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Thu 18 Dec 2025

Bivalvia | ... | Unionidae | Ambleminae | Quadrulini | Quadrulina | Quadrula couchiana

nominal species Unio couchianus Lea, 1860

Unio couchianus Lea, 1860 = Quadrula couchiana

Lea, 1860. Proc. Acad. Nat. Sci. 12: 305.
Lea, 1860. Jour. Acad. Nat. Sci. 4 [O. 8]: 371 [53], pl. 66, fig. 196.
Type(s): LECTOTYPE [AS HOLOTYPE] USNM_85728 (by R.I. Johnson, 1974: 38), Rio Salado, Mexico.

= Quadrula couchiana

Taxonomic Opinions

= Quadrula couchiana (Lea, 1860) — FMCS, 2025

= Quadrula couchiana (Lea, 1860) — Cummings et al., 2025

= Quadrula couchiana (I. Lea, 1860) — Czaja et al., 2023

= Quadrula couchiana (Lea, 1860) — Neemuchwala et al., 2023

= Quadrula couchiana (Lea, 1860) — Hopper et al., 2023

= Quadrula couchiana (Lea, 1860) — Graf & Cummings, 2021

= Quadrula couchiana (Lea, 1860) — FMCS, 2021

= Quadrula couchiana — Tiemann et al., 2020

= Quadrula couchiana (Lea, 1860) — Lopes-Lima et al., 2019

= Quadrula couchiana (Lea, 1860) — Williams et al., 2017

= Quadrula couchiana (I. Lea, 1860) — ITIS, 2012

= Quadrula couchiana — Haag, 2012

= Quadrula couchiana Lea, 1860 — Bogan, 2010

= Amphinaias couchiana — Cummings & Graf, 2010

= Amphinaias couchiana (Lea, 1860) — Graf & Cummings, 2007

= Quadrula counchiana [sic] Lea, 1860 — Contreras-Arquieta, 2000

= Quadrula couchiana (Lea, 1860) — R.I. Johnson, 1999

= Quadrula couchiana (Lea, 1860) — Turgeon et al., 1998

= Quadrula couchiana (Lea, 1860) — Howells et al., 1996

= Amphinaias couchiana (Lea, 1860) — IUCN, 1996

= Quadrula couchiana (I. Lea) — Williams et al., 1993

= Quadrula couchiana (Lea, 1860) — Turgeon et al., 1988

= Quadrula (Pustulosa) couchiana (Lea, 1860) — Haas, 1969

= Quadrula (Amphinaias) couchianus (Lea, 1860) — Haas, 1969

= Quadrula (sec. Amphinaias) couchiana (Lea) — Thiele, 1934

= Quadrula (Bullata) couchiana (Lea, 1860) — Frierson, 1927

= Quadrula (sec. Theliderma) couchiana (Lea, 1860) — Simpson, 1914

= Quadrula (sec. Theliderma) couchiana (Lea, 1860) — Simpson, 1900

= Unio (Pleurobema) couchianus Lea, 1860 — Martens, 1900

= Unio (Amphinaias) couchianus I. Lea, 1860 — Fischer & Crosse, 1894

= Unio couchianus Lea — B.H. Wright, 1888

= Margaron (Unio) couchianus (Lea) — Lea, 1870

= Unio couchianus (Lea, 1860) — Sowerby, 1868