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The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!
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Bivalvia | Unionoida | Unionoidea | Unionidae | Ambleminae | Popenaiadini | Psoronaias species Psoronaias semigranosa (von dem Busch in Philippi, 1843): synonymy 149 specimens + 3 Genbank vouchers Unio semigranosus Philippi, 1843
von dem Busch in Philippi, 1843. Abbild. Beschr. Conch. 1: 19, pl. 1, figs. 1-3. Type(s): PARATYPES MCZ_152861, near Tampico, Mexico.
Unio (sec. Elliptio) semigranosus — Simpson (1900), Simpson (1914) Unio (Psoronaias) semigranosus — Martens (1900) Quadrula semigranosa — Pilsbry (1910) Psoronaias semigranosus — Frierson (1917), Frierson (1927), Haas (1930), Goodrich & van der Schalie (1937) Psoronaias semigranosa — Haas (1929), Graf & Cummings (2007), Bogan (2010), Pfeiffer et al. (2019), Pfeiffer et al. (2019), Inoue et al. (2020), Smith et al. (2020), Graf & Cummings (2021), Czaja et al. (2023), Freeman et al. (2025) Psoronaias semigranosa semigranosa — Haas (1969) | |
Unio crocodilorum Morelet, 1849
Morelet, 1849. Test. Nov. 1: 28. Type(s): SYNTYPE BMNH_1893-2-4-2030, River Usumacinta, Central America.
Unio (sec. Elliptio) crocodilarum [sic] — Simpson (1900), Simpson (1914) Unio (Psoronaias) crocodilorum — Martens (1900) Psoronaias crocodilorum — Frierson (1917), Frierson (1927), Haas (1930), Haas (1969), Graf & Cummings (2007), Bogan (2010) Psoronaias crocodilarum [sic] — Haas (1929) Psoronaias semigranosus — Goodrich & van der Schalie (1937) Psoronaias semigranosa — Graf & Cummings (2021) | |
Unio testudineus Morelet, 1849
Morelet, 1849. Test. Nov. 1: 28. Type(s): HOLOTYPE BMNH_94-3-22-1, flumen Usumacinta, Guatemala.
Unio (sec. Elliptio) testudineus — Simpson (1900), Simpson (1914) Unio (Psoronaias) semigranosus var. testudineus — Martens (1900) Psoronaias semigranosus — Frierson (1927) Psoronaias testudinea — Haas (1929), Haas (1969) Psoronaias testudineus — Haas (1930) Psoronaias testudinarea [sic] — Bogan (2010) | |
Unio morini Morelet, 1851
Morelet, 1851. Test. Nov. 2: 25. Type(s): HOLOTYPE BMNH_93-2-4-2026, Rio Usumacinta, Guatemala.
Unio (sec. Elliptio) morini — Simpson (1900), Simpson (1914) Unio (Psoronaias) morini — Martens (1900) Psoronaias morini — Frierson (1927), Haas (1929), Haas (1930), Haas (1969), Graf & Cummings (2007), Bogan (2010) Psoronaias semigranosus — Goodrich & van der Schalie (1937) Psoronaias semigranosa — Graf & Cummings (2021) | |
Unio psoricus Morelet, 1851
Morelet, 1851. Test. Nov. 2: 25. Type(s): HOLOTYPE BMNH_93-2-4-2033, Rio Usumacinta, Tabasco.
Unio (sec. Elliptio) psoricus — Simpson (1900), Simpson (1914) Unio (Psoronaias) psoricus — Martens (1900) Psoronaias psoricus — Frierson (1917), Haas (1930), Thiele (1934) Psoronaias semigranosus-psoricus — Frierson (1927) Psoronaias psorica — Haas (1929), Haas (1969), Haas (1969), Bogan (2010) Psoronaias semigranosus — Goodrich & van der Schalie (1937) Psoronaias semigranosa — Graf & Cummings (2007), Pfeiffer et al. (2019), Graf & Cummings (2021) | |
Unio corium Reeve, 1864
 Reeve, 1864. Conch. Icon. 16: pl. 10, sp. 39. Type(s): HOLOTYPE BMNH_1965209, Chiapas, Mexico.
Unio (sec. Elliptio) corium — Simpson (1900), Simpson (1914) Unio (Psoronaias) corium — Martens (1900) Unio corium — Pilsbry (1910) Psoronaias semigranosus-corium — Frierson (1927) Psoronaias semigranosus — Haas (1930), Goodrich & van der Schalie (1937) Psoronaias semigranosa corium — Haas (1969) | |
Unio crocodilorum var. B semipustulata Fischer & Crosse, 1894 Fischer & Crosse, 1894. Miss. Sci. Mexique, Rech. Zool. 2(16): 578, pl. 60, fig. 4. (intro'd as intra-sp. of Unio crocodilorum Morelet, 1849) Type(s): PARATYPES BMNH_1893-2-4-2032, River Usumacinta, Guatemala.
Unio (Psoronaias) crocodilorum — Martens (1900) Psoronaias crocodilorum — Frierson (1927), Haas (1969) Psoronaias crocodilarum [sic] — Haas (1929) | |
Unio crocodilorum var. G praestricta Fischer & Crosse, 1894
Fischer & Crosse, 1894. Miss. Sci. Mexique, Rech. Zool. 2(16): 578, pl. 60, fig. 5. (intro'd as intra-sp. of Unio crocodilorum Morelet, 1849) Type(s): SYNTYPES BMNH_1893-2-4-2031, River Usumacinta, Guatemala.
Unio (Psoronaias) crocodilorum — Martens (1900) Psoronaias crocodilorum — Frierson (1927), Haas (1969) | |
Quadrula rudis Simpson, 1900
 Simpson, 1900. Proc. Acad. Nat. Sci. 52: 82, pl. 3, fig. 2. Simpson, 1914. Descr. Cat. Naiades: 910. Type(s): HOLOTYPE (MONOTYPY) USNM_84210, Rio Taxtunilha, (tributary to the Rio de la Pasion), Guatemala.
— Simpson (1900) Quadrula rudis — Simpson (1900) Quadrula (Rotundaria) rudis — Simpson (1914) Psoronaias rudis — Frierson (1927) Rotundaria rudis — Haas (1929) Amblema (Psorula) rudis — Haas (1930) Amblema (sec. Psorula) rudis — Thiele (1934) Psoronaias semigranosus — Goodrich & van der Schalie (1937) Psorula rudis — Haas (1969), Graf & Cummings (2007), Bogan (2010), Pfeiffer et al. (2019), Inoue et al. (2020), Freeman et al. (2025) Psoronaias semigranosa — Graf & Cummings (2021) | |
Unio crocodilarum Simpson, 1900 ‘Morelet’ of Authors. Simpson, 1900. Proc. U.S. Nat. Mus. 22: 702. Simpson, 1914. Descr. Cat. Naiades: 591. Haas, 1929. Senckenbergiana 11: 332. (error for Unio crocodilorum Morelet, 1849) | |
Unio crocolilorum Lea, 1852 ‘Morelet’ Lea, 1852. Synopsis Fam. Naiades: 39. (error for Unio crocodilorum Morelet, 1849) | |
Unio testidineus Lea, 1852 ‘Morelet’ Lea, 1852. Synopsis Fam. Naiades: 40. (error for Unio testudineus Morelet, 1849) | |
Unio pliciferus Reeve, 1864
‘Lea’ Reeve, 1864. Conch. Icon. 16: pl. 10, sp. 37; errata [changed to ‘U. crocodilorum Morelet’]. (misidentified reference to Unio pliciferus Lea, 1838; error for Unio crocodilorum Morelet, 1849)
Unio (Psoronaias) crocodilorum — Martens (1900) Unio (sec. Elliptio) crocodilarum [sic] — Simpson (1900), Simpson (1914) Psoronaias crocodilorum — Frierson (1927) — Haas (1969) | |
Unio semigranosus Reeve, 1864
 ‘Philippi’ Reeve, 1864. Conch. Icon. 16: pl. 10, sp. 36; errata. (reference to Unio semigranosus Philippi, 1843) Type(s): FIGURED SPECIMEN BMNH_196496, Mexico.
Unio (sec. Elliptio) testudineus — Simpson (1900), Simpson (1914) Unio (Psoronaias) semigranosus var. testudineus — Martens (1900) | |
Psoronaias testudinarea Bogan, 2010 ‘Morelet’ Bogan, 2010. FADA: [web]. (error for Unio testudineus Morelet, 1849) | |
Taxonomic History (since 1900)Unio (sec. Elliptio) semigranosus — Simpson (1900), Simpson (1914) Unio (Psoronaias) semigranosus — Martens (1900) Unio (sec. Elliptio) crocodilarum [sic] — Simpson (1900), Simpson (1914) Unio (Psoronaias) crocodilorum — Martens (1900) Unio (sec. Elliptio) testudineus — Simpson (1900), Simpson (1914) Unio (Psoronaias) semigranosus var. testudineus — Martens (1900) Unio (sec. Elliptio) morini — Simpson (1900), Simpson (1914) Unio (Psoronaias) morini — Martens (1900) Unio (sec. Elliptio) psoricus — Simpson (1900), Simpson (1914) Unio (Psoronaias) psoricus — Martens (1900) Unio (sec. Elliptio) corium — Simpson (1900), Simpson (1914) Unio (Psoronaias) corium — Martens (1900) — Simpson (1900), Haas (1969) Quadrula rudis — Simpson (1900) Quadrula semigranosa — Pilsbry (1910) Unio corium — Pilsbry (1910) Quadrula (Rotundaria) rudis — Simpson (1914) Psoronaias semigranosus — Frierson (1917), Frierson (1927), Haas (1930), Goodrich & van der Schalie (1937) Psoronaias crocodilorum — Frierson (1917), Frierson (1927), Haas (1930), Haas (1969), Graf & Cummings (2007), Bogan (2010) Psoronaias psoricus — Frierson (1917), Haas (1930), Thiele (1934) Psoronaias morini — Frierson (1927), Haas (1929), Haas (1930), Haas (1969), Graf & Cummings (2007), Bogan (2010) Psoronaias semigranosus-psoricus — Frierson (1927) Psoronaias semigranosus-corium — Frierson (1927) Psoronaias rudis — Frierson (1927) Psoronaias semigranosa — Haas (1929), Graf & Cummings (2007), Bogan (2010), Pfeiffer et al. (2019), Pfeiffer et al. (2019), Inoue et al. (2020), Smith et al. (2020), Graf & Cummings (2021), Czaja et al. (2023), Freeman et al. (2025) Psoronaias crocodilarum [sic] — Haas (1929) Psoronaias testudinea — Haas (1929), Haas (1969) Psoronaias psorica — Haas (1929), Haas (1969), Haas (1969), Bogan (2010) Rotundaria rudis — Haas (1929) Psoronaias testudineus — Haas (1930) Amblema (Psorula) rudis — Haas (1930) Amblema (sec. Psorula) rudis — Thiele (1934) Psoronaias semigranosa semigranosa — Haas (1969) Psoronaias semigranosa corium — Haas (1969) Psorula rudis — Haas (1969), Graf & Cummings (2007), Bogan (2010), Pfeiffer et al. (2019), Inoue et al. (2020), Freeman et al. (2025) Psoronaias testudinarea [sic] — Bogan (2010)
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