Khalloufi, N., C. Toledo, A. Machordom, M. Boumaïza & R. Araujo. 2011. The unionids of Tunisia: taxonomy and phylogenetic relationships, with redescription of Unio ravoisieri Deshayes, 1847 and U. durieui Deshayes, 1847. Journal of Molluscan Studies 77(2): 103-115. https://doi.org/10.1093/mollus/eyq046.
(click the figure number to see the cladogram)
+----------------- Amblema_plicata
|
--+ +-------------- Potomida_littoralis
| |
| | +----- Unio_tumidus
+--+ +-----+
| | | +-- Unio_gibbus
| | +--+
| | +-- Unio_durieui
+--+
| +-- Unio_tumidiformis
| +-----+
| | +-- Unio_crassus
+--+
| +----- Unio
| |
+--+----- Unio_pictorum
|
| +-- Unio_delphinus
+--+
+-- Unio_mancus