadrula fragosa (Conrad, 1836) — Turgeon et al., 1998: 37.

= Quadrula fragosa (Conrad, 1835) — Cummings & Mayer, 1997: 137.

+ Quadrula quadrula (Rafinesque, 1820) — Howells et al., 1996: 124.

= Quadrula fragosa (Conrad, 1835) — IUCN, 1996: [web].

= Quadrula fragosa (Conrad) — Williams et al., 1993: 14.

= Quadrula fragosa (Conrad, 1835) — Cummings & Mayer, 1992: 28, text fig..

= Quadrula fragosa (Conrad, 1836) — Turgeon et al., 1988: 32.

+ Quadrula quadrula (Rafinesque, 1820) — R.I. Johnson, 1980: 90.

+ Quadrula quadrula (Rafinesque, 1820) — Burch, 1975: 177.

= Quadrula (s.s.) quadrula fragosa (Conrad, 1836) — Haas, 1969: 290.

= Quadrula (s.s.) fragosus (Conrad, 1836) — F.C. Baker, 1928: 88, pl. 46, figs. 5-6; pl. 47, fig. 1; pl. 48, fig. 1.

= Quadrula (s.s.) quadrula-fragosa (Conrad, 1836) — Frierson, 1927: 47, sp. 207a.

= Quadrula quadrula fragosa (Conrad) — Ortmann, 1925: 335.

= Quadrula fragosa (Conrad) — Ortmann, 1924: 16.

= Quadrula fragosa (Conrad) — Utterback, 1915: 140, pl. 18, figs. 48a-b.

= Quadrula (sec. Theliderma) fragosa (Conrad, 1836) — Simpson, 1914: 843.

= Quadrula (sec. Theliderma) fragosus (Conrad, 1836) — Simpson, 1900: 777.

= Unio (s.s.) fragosus Conrad, 1836 — Call, 1900: 490.

= Unio fragosus Con. — B.H. Wright, 1888: [3].

= Margaron (Unio) fragosus (Con.) — Lea, 1870: 33.

= Unio fragosus Conrad, 1836 — Reeve, 1864: pl. 1, sp. 2; pl. 7, sp. 27.

= Unio fragosus Conrad — Küster, 1861: 173, pl. 55, fig. 1.

= Unio (Quadrula) fragosus Conr. — Adams & Adams, 1857: 497.

= Unio (Theliderma) fragosus Con. — Conrad, 1853: 249, 269.

= Quadrula fragosus — Agassiz, 1852: 48.

= Margaron (Unio) fragosus (Con.) — Lea, 1852: 22.

= Unio tragosus [sic] Conr. — Hanley, 1843: 178, pl. 20, fig. 40.

= Margarita (Unio) fragosus Con. — Lea, 1838: 15.

= Margarita (Unio) fragrosus [sic] (Conrad, 1836) — Lea, 1836: 14.

= Unio fragosus Conrad, 1835 — Conrad, 1835: 12, pl. 6, fig.2.