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species Quadrula fragosa (Conrad, 1836): synonymy

1 specimen + 18 Genbank vouchers

Synonymy

Unio fragosus Conrad, 1835

Conrad, 1835. Monogr. Fam. Unionidae 1(1): 12, pl. 6, fig. 2.
Conrad, 1853. Proc. Acad. Nat. Sci. 6: 249.

Quadrula (sec. Theliderma) fragosus — Simpson (1900)
Unio (s.s.) fragosus — Call (1900)
Quadrula (sec. Theliderma) fragosa — Simpson (1914)
Quadrula fragosa — Utterback (1915), Ortmann (1924), Turgeon et al. (1988), Cummings & Mayer (1992), Williams et al. (1993), ... FMCS (2021), Neemuchwala et al. (2023), Hopper et al. (2023), Keogh et al. (2024), FMCS (2025)
Quadrula quadrula fragosa — Ortmann (1925)
Quadrula (s.s.) quadrula-fragosa — Frierson (1927)
Quadrula (s.s.) fragosus — F.C. Baker (1928)
Quadrula (s.s.) quadrula fragosa — Haas (1969)
Quadrula quadrula — Burch (1975), R.I. Johnson (1980), Howells et al. (1996)

Margarita (Unio) fragrosus Lea, 1836

‘Con.’ Lea, 1836. Synopsis Fam. Naiades: 14 [corrected in the errata].
(error for Unio fragosus Conrad, 1835)

Quadrula fragosa — Watters et al. (2009)

Unio tragosus Hanley, 1843

‘Conrad’ Hanley, 1843. Illustr. Descr. Cat. Rec. Bivalve Shells: 178, pl. 20, fig. 40.
(error for Unio fragosus Conrad, 1835)

Quadrula (sec. Theliderma) fragosus — Simpson (1900)
Quadrula (sec. Theliderma) fragosa — Simpson (1914)
Quadrula fragosa — Watters et al. (2009)

Quadrula fragosa Scammon, 1906

‘Conrad’ Scammon, 1906. KS Univ. Sci. Bull. 3(9): ??.
(reference to Unio fragosus Conrad, 1835)

Quadrula quadrula — Murray & Leonard (1962), Vidrine (1993)

Taxonomic History (since 1900)

Quadrula (sec. Theliderma) fragosus — Simpson (1900)
Unio (s.s.) fragosus — Call (1900)
Quadrula (sec. Theliderma) fragosa — Simpson (1914)
Quadrula fragosa — Utterback (1915), Ortmann (1924), Turgeon et al. (1988), Cummings & Mayer (1992), Williams et al. (1993), ... FMCS (2021), Neemuchwala et al. (2023), Hopper et al. (2023), Keogh et al. (2024), FMCS (2025)
Quadrula quadrula fragosa — Ortmann (1925)
Quadrula (s.s.) quadrula-fragosa — Frierson (1927)
Quadrula (s.s.) fragosus — F.C. Baker (1928)
Quadrula quadrula — Murray & Leonard (1962), Burch (1975), R.I. Johnson (1980), Vidrine (1993), Howells et al. (1996)
Quadrula (s.s.) quadrula fragosa — Haas (1969)