MUSSELpdb
Taxonomy
Species & Genus Richness of
freshwater mussels
by Taxonomy
Species & Genus Richness of
freshwater bivalves
by Family
Geography
Species Richness & Endemism of
freshwater mussels
by Geography
Data last updated
Mon 2 March 2026
Bivalvia | Unionida | Unionoidea | Margaritiferidae | Gibbosulinae | Gibbosula laosensis
nominal species Unio laosensis Lea, 1863
Unio laosensis Lea, 1863 = Gibbosula laosensis
Lea, 1863. Proc. Acad. Nat. Sci. 15: 190.
Lea, 1866. Jour. Acad. Nat. Sci. 6 [O. 11]: 63, pl. 21, fig. 61.
Type(s): HOLOTYPE USNM_86160, Laos Mts., Cambodia.
Taxonomic Opinions
= Gibbosula laosensis (Lea, 1863) — Dai et al., 2026: Table 1.
= Gibbosula laosensis (Lea, 1863) — Campbell & Lydeard, 2026: 287.
= Gibbosula laosensis — Dai et al., 2025: Table 1.
= Gibbosula laosensis (Lea, 1863) — Chen et al., 2025: Table 1.
= Gibbosula laosensis — Chen et al., 2025: Table 1.
= Gibbosula laosensis (Lea, 1863) — Dai et al., 2024: 1194, Table 1.
= Gibbosula laosensis — Dai et al., 2024: Table 2.
= Gibbosula laosensis — Dai et al., 2024: Table 2.
= Gibbosula laosensis (Lea, 1863) — Konopleva et al., 2023: Table S1.
= Gibbosula laosensis — Dai et al., 2023: 156, Table 1.
= Gibbosula laosensis (Lea, 1863) — Bolotov et al., 2022: Table S1.
= Gibbosula laosensis (Lea, 1863) — Jeratthitikul et al., 2022: Table S1.
= Gibbosula laosensis (Lea, 1863) — Graf & Cummings, 2021: 26.
= Gibbosula laosensis (Lea, 1863) — Wu et al., 2021: attached Table 1.
= Gibbosula laosensis (Lea, 1863) — Bolotov et al., 2020: Table S1.
= Gibbosula laosensis — Wu et al., 2020: 195.
= Margaritifera laosensis — Sano et al., 2020: 99.
= Margaritifera laosensis (Lea, 1863) — Huang et al., 2019: Table S4.
= Gibbosula laosensis (Lea, 1863) — Konopleva et al., 2019: Table S1.
= Gibbosula laosensis (Lea, 1863) — Bolotov et al., 2019: 23.
= Gibbosula laosensis — Lyubas et al., 2019: 14.
= Gibbosula laosensis (Lea, 1863) — Bolotov et al., 2019: Table S1.
= Gibbosula laosensis (Lea, 1863) — Bolotov et al., 2019: 443.
= Gibbosula laosensis — Vikhrev et al., 2019: 7, fig. 5, Table 3.
= Margaritifera laosensis (Lea, 1863) — Zieritz et al., 2018: online Table 1.
= Margaritifera laosensis — Vikhrev et al., 2018: 73, appendix 5.
= Margaritifera laosensis (Lea, 1863) — Do et al., 2018: 5, Table 2.
= Margaritifera laosensis (Lea, 1863) — Bolotov et al., 2018: Table S1.
= Margaritifera laosensis — Huang et al., 2018: 302.
= Gibbosula laosensis (Lea, 1863) — Lopes-Lima et al., 2018: 103, Table 3, S1, sp. 3.
= Margaritifera laosensis (Lea, 1863) — Bespalaya et al., 2018: 97.
= Margaritifera laosensis (Lea, 1863) — Bolotov et al., 2017: [16], Table S1.
= Margaritifera laosensis (Lea, 1863) — Bolotov et al., 2017: Table S1.
= Margaritifera laosensis (Lea, 1863) — Vikhrev et al., 2017: 5.
= Margaritifera laosensis — Bolotov et al., 2016: 59.
= Margaritifera (Margaritanopsis) laosensis — Bolotov et al., 2016: 107, 111.
= Margaritifera laosensis (Lea, 1863) — Araujo et al., 2016: 290.
= Margaritifera laosensis (Lea, 1863) — Bolotov et al., 2014: 706, figs. 2-3.
= Margaritifera laosensis (Lea, 1863) — IUCN, 2011: [web].
= Margaritifera laosensis Lea, 1863 — Bogan, 2010: [20].
= Margaritifera laosensis (Lea, 1863) — Araujo et al., 2009: 95.
= Margaritifera laosensis (Lea, 1863) — Graf & Cummings, 2007: 312.
= Margaritanopsis laosensis — Kittivorachate & Yangyuen, 2004: 134.
= Margaritanopsis laosensis (Lea, 1863) — D.G. Smith, 2001: 41, 42.
= Margaritifera laosensis (Lea, 1863) — Ziuganov et al., 1994: 60.
= Margaritifera (Margaritanopsis) laosensis (Lea, 1863) — Subba Rao, 1989: 199, sp. 240, figs. 546-547.
= Margaritanopsis laosensis (Lea, 1863) — Brandt, 1974: 261.
= Margaritifera (Margaritanopsis) laosensis (Lea, 1863) — Haas, 1969: 13.
= Margaritifera (Margaritanopsis) laosensis (Lea, 1863) — Haas, 1969: N415, fig. D16:1.
= Margaritanopsis laosensis (Lea, 1863) — Prashad, 1922: 93, pl. 2, figs. 1-4.
= Margaritana laosensis (Lea, 1863) — Simpson, 1914: 520.
+ — de Gregorio, 1914: 48.
= Margaritanopsis laosensis (Lea, 1863) — Haas, 1912: 122, pl. 12, figs. 1-2.
= Margaritana laosensis (Lea, 1863) — Simpson, 1900: 678.
= Unio laosensis Lea — Fischer, 1891: 226.
= Margaron (Unio) laosensis (Lea) — Lea, 1870: 62.
= Unio laosensis Lea, 1863 — Lea, 1866: 67, pl. 21, fig. 61.
