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Data last updated
Mon 1 Dec 2025
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Bivalvia
| Unionoida
| Unionoidea
| Unionidae
| Ambleminae
| Popenaiadini
| Psoronaias
species Psoronaias crocodilorum (Morelet, 1849): specimens17 available nominal species synonyms (25 total synonyms)
Recent Taxonomic Opinions
Barynaias pigerrima — Graf & Cummings (2007), Bogan (2010), Graf & Cummings (2021)
Psoronaias semigranosa — Graf & Cummings (2007), Pfeiffer et al. (2019), Pfeiffer et al. (2019), Inoue et al. (2020), Smith et al. (2020), Graf & Cummings (2021), Freeman et al. (2025)
Psoronaias crocodilorum — Graf & Cummings (2007), Bogan (2010), Cummings et al. (2025)
Psoronaias martensi — Graf & Cummings (2007), Bogan (2010), Graf & Cummings (2021)
Psoronaias morini — Graf & Cummings (2007), Bogan (2010)
Psoronaias ostreata — Graf & Cummings (2007), Bogan (2010), Graf & Cummings (2021), Czaja et al. (2023)
Psorula rudis — Graf & Cummings (2007), Bogan (2010), Pfeiffer et al. (2019), Inoue et al. (2020), Freeman et al. (2025)
Psorula percompressa — Graf & Cummings (2007), Bogan (2010), Pfeiffer et al. (2019), Inoue et al. (2020)
Psorula profunda — Graf & Cummings (2007), Bogan (2010), Pfeiffer et al. (2019), Inoue et al. (2020), Freeman et al. (2025)
Psorula quadrata — Graf & Cummings (2007), Bogan (2010)
Psorula salinarum — Graf & Cummings (2007), Bogan (2010)
Psorula usumasintae — Graf & Cummings (2007), Bogan (2010), Pfeiffer et al. (2019), Inoue et al. (2020), Freeman et al. (2025)
Psoronaias psorica — Bogan (2010)
Psoronaias testudinarea [sic] — Bogan (2010)
Psoronaias percompressa — Graf & Cummings (2021), Czaja et al. (2023)
Psoronaias profunda — Graf & Cummings (2021), Czaja et al. (2023)
Psoronaias quadrata — Graf & Cummings (2021)
Psoronaias salinarum — Graf & Cummings (2021)
Psoronaias usumasintae — Graf & Cummings (2021), Czaja et al. (2023)
Barynaias pigerrimus — Czaja et al. (2023)
Psorula percompressus — Freeman et al. (2025)
Range
Endemic to the Usumacinta basin, México and Guatemala
Map of Georeferenced Specimens
MUSSELp Specimen Records
Guatemala
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AMNH_140555
1 league [3 miles] above mouth Rio Santa Amelia, Peten, Guatemala
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MCZ_270070
3 mi above mouth Rio Santa Amelia, Peten, Guatemala
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MCZ_270071
3 mi above mouth Rio Santa Amelia, Peten, Guatemala
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MCZ_270073
3 mi above mouth Rio Santa Amelia, Peten, Guatemala
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MCZ_270074
3 mi above mouth Rio Santa Amelia, Peten, Guatemala
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MCZ_270078
3 mi above mouth Rio Santa Amelia, Peten, Guatemala
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UMMZ_65556
Arroyo Chocop, about 12 mi above Mactun, Dept. Peten, Guatemala
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BMNH_1965161-162
flumen Usumacinta, Guatemala
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BMNH_1965161-162
flumen Usumacinta, Guatemala
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BMNH_94-3-22-1
flumen Usumacinta, Guatemala
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BMNH_94-3-22-1
flumen Usumacinta, Guatemala
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BMNH_94-3-22-2
flumen Usumacinta, Guatemala
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BMNH_94-3-22-2
flumen Usumacinta, Guatemala
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BMNH_94-3-22-2
flumen Usumacinta, Guatemala
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UMMZ_76912
Guatemala
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UMMZ_76912
Guatemala
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UMMZ_65549
Pasion River, about 13 mi above Sayache [=Sayaxche], Dept. Peten, Guatemala
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UMMZ_65549
Pasion River, about 13 mi above Sayache [=Sayaxche], Dept. Peten, Guatemala
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UMMZ_65551
Pasion River, above mouth of Rio Santa Monica, Dept. Peten, Guatemala
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UMMZ_65545
Pasion River, at Cambio, Dept. Peten, Guatemala
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UMMZ_65543
Pasion River, at Embuido rapids, above Porvenir, Dept. Alta Vera Paz, Guatemala
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UMMZ_65541
Pasion River, at first arroyo below mouth of Rio San Simon, Dept. Alta Vera Paz, Guatemala
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UMMZ_65552
Pasion River, at mouth of Rio Martin, Dept. Peten, Guatemala
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UMMZ_65546
Pasion River, at road above Tres Islas, Dept. Peten, Guatemala
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UMMZ_65542
Pasion River, at rodal de Caresal, above Porvenir, Dept. Alta Vera Paz, Guatemala
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UMMZ_65548
Pasion River, at Santa Amelia, Dept. Peten, Guatemala
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UMMZ_65550
Pasion River, at Sayanche [=Sayaxche], Dept. Peten, Guatemala
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UMMZ_65547
Pasion River, at Tres Islas, Dept. Peten, Guatemala
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UMMZ_65553
Pasion River, below Sayarche [=Sayaxche], Dept. Peten, Guatemala
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UMMZ_65544
Pasion River, Rodal “Grande”, Alta Vera Paz, Guatemala
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ANSP_159973
Piedras Negras, Guatemala
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ANSP_159973
Piedras Negras, Guatemala
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ANSP_159973
Piedras Negras, Guatemala
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ANSP_159973
Piedras Negras, Guatemala
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SMF_15027
Rio Cancuen, Nebenfluss d. Rio Usumacinta, Guatemala
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ANSP_41665
Rio de la Pasion
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UF_507905
Rio De La Pasion, 5.5 miles east of Sayaxche, El Peten Department, Guatemala 16.536339, -90.103900
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UF_507905
Rio De La Pasion, 5.5 miles east of Sayaxche, El Peten Department, Guatemala 16.536339, -90.103900
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UF_507906
Rio De La Pasion, 5.5 miles east of Sayaxche, El Peten Department, Guatemala 16.536339, -90.103900
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UF_507907
Rio De La Pasion, 5.5 miles east of Sayaxche, El Peten Department, Guatemala 16.536339, -90.103900
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UF_507908
Rio De La Pasion, 5.5 miles east of Sayaxche, El Peten Department, Guatemala 16.536339, -90.103900
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UF_507908
Rio De La Pasion, 5.5 miles east of Sayaxche, El Peten Department, Guatemala 16.536339, -90.103900
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UF_507909
Rio De La Pasion, 5.5 miles east of Sayaxche, El Peten Department, Guatemala 16.536339, -90.103900
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SMF_15030
Rio de la Pasion, Guatemala (Paso Real)
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AMNH_140570
Rio de la Pasion, Peten, Guatemala
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MCZ_270076
Rio de la Pasion, Peten, Guatemala
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OSUM_18868
Rio de la Pasion, Peten, Guatemala
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USNM_783055
Rio de la Pasion, Santa Amelia, Guatemala
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USNM_783055
Rio de la Pasion, Santa Amelia, Guatemala
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USNM_783058
Rio de la Pasion, Santa Amelia, Guatemala
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USNM_783058
Rio de la Pasion, Santa Amelia, Guatemala
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USNM_783059
Rio de la Pasion, Santa Amelia, Guatemala
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USNM_783059
Rio de la Pasion, Santa Amelia, Guatemala
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USNM_783061
Rio de la Pasion, Santa Amelia, Guatemala
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USNM_783061
Rio de la Pasion, Santa Amelia, Guatemala
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USNM_783062
Rio de la Pasion, Santa Amelia, Guatemala
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USNM_783062
Rio de la Pasion, Santa Amelia, Guatemala
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USNM_783063
Rio de la Pasion, Santa Amelia, Guatemala
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USNM_783063
Rio de la Pasion, Santa Amelia, Guatemala
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AMNH_140556
Rio de la Pasion, Santa Amelia, Peten, Guatemala
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AMNH_140557
Rio de la Pasion, Santa Amelia, Peten, Guatemala
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AMNH_140558
Rio de la Pasion, Santa Amelia, Peten, Guatemala
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AMNH_140559
Rio de la Pasion, Santa Amelia, Peten, Guatemala
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AMNH_140560
Rio de la Pasion, Santa Amelia, Peten, Guatemala
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AMNH_140560
Rio de la Pasion, Santa Amelia, Peten, Guatemala
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AMNH_140561
Rio de la Pasion, Santa Amelia, Peten, Guatemala
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AMNH_140562
Rio de la Pasion, Santa Amelia, Peten, Guatemala
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AMNH_140564
Rio de la Pasion, Santa Amelia, Peten, Guatemala
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AMNH_140565
Rio de la Pasion, Santa Amelia, Peten, Guatemala
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AMNH_140571
Rio de la Pasion, Santa Amelia, Peten, Guatemala
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MCZ_270072
Rio de la Pasion, Santa Amelia, Peten, Guatemala
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USNM_783047
Rio de la Pasion, Santa Amelia, Peten, Guatemala
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USNM_783047
Rio de la Pasion, Santa Amelia, Peten, Guatemala
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USNM_783048
Rio de la Pasion, Santa Amelia, Peten, Guatemala
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USNM_783048
Rio de la Pasion, Santa Amelia, Peten, Guatemala
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USNM_783049
Rio de la Pasion, Santa Amelia, Peten, Guatemala
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USNM_783049
Rio de la Pasion, Santa Amelia, Peten, Guatemala
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USNM_783049
Rio de la Pasion, Santa Amelia, Peten, Guatemala
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USNM_783049
Rio de la Pasion, Santa Amelia, Peten, Guatemala
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USNM_783049
Rio de la Pasion, Santa Amelia, Peten, Guatemala
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USNM_783049
Rio de la Pasion, Santa Amelia, Peten, Guatemala
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USNM_783050
Rio de la Pasion, Santa Amelia, Peten, Guatemala
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USNM_783050
Rio de la Pasion, Santa Amelia, Peten, Guatemala
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USNM_783051
Rio de la Pasion, Santa Amelia, Peten, Guatemala
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USNM_783051
Rio de la Pasion, Santa Amelia, Peten, Guatemala
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USNM_783057
Rio de la Pasion, Santa Amelia, Peten, Guatemala
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USNM_783057
Rio de la Pasion, Santa Amelia, Peten, Guatemala
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USNM_783057
Rio de la Pasion, Santa Amelia, Peten, Guatemala
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MCZ_270075
Rio de la Pasion, Treasure Island, Peten, Guatemala
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MCZ_270077
Rio de la Pasion, Treasure Island, Peten, Guatemala
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SMF_15026
Rio de la Passion, Guatemala
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ZMB_108849
Rio de las Salinas, Guatemala
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ZMB_108849
Rio de las Salinas, Guatemala
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ZMB_39463
Rio de las Salinas, Guatemala
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ZMB_39463
Rio de las Salinas, Guatemala
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ZMB_39463
Rio de las Salinas, Guatemala
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ZMB_39466
Rio de las Salinas, Guatemala
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SMF_15023
Rio de las Salinas, N. Guatemala
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SMF_3487
Rio de las Salinas, N. Guatemala
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SMF_3487
Rio de las Salinas, N. Guatemala
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ANSP_185944
Rio Pasion, between Sabol and Sayaxche, Guatemala
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INHS_24857
Rio Petexbatum, Peten, Guatemala
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AMNH_140563
Rio San Pedro, Mactun, 96 km below Paso Caballas, Peten, Guatemala
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AMNH_140566
Rio San Pedro, Mactun, 96 km below Paso Caballas, Peten, Guatemala
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USNM_783046
Rio San Pedro, Mactun, 96 km below Paso Caballas, Peten, Guatemala
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USNM_783046
Rio San Pedro, Mactun, 96 km below Paso Caballas, Peten, Guatemala
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USNM_783046
Rio San Pedro, Mactun, 96 km below Paso Caballas, Peten, Guatemala
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USNM_795111
Rio San Pedro, Mactun, 96 km below Paso Caballas, Peten, Guatemala
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USNM_795111
Rio San Pedro, Mactun, 96 km below Paso Caballas, Peten, Guatemala
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USNM_795111
Rio San Pedro, Mactun, 96 km below Paso Caballas, Peten, Guatemala
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ANSP_125582
Rio Taxtunilha, (trib. to Rio de la Pasion), Guatemala
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USNM_84210
Rio Taxtunilha, (tributary to the Rio de la Pasion), Guatemala
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USNM_84210
Rio Taxtunilha, (tributary to the Rio de la Pasion), Guatemala
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USNM_84210
Rio Taxtunilha, (tributary to the Rio de la Pasion), Guatemala
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USNM_84210
Rio Taxtunilha, (tributary to the Rio de la Pasion), Guatemala
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MNHN-IM-2000-2102
Rio Usumacinta superior, Guatemala
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MNHN-IM-2000-2102
Rio Usumacinta superior, Guatemala
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INHS_24858
Rio Usumacinta, Alter of Sacrifice, Peten, Guatemala
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BMNH_93-2-4-2026
Rio Usumacinta, Guatemala
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BMNH_93-2-4-2026
Rio Usumacinta, Guatemala
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BMNH_93-2-4-2026
Rio Usumacinta, Guatemala
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MNHN_MP_0023
Rio Usumacinta, Guatemala
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MNHN_MP_0023
Rio Usumacinta, Guatemala
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MNHN_MP_1343
Rio Usumacinta, Guatemala
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MNHN-IM-2000-1667
Rio Usumacinta, Guatemala
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SMF_15024
Rio Usumacinta, Guatemala
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SMF_15028
Rio Usumacinta, Guatemala
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SMF_15029
Rio Usumacinta, Guatemala
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SMF_3475
Rio Usumacinta, Guatemala
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SMF_3476
Rio Usumacinta, Guatemala
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USNM_128984
Rio Usumacinta, Guatemala
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USNM_128984
Rio Usumacinta, Guatemala
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USNM_128988
Rio Usumacinta, Guatemala
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USNM_128988
Rio Usumacinta, Guatemala
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BMNH_93-2-4-2045
Rio Usumacinta, Tabasco Superieur, Mexico
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BMNH_93-2-4-2045
Rio Usumacinta, Tabasco Superieur, Mexico
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USNM_347187
Rio Usumacinta, Tabasco, Mexico
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USNM_347187
Rio Usumacinta, Tabasco, Mexico
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USNM_792459
Rio Usumasinta, Guatemala
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USNM_792459
Rio Usumasinta, Guatemala
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BMNH_1893-2-4-2030
River Usumacinta, Central America
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BMNH_1893-2-4-2031
River Usumacinta, Guatemala
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BMNH_1908-12-15-4
River Usumacinta, Guatemala
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UMMZ_65558
San Pedro River, 4-6 mi E Paso Caballo, Dept. Peten, Guatemala
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UMMZ_65557
San Pedro River, Mactun, Dept. Peten, Guatemala
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UMMZ_65554
Subin River, about 2 mi above mouth, Dept. Peten, Guatemala
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UMMZ_65555
Subin River, Santa Teresa, Dept. Peten, Guatemala
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USNM_128992_01
Usumacinta R., Guatemala
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USNM_128992_01
Usumacinta R., Guatemala
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USNM_128987
Usumacinta River, Guatemala
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USNM_128987
Usumacinta River, Guatemala
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USNM_128987
Usumacinta River, Guatemala
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USNM_128996
Usumacinta River, Guatemala
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USNM_128996
Usumacinta River, Guatemala
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OSUM_18870
Usumacinta River, Machtun, 96 km below Paso Caballos [at or very near Mexican border]
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MCZ_293588
Usumacinta River, Yaxchilan, 106 mi downstream from Sayache, Mexico
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MCZ_293589
Usumacinta River, Yaxchilan, 106 mi downstream from Sayache, Mexico
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COMA_2048. Rio Usumacinta, Tabasco, Mexico BMNH_1893-2-4-2032. River Usumacinta, Guatemala
Mexico
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MCZ_72567
Balancan in Usumacinta R., Tabasco, Mexico
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BMNH_1965209
Chiapas, Mexico
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BMNH_1965209
Chiapas, Mexico
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UF_270399
Chiapas, Mexico
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ANSP_138664
Hacienda Reforma on the San Pedro River, trib. of the Usumacinta above Balancan, Mexico
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ANSP_138664
Hacienda Reforma on the San Pedro River, trib. of the Usumacinta above Balancan, Mexico
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ANSP_62937
lower part of Rio Tulija, State Tabasco, Mexico
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BMNH_1907-10-28-181
Mexico
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BMNH_196496
Mexico
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BMNH_196496
Mexico
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MNHN_MP_1340
Mexico
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UF_270520
Mexico
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USNM_84335
Mexico
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USNM_84335
Mexico
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ZMB_MP_0486
Mexico
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MNHN-IM-2000-1761
Mexique
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MNHN-IM-2000-1761
Mexique
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USNM_161475
Monte Cristo, Tabasco, Mexico
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USNM_161475
Monte Cristo, Tabasco, Mexico
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USNM_161475
Monte Cristo, Tabasco, Mexico
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USNM_161475
Monte Cristo, Tabasco, Mexico
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USNM_359337
near San Cristobal, Mexico
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USNM_359337
near San Cristobal, Mexico
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USNM_795617
pond W of Rio Grijalva, about 3 km above Villahermosa, Tabasco, Mexico
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USNM_795617
pond W of Rio Grijalva, about 3 km above Villahermosa, Tabasco, Mexico
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INHS_91077
Rio Saguila, Gulf of Mexico - Grijalva - Usumacinta - Chacamax, SE Emiliano Zapata, off Rt. 10, fish camp, Tabasco, Mexico
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INHS_91078
Rio Saguila, Gulf of Mexico - Grijalva - Usumacinta - Chacamax, SE Emiliano Zapata, off Rt. 10, fish camp, Tabasco, Mexico
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INHS_91078
Rio Saguila, Gulf of Mexico - Grijalva - Usumacinta - Chacamax, SE Emiliano Zapata, off Rt. 10, fish camp, Tabasco, Mexico
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INHS_91079
Rio Saguila, Gulf of Mexico - Grijalva - Usumacinta - Chacamax, SE Emiliano Zapata, off Rt. 10, fish camp, Tabasco, Mexico
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UF_164057
Rio San Pedro, 3 km N La Palma, Tabasco, Mexico.
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UF_164058
Rio San Pedro, 3 km N La Palma, Tabasco, Mexico.
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UF_164059
Rio San Pedro, 3 km N La Palma, Tabasco, Mexico.
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UF_517941
Rio San Pedro, Bahia Campeche 3 km N of La Palma Tabasco, Mexico
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UF_164065
Rio San Pedro, Cascadas de la Reforma, 7 km N and 7 km E El Aguila, Tabasco, Mexico
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UF_164066
Rio San Pedro, Cascadas de la Reforma, 7 km N and 7 km E El Aguila, Tabasco, Mexico
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UF_164067
Rio San Pedro, Limon, Tabasco, Mexico
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USNM_675261
Rio Usumacinta, 9.4 mi NNW Emillano Zapata, Mexico
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USNM_675261
Rio Usumacinta, 9.4 mi NNW Emillano Zapata, Mexico
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USNM_675261
Rio Usumacinta, 9.4 mi NNW Emillano Zapata, Mexico
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USNM_267924
Rio Usumacinta, 9.4 mi NNW Emillano Zapata, Tabasco, Mexico
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INHS_24859
Rio Usumacinta, Boca Lacantum, Chiapas, Mexico
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INHS_24859
Rio Usumacinta, Boca Lacantum, Chiapas, Mexico
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INHS_24860
Rio Usumacinta, Boca Lacantum, Chiapas, Mexico
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AMS_102111
Rio Usumacinta, Emiliano Zapata, Tabasco, Mexico
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MCZ_288412
Rio Usumacinta, Emiliano Zapata, Tabasco, Mexico
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UF_190664
Rio Usumacinta, Emiliano Zapata, Tabasco, Mexico
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UF_378849
Rio Usumacinta, Emilliano Zapata, Tabasco, Mexico 17.7333, -91.8
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USNM_251192
Rio Usumacinta, near Jonuta, Tabasco, Mexico
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USNM_251192
Rio Usumacinta, near Jonuta, Tabasco, Mexico
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BMNH_93-2-4-2033
Rio Usumacinta, Tabasco
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BMNH_93-2-4-2033
Rio Usumacinta, Tabasco
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USNM_675260
Rio Usumacinta, Tabasco, Emillano Zapata, Mexico
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USNM_675260
Rio Usumacinta, Tabasco, Emillano Zapata, Mexico
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UF_270396
Tabasco, Mexico
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USNM_795616
trib. of Rio Grijalva, 1 km above Villa Hermosa, Tabasco, Mexico
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USNM_795616
trib. of Rio Grijalva, 1 km above Villa Hermosa, Tabasco, Mexico
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UF_507899
Usumacinta River, opposite of Emiliano Zapata. Tabasco, Mexico. 17.749685, -91.766995
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COMA_B.1053. Río Candelaria, Campeche, México COMA_B.0392. Rio González, Laguna del Horizonte, Tabasco, México UF_529129. Rio San Pedro, Bahia Campeche 3 km N of La Palma Tabasco, Mexico BMNH_93-2-4-2034. Rio Usumacinta, Tabasco COMA_B.0720. Río Usumacinta, Tabasco, México MCZ_288418B. small river, 3 mi SE Emiliano Zapata, Tabasco, Mexico
unknown
Genbank Vouchers
Georeferenced Genbank Vouchers
Mexico
UF_507899. (SRA: SRR8473037, go to Genbank) Mexico, Tabasco, Usumacinta River, opposite Emiliano Zapata (17.749685, -91.766995) .
UF_507899 (2015-0722). (16S: MK001775, 28S: MK001809, COI: MK001758, go to Genbank) ditto.
UF_507899 (2015-0726). (16S: MK001776, 28S: MK001810, COI: MK001759, go to Genbank) ditto.
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