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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

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Bivalvia | Unionoida | Unionoidea | Unionidae | Ambleminae | Popenaiadini | Psoronaias

species Psoronaias semigranosa (von dem Busch in Philippi, 1843): synonymy
     149 specimens + 3 Genbank vouchers

Unio semigranosus Philippi, 1843
von dem Busch in Philippi, 1843. Abbild. Beschr. Conch. 1: 19, pl. 1, figs. 1-3.
Type(s): PARATYPES MCZ_152861, near Tampico, Mexico.

Unio (sec. Elliptio) semigranosusSimpson (1900), Simpson (1914)
Unio (Psoronaias) semigranosusMartens (1900)
Quadrula semigranosaPilsbry (1910)
Psoronaias semigranosusFrierson (1917), Frierson (1927), Haas (1930), Goodrich & van der Schalie (1937)
Psoronaias semigranosaHaas (1929), Graf & Cummings (2007), Bogan (2010), Pfeiffer et al. (2019), Pfeiffer et al. (2019), Inoue et al. (2020), Smith et al. (2020), Graf & Cummings (2021), Czaja et al. (2023), Freeman et al. (2025)
Psoronaias semigranosa semigranosaHaas (1969)

Unio crocodilorum Morelet, 1849
Morelet, 1849. Test. Nov. 1: 28.
Type(s): SYNTYPE BMNH_1893-2-4-2030, River Usumacinta, Central America.

Unio (sec. Elliptio) crocodilarum [sic]Simpson (1900), Simpson (1914)
Unio (Psoronaias) crocodilorumMartens (1900)
Psoronaias crocodilorumFrierson (1917), Frierson (1927), Haas (1930), Haas (1969), Graf & Cummings (2007), Bogan (2010)
Psoronaias crocodilarum [sic]Haas (1929)
Psoronaias semigranosusGoodrich & van der Schalie (1937)
Psoronaias semigranosaGraf & Cummings (2021)

Unio testudineus Morelet, 1849
Morelet, 1849. Test. Nov. 1: 28.
Type(s): HOLOTYPE BMNH_94-3-22-1, flumen Usumacinta, Guatemala.

Unio (sec. Elliptio) testudineusSimpson (1900), Simpson (1914)
Unio (Psoronaias) semigranosus var. testudineusMartens (1900)
Psoronaias semigranosusFrierson (1927)
Psoronaias testudineaHaas (1929), Haas (1969)
Psoronaias testudineusHaas (1930)
Psoronaias testudinarea [sic]Bogan (2010)

Unio morini Morelet, 1851
Morelet, 1851. Test. Nov. 2: 25.
Type(s): HOLOTYPE BMNH_93-2-4-2026, Rio Usumacinta, Guatemala.

Unio (sec. Elliptio) moriniSimpson (1900), Simpson (1914)
Unio (Psoronaias) moriniMartens (1900)
Psoronaias moriniFrierson (1927), Haas (1929), Haas (1930), Haas (1969), Graf & Cummings (2007), Bogan (2010)
Psoronaias semigranosusGoodrich & van der Schalie (1937)
Psoronaias semigranosaGraf & Cummings (2021)

Unio psoricus Morelet, 1851
Morelet, 1851. Test. Nov. 2: 25.
Type(s): HOLOTYPE BMNH_93-2-4-2033, Rio Usumacinta, Tabasco.

Unio (sec. Elliptio) psoricusSimpson (1900), Simpson (1914)
Unio (Psoronaias) psoricusMartens (1900)
Psoronaias psoricusFrierson (1917), Haas (1930), Thiele (1934)
Psoronaias semigranosus-psoricusFrierson (1927)
Psoronaias psoricaHaas (1929), Haas (1969), Haas (1969), Bogan (2010)
Psoronaias semigranosusGoodrich & van der Schalie (1937)
Psoronaias semigranosaGraf & Cummings (2007), Pfeiffer et al. (2019), Graf & Cummings (2021)

Unio corium Reeve, 1864
Reeve, 1864. Conch. Icon. 16: pl. 10, sp. 39.
Type(s): HOLOTYPE BMNH_1965209, Chiapas, Mexico.

Unio (sec. Elliptio) coriumSimpson (1900), Simpson (1914)
Unio (Psoronaias) coriumMartens (1900)
Unio coriumPilsbry (1910)
Psoronaias semigranosus-coriumFrierson (1927)
Psoronaias semigranosusHaas (1930), Goodrich & van der Schalie (1937)
Psoronaias semigranosa coriumHaas (1969)

Unio crocodilorum var. B semipustulata Fischer & Crosse, 1894
Fischer & Crosse, 1894. Miss. Sci. Mexique, Rech. Zool. 2(16): 578, pl. 60, fig. 4.
(intro'd as intra-sp. of Unio crocodilorum Morelet, 1849)
Type(s): PARATYPES BMNH_1893-2-4-2032, River Usumacinta, Guatemala.

Unio (Psoronaias) crocodilorumMartens (1900)
Psoronaias crocodilorumFrierson (1927), Haas (1969)
Psoronaias crocodilarum [sic]Haas (1929)

Unio crocodilorum var. G praestricta Fischer & Crosse, 1894
Fischer & Crosse, 1894. Miss. Sci. Mexique, Rech. Zool. 2(16): 578, pl. 60, fig. 5.
(intro'd as intra-sp. of Unio crocodilorum Morelet, 1849)
Type(s): SYNTYPES BMNH_1893-2-4-2031, River Usumacinta, Guatemala.

Unio (Psoronaias) crocodilorumMartens (1900)
Psoronaias crocodilorumFrierson (1927), Haas (1969)

Quadrula rudis Simpson, 1900
Simpson, 1900. Proc. Acad. Nat. Sci. 52: 82, pl. 3, fig. 2.
Simpson, 1914. Descr. Cat. Naiades: 910.
Type(s): HOLOTYPE (MONOTYPY) USNM_84210, Rio Taxtunilha, (tributary to the Rio de la Pasion), Guatemala.

Simpson (1900)
Quadrula rudisSimpson (1900)
Quadrula (Rotundaria) rudisSimpson (1914)
Psoronaias rudisFrierson (1927)
Rotundaria rudisHaas (1929)
Amblema (Psorula) rudisHaas (1930)
Amblema (sec. Psorula) rudisThiele (1934)
Psoronaias semigranosusGoodrich & van der Schalie (1937)
Psorula rudisHaas (1969), Graf & Cummings (2007), Bogan (2010), Pfeiffer et al. (2019), Inoue et al. (2020), Freeman et al. (2025)
Psoronaias semigranosaGraf & Cummings (2021)

Unio crocodilarum Simpson, 1900
‘Morelet’ of Authors.
Simpson, 1900. Proc. U.S. Nat. Mus. 22: 702.
Simpson, 1914. Descr. Cat. Naiades: 591.
Haas, 1929. Senckenbergiana 11: 332.
(error for Unio crocodilorum Morelet, 1849)

Unio crocolilorum Lea, 1852
‘Morelet’ Lea, 1852. Synopsis Fam. Naiades: 39.
(error for Unio crocodilorum Morelet, 1849)

Unio testidineus Lea, 1852
‘Morelet’ Lea, 1852. Synopsis Fam. Naiades: 40.
(error for Unio testudineus Morelet, 1849)

Unio pliciferus Reeve, 1864
‘Lea’ Reeve, 1864. Conch. Icon. 16: pl. 10, sp. 37; errata [changed to ‘U. crocodilorum Morelet’].
(misidentified reference to Unio pliciferus Lea, 1838; error for Unio crocodilorum Morelet, 1849)

Unio (Psoronaias) crocodilorumMartens (1900)
Unio (sec. Elliptio) crocodilarum [sic]Simpson (1900), Simpson (1914)
Psoronaias crocodilorumFrierson (1927)
Haas (1969)

Unio semigranosus Reeve, 1864
‘Philippi’ Reeve, 1864. Conch. Icon. 16: pl. 10, sp. 36; errata.
(reference to Unio semigranosus Philippi, 1843)
Type(s): FIGURED SPECIMEN BMNH_196496, Mexico.

Unio (sec. Elliptio) testudineusSimpson (1900), Simpson (1914)
Unio (Psoronaias) semigranosus var. testudineusMartens (1900)

Psoronaias testudinarea Bogan, 2010
‘Morelet’ Bogan, 2010. FADA: [web].
(error for Unio testudineus Morelet, 1849)

Taxonomic History (since 1900)

Unio (sec. Elliptio) semigranosusSimpson (1900), Simpson (1914)
Unio (Psoronaias) semigranosusMartens (1900)
Unio (sec. Elliptio) crocodilarum [sic]Simpson (1900), Simpson (1914)
Unio (Psoronaias) crocodilorumMartens (1900)
Unio (sec. Elliptio) testudineusSimpson (1900), Simpson (1914)
Unio (Psoronaias) semigranosus var. testudineusMartens (1900)
Unio (sec. Elliptio) moriniSimpson (1900), Simpson (1914)
Unio (Psoronaias) moriniMartens (1900)
Unio (sec. Elliptio) psoricusSimpson (1900), Simpson (1914)
Unio (Psoronaias) psoricusMartens (1900)
Unio (sec. Elliptio) coriumSimpson (1900), Simpson (1914)
Unio (Psoronaias) coriumMartens (1900)
Simpson (1900), Haas (1969)
Quadrula rudisSimpson (1900)
Quadrula semigranosaPilsbry (1910)
Unio coriumPilsbry (1910)
Quadrula (Rotundaria) rudisSimpson (1914)
Psoronaias semigranosusFrierson (1917), Frierson (1927), Haas (1930), Goodrich & van der Schalie (1937)
Psoronaias crocodilorumFrierson (1917), Frierson (1927), Haas (1930), Haas (1969), Graf & Cummings (2007), Bogan (2010)
Psoronaias psoricusFrierson (1917), Haas (1930), Thiele (1934)
Psoronaias moriniFrierson (1927), Haas (1929), Haas (1930), Haas (1969), Graf & Cummings (2007), Bogan (2010)
Psoronaias semigranosus-psoricusFrierson (1927)
Psoronaias semigranosus-coriumFrierson (1927)
Psoronaias rudisFrierson (1927)
Psoronaias semigranosaHaas (1929), Graf & Cummings (2007), Bogan (2010), Pfeiffer et al. (2019), Pfeiffer et al. (2019), Inoue et al. (2020), Smith et al. (2020), Graf & Cummings (2021), Czaja et al. (2023), Freeman et al. (2025)
Psoronaias crocodilarum [sic]Haas (1929)
Psoronaias testudineaHaas (1929), Haas (1969)
Psoronaias psoricaHaas (1929), Haas (1969), Haas (1969), Bogan (2010)
Rotundaria rudisHaas (1929)
Psoronaias testudineusHaas (1930)
Amblema (Psorula) rudisHaas (1930)
Amblema (sec. Psorula) rudisThiele (1934)
Psoronaias semigranosa semigranosaHaas (1969)
Psoronaias semigranosa coriumHaas (1969)
Psorula rudisHaas (1969), Graf & Cummings (2007), Bogan (2010), Pfeiffer et al. (2019), Inoue et al. (2020), Freeman et al. (2025)
Psoronaias testudinarea [sic]Bogan (2010)

 
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