MUSSELpdb | valid sp. Psoronaias crocodilorum specimens

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Page last updated
Wed 1 October 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

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Bivalvia | Unionoida | Unionoidea | Unionidae | Ambleminae | Popenaiadini | Psoronaias

species Psoronaias crocodilorum (Morelet, 1849): specimens
17 available nominal species synonyms (25 total synonyms)

Recent Taxonomic Opinions

Psoronaias crocodilorum — Graf & Cummings (2007), Bogan (2010), Cummings et al. (2025)
Psoronaias ostreata — Graf & Cummings (2007), Bogan (2010), Graf & Cummings (2021), Czaja et al. (2023)
Psoronaias morini — Graf & Cummings (2007), Bogan (2010)
Psoronaias semigranosa — Graf & Cummings (2007), Pfeiffer et al. (2019), Graf & Cummings (2021)
Psorula percompressa — Graf & Cummings (2007), Bogan (2010), Pfeiffer et al. (2019), Inoue et al. (2020)
Psorula usumasintae — Graf & Cummings (2007), Bogan (2010), Pfeiffer et al. (2019), Inoue et al. (2020), Freeman et al. (2025)
Barynaias pigerrima — Graf & Cummings (2007), Bogan (2010), Graf & Cummings (2021)
Psorula rudis — Graf & Cummings (2007), Bogan (2010), Pfeiffer et al. (2019), Inoue et al. (2020), Freeman et al. (2025)
Psorula profunda — Graf & Cummings (2007), Bogan (2010), Pfeiffer et al. (2019), Inoue et al. (2020), Freeman et al. (2025)
Psorula quadrata — Graf & Cummings (2007), Bogan (2010)
Psoronaias martensi — Graf & Cummings (2007), Bogan (2010), Graf & Cummings (2021)
Psorula salinarum — Graf & Cummings (2007), Bogan (2010)
Psoronaias testudinarea [sic] — Bogan (2010)
Psoronaias psorica — Bogan (2010)
Psoronaias percompressa — Graf & Cummings (2021), Czaja et al. (2023)
Psoronaias usumasintae — Graf & Cummings (2021), Czaja et al. (2023)
Psoronaias profunda — Graf & Cummings (2021), Czaja et al. (2023)
Psoronaias quadrata — Graf & Cummings (2021)
Psoronaias salinarum — Graf & Cummings (2021)
Barynaias pigerrimus — Czaja et al. (2023)
Psorula percompressus — Freeman et al. (2025)

Range

Endemic to the Usumacinta basin, México and Guatemala

Map of Georeferenced Specimens

species map

SMF_15025
Cochinchine Jung ?

Guatemala

AMNH_140555 1 league [3 miles] above mouth Rio Santa Amelia, Peten, Guatemala	MCZ_270070 3 mi above mouth Rio Santa Amelia, Peten, Guatemala	MCZ_270071 3 mi above mouth Rio Santa Amelia, Peten, Guatemala
MCZ_270073 3 mi above mouth Rio Santa Amelia, Peten, Guatemala	MCZ_270074 3 mi above mouth Rio Santa Amelia, Peten, Guatemala	MCZ_270078 3 mi above mouth Rio Santa Amelia, Peten, Guatemala
UMMZ_65556 Arroyo Chocop, about 12 mi above Mactun, Dept. Peten, Guatemala	BMNH_1965161-162 flumen Usumacinta, Guatemala	BMNH_1965161-162 flumen Usumacinta, Guatemala
BMNH_94-3-22-1 flumen Usumacinta, Guatemala	BMNH_94-3-22-1 flumen Usumacinta, Guatemala	BMNH_94-3-22-2 flumen Usumacinta, Guatemala
BMNH_94-3-22-2 flumen Usumacinta, Guatemala	BMNH_94-3-22-2 flumen Usumacinta, Guatemala	UMMZ_76912 Guatemala
UMMZ_76912 Guatemala	UMMZ_65549 Pasion River, about 13 mi above Sayache [=Sayaxche], Dept. Peten, Guatemala	UMMZ_65549 Pasion River, about 13 mi above Sayache [=Sayaxche], Dept. Peten, Guatemala
UMMZ_65551 Pasion River, above mouth of Rio Santa Monica, Dept. Peten, Guatemala	UMMZ_65545 Pasion River, at Cambio, Dept. Peten, Guatemala	UMMZ_65543 Pasion River, at Embuido rapids, above Porvenir, Dept. Alta Vera Paz, Guatemala
UMMZ_65541 Pasion River, at first arroyo below mouth of Rio San Simon, Dept. Alta Vera Paz, Guatemala	UMMZ_65552 Pasion River, at mouth of Rio Martin, Dept. Peten, Guatemala	UMMZ_65546 Pasion River, at road above Tres Islas, Dept. Peten, Guatemala
UMMZ_65542 Pasion River, at rodal de Caresal, above Porvenir, Dept. Alta Vera Paz, Guatemala	UMMZ_65548 Pasion River, at Santa Amelia, Dept. Peten, Guatemala	UMMZ_65550 Pasion River, at Sayanche [=Sayaxche], Dept. Peten, Guatemala
UMMZ_65547 Pasion River, at Tres Islas, Dept. Peten, Guatemala	UMMZ_65553 Pasion River, below Sayarche [=Sayaxche], Dept. Peten, Guatemala	UMMZ_65544 Pasion River, Rodal “Grande”, Alta Vera Paz, Guatemala
ANSP_159973 Piedras Negras, Guatemala	ANSP_159973 Piedras Negras, Guatemala	ANSP_159973 Piedras Negras, Guatemala
ANSP_159973 Piedras Negras, Guatemala	SMF_15027 Rio Cancuen, Nebenfluss d. Rio Usumacinta, Guatemala	ANSP_41665 Rio de la Pasion
UF_507905 Rio De La Pasion, 5.5 miles east of Sayaxche, El Peten Department, Guatemala 16.536339, -90.103900	UF_507905 Rio De La Pasion, 5.5 miles east of Sayaxche, El Peten Department, Guatemala 16.536339, -90.103900	UF_507906 Rio De La Pasion, 5.5 miles east of Sayaxche, El Peten Department, Guatemala 16.536339, -90.103900
UF_507907 Rio De La Pasion, 5.5 miles east of Sayaxche, El Peten Department, Guatemala 16.536339, -90.103900	UF_507908 Rio De La Pasion, 5.5 miles east of Sayaxche, El Peten Department, Guatemala 16.536339, -90.103900	UF_507908 Rio De La Pasion, 5.5 miles east of Sayaxche, El Peten Department, Guatemala 16.536339, -90.103900
UF_507909 Rio De La Pasion, 5.5 miles east of Sayaxche, El Peten Department, Guatemala 16.536339, -90.103900	SMF_15030 Rio de la Pasion, Guatemala (Paso Real)	AMNH_140570 Rio de la Pasion, Peten, Guatemala
MCZ_270076 Rio de la Pasion, Peten, Guatemala	OSUM_18868 Rio de la Pasion, Peten, Guatemala	USNM_783055 Rio de la Pasion, Santa Amelia, Guatemala
USNM_783055 Rio de la Pasion, Santa Amelia, Guatemala	USNM_783058 Rio de la Pasion, Santa Amelia, Guatemala	USNM_783058 Rio de la Pasion, Santa Amelia, Guatemala
USNM_783059 Rio de la Pasion, Santa Amelia, Guatemala	USNM_783059 Rio de la Pasion, Santa Amelia, Guatemala	USNM_783061 Rio de la Pasion, Santa Amelia, Guatemala
USNM_783061 Rio de la Pasion, Santa Amelia, Guatemala	USNM_783062 Rio de la Pasion, Santa Amelia, Guatemala	USNM_783062 Rio de la Pasion, Santa Amelia, Guatemala
USNM_783063 Rio de la Pasion, Santa Amelia, Guatemala	USNM_783063 Rio de la Pasion, Santa Amelia, Guatemala	AMNH_140556 Rio de la Pasion, Santa Amelia, Peten, Guatemala
AMNH_140557 Rio de la Pasion, Santa Amelia, Peten, Guatemala	AMNH_140558 Rio de la Pasion, Santa Amelia, Peten, Guatemala	AMNH_140559 Rio de la Pasion, Santa Amelia, Peten, Guatemala
AMNH_140560 Rio de la Pasion, Santa Amelia, Peten, Guatemala	AMNH_140560 Rio de la Pasion, Santa Amelia, Peten, Guatemala	AMNH_140561 Rio de la Pasion, Santa Amelia, Peten, Guatemala
AMNH_140562 Rio de la Pasion, Santa Amelia, Peten, Guatemala	AMNH_140564 Rio de la Pasion, Santa Amelia, Peten, Guatemala	AMNH_140565 Rio de la Pasion, Santa Amelia, Peten, Guatemala
AMNH_140571 Rio de la Pasion, Santa Amelia, Peten, Guatemala	MCZ_270072 Rio de la Pasion, Santa Amelia, Peten, Guatemala	USNM_783047 Rio de la Pasion, Santa Amelia, Peten, Guatemala
USNM_783047 Rio de la Pasion, Santa Amelia, Peten, Guatemala	USNM_783048 Rio de la Pasion, Santa Amelia, Peten, Guatemala	USNM_783048 Rio de la Pasion, Santa Amelia, Peten, Guatemala
USNM_783049 Rio de la Pasion, Santa Amelia, Peten, Guatemala	USNM_783049 Rio de la Pasion, Santa Amelia, Peten, Guatemala	USNM_783049 Rio de la Pasion, Santa Amelia, Peten, Guatemala
USNM_783049 Rio de la Pasion, Santa Amelia, Peten, Guatemala	USNM_783049 Rio de la Pasion, Santa Amelia, Peten, Guatemala	USNM_783049 Rio de la Pasion, Santa Amelia, Peten, Guatemala
USNM_783050 Rio de la Pasion, Santa Amelia, Peten, Guatemala	USNM_783050 Rio de la Pasion, Santa Amelia, Peten, Guatemala	USNM_783051 Rio de la Pasion, Santa Amelia, Peten, Guatemala
USNM_783051 Rio de la Pasion, Santa Amelia, Peten, Guatemala	USNM_783057 Rio de la Pasion, Santa Amelia, Peten, Guatemala	USNM_783057 Rio de la Pasion, Santa Amelia, Peten, Guatemala
USNM_783057 Rio de la Pasion, Santa Amelia, Peten, Guatemala	MCZ_270075 Rio de la Pasion, Treasure Island, Peten, Guatemala	MCZ_270077 Rio de la Pasion, Treasure Island, Peten, Guatemala
SMF_15026 Rio de la Passion, Guatemala	ZMB_108849 Rio de las Salinas, Guatemala	ZMB_108849 Rio de las Salinas, Guatemala
ZMB_39463 Rio de las Salinas, Guatemala	ZMB_39463 Rio de las Salinas, Guatemala	ZMB_39463 Rio de las Salinas, Guatemala
ZMB_39466 Rio de las Salinas, Guatemala	SMF_15023 Rio de las Salinas, N. Guatemala	SMF_3487 Rio de las Salinas, N. Guatemala
SMF_3487 Rio de las Salinas, N. Guatemala	ANSP_185944 Rio Pasion, between Sabol and Sayaxche, Guatemala	INHS_24857 Rio Petexbatum, Peten, Guatemala
AMNH_140563 Rio San Pedro, Mactun, 96 km below Paso Caballas, Peten, Guatemala	AMNH_140566 Rio San Pedro, Mactun, 96 km below Paso Caballas, Peten, Guatemala	USNM_783046 Rio San Pedro, Mactun, 96 km below Paso Caballas, Peten, Guatemala
USNM_783046 Rio San Pedro, Mactun, 96 km below Paso Caballas, Peten, Guatemala	USNM_783046 Rio San Pedro, Mactun, 96 km below Paso Caballas, Peten, Guatemala	USNM_795111 Rio San Pedro, Mactun, 96 km below Paso Caballas, Peten, Guatemala
USNM_795111 Rio San Pedro, Mactun, 96 km below Paso Caballas, Peten, Guatemala	USNM_795111 Rio San Pedro, Mactun, 96 km below Paso Caballas, Peten, Guatemala	ANSP_125582 Rio Taxtunilha, (trib. to Rio de la Pasion), Guatemala
USNM_84210 Rio Taxtunilha, (tributary to the Rio de la Pasion), Guatemala	USNM_84210 Rio Taxtunilha, (tributary to the Rio de la Pasion), Guatemala	USNM_84210 Rio Taxtunilha, (tributary to the Rio de la Pasion), Guatemala
USNM_84210 Rio Taxtunilha, (tributary to the Rio de la Pasion), Guatemala	MNHN-IM-2000-2102 Rio Usumacinta superior, Guatemala	MNHN-IM-2000-2102 Rio Usumacinta superior, Guatemala
INHS_24858 Rio Usumacinta, Alter of Sacrifice, Peten, Guatemala	BMNH_93-2-4-2026 Rio Usumacinta, Guatemala	BMNH_93-2-4-2026 Rio Usumacinta, Guatemala
BMNH_93-2-4-2026 Rio Usumacinta, Guatemala	MNHN_MP_0023 Rio Usumacinta, Guatemala	MNHN_MP_0023 Rio Usumacinta, Guatemala
MNHN_MP_1343 Rio Usumacinta, Guatemala	MNHN-IM-2000-1667 Rio Usumacinta, Guatemala	SMF_15024 Rio Usumacinta, Guatemala
SMF_15028 Rio Usumacinta, Guatemala	SMF_15029 Rio Usumacinta, Guatemala	SMF_3475 Rio Usumacinta, Guatemala
SMF_3476 Rio Usumacinta, Guatemala	USNM_128984 Rio Usumacinta, Guatemala	USNM_128984 Rio Usumacinta, Guatemala
USNM_128988 Rio Usumacinta, Guatemala	USNM_128988 Rio Usumacinta, Guatemala	BMNH_93-2-4-2045 Rio Usumacinta, Tabasco Superieur, Mexico
BMNH_93-2-4-2045 Rio Usumacinta, Tabasco Superieur, Mexico	USNM_347187 Rio Usumacinta, Tabasco, Mexico	USNM_347187 Rio Usumacinta, Tabasco, Mexico
USNM_792459 Rio Usumasinta, Guatemala	USNM_792459 Rio Usumasinta, Guatemala	BMNH_1893-2-4-2030 River Usumacinta, Central America
BMNH_1893-2-4-2031 River Usumacinta, Guatemala	BMNH_1908-12-15-4 River Usumacinta, Guatemala	UMMZ_65558 San Pedro River, 4-6 mi E Paso Caballo, Dept. Peten, Guatemala
UMMZ_65557 San Pedro River, Mactun, Dept. Peten, Guatemala	UMMZ_65554 Subin River, about 2 mi above mouth, Dept. Peten, Guatemala	UMMZ_65555 Subin River, Santa Teresa, Dept. Peten, Guatemala
USNM_128992_01 Usumacinta R., Guatemala	USNM_128992_01 Usumacinta R., Guatemala	USNM_128987 Usumacinta River, Guatemala
USNM_128987 Usumacinta River, Guatemala	USNM_128987 Usumacinta River, Guatemala	USNM_128996 Usumacinta River, Guatemala
USNM_128996 Usumacinta River, Guatemala	OSUM_18870 Usumacinta River, Machtun, 96 km below Paso Caballos [at or very near Mexican border]	MCZ_293588 Usumacinta River, Yaxchilan, 106 mi downstream from Sayache, Mexico
MCZ_293589 Usumacinta River, Yaxchilan, 106 mi downstream from Sayache, Mexico

COMA_2048. Rio Usumacinta, Tabasco, Mexico

BMNH_1893-2-4-2032. River Usumacinta, Guatemala

Mexico

MCZ_72567 Balancan in Usumacinta R., Tabasco, Mexico	BMNH_1965209 Chiapas, Mexico	BMNH_1965209 Chiapas, Mexico
UF_270399 Chiapas, Mexico	ANSP_138664 Hacienda Reforma on the San Pedro River, trib. of the Usumacinta above Balancan, Mexico	ANSP_138664 Hacienda Reforma on the San Pedro River, trib. of the Usumacinta above Balancan, Mexico
ANSP_62937 lower part of Rio Tulija, State Tabasco, Mexico	BMNH_1907-10-28-181 Mexico	BMNH_196496 Mexico
BMNH_196496 Mexico	MNHN_MP_1340 Mexico	UF_270520 Mexico
USNM_84335 Mexico	USNM_84335 Mexico	ZMB_MP_0486 Mexico
MNHN-IM-2000-1761 Mexique	MNHN-IM-2000-1761 Mexique	USNM_161475 Monte Cristo, Tabasco, Mexico
USNM_161475 Monte Cristo, Tabasco, Mexico	USNM_161475 Monte Cristo, Tabasco, Mexico	USNM_161475 Monte Cristo, Tabasco, Mexico
USNM_359337 near San Cristobal, Mexico	USNM_359337 near San Cristobal, Mexico	USNM_795617 pond W of Rio Grijalva, about 3 km above Villahermosa, Tabasco, Mexico
USNM_795617 pond W of Rio Grijalva, about 3 km above Villahermosa, Tabasco, Mexico	INHS_91077 Rio Saguila, Gulf of Mexico - Grijalva - Usumacinta - Chacamax, SE Emiliano Zapata, off Rt. 10, fish camp, Tabasco, Mexico	INHS_91078 Rio Saguila, Gulf of Mexico - Grijalva - Usumacinta - Chacamax, SE Emiliano Zapata, off Rt. 10, fish camp, Tabasco, Mexico
INHS_91078 Rio Saguila, Gulf of Mexico - Grijalva - Usumacinta - Chacamax, SE Emiliano Zapata, off Rt. 10, fish camp, Tabasco, Mexico	INHS_91079 Rio Saguila, Gulf of Mexico - Grijalva - Usumacinta - Chacamax, SE Emiliano Zapata, off Rt. 10, fish camp, Tabasco, Mexico	UF_164057 Rio San Pedro, 3 km N La Palma, Tabasco, Mexico.
UF_164058 Rio San Pedro, 3 km N La Palma, Tabasco, Mexico.	UF_164059 Rio San Pedro, 3 km N La Palma, Tabasco, Mexico.	UF_517941 Rio San Pedro, Bahia Campeche 3 km N of La Palma Tabasco, Mexico
UF_164065 Rio San Pedro, Cascadas de la Reforma, 7 km N and 7 km E El Aguila, Tabasco, Mexico	UF_164066 Rio San Pedro, Cascadas de la Reforma, 7 km N and 7 km E El Aguila, Tabasco, Mexico	UF_164067 Rio San Pedro, Limon, Tabasco, Mexico
USNM_675261 Rio Usumacinta, 9.4 mi NNW Emillano Zapata, Mexico	USNM_675261 Rio Usumacinta, 9.4 mi NNW Emillano Zapata, Mexico	USNM_675261 Rio Usumacinta, 9.4 mi NNW Emillano Zapata, Mexico
USNM_267924 Rio Usumacinta, 9.4 mi NNW Emillano Zapata, Tabasco, Mexico	INHS_24859 Rio Usumacinta, Boca Lacantum, Chiapas, Mexico	INHS_24859 Rio Usumacinta, Boca Lacantum, Chiapas, Mexico
INHS_24860 Rio Usumacinta, Boca Lacantum, Chiapas, Mexico	AMS_102111 Rio Usumacinta, Emiliano Zapata, Tabasco, Mexico	MCZ_288412 Rio Usumacinta, Emiliano Zapata, Tabasco, Mexico
UF_190664 Rio Usumacinta, Emiliano Zapata, Tabasco, Mexico	UF_378849 Rio Usumacinta, Emilliano Zapata, Tabasco, Mexico 17.7333, -91.8	USNM_251192 Rio Usumacinta, near Jonuta, Tabasco, Mexico
USNM_251192 Rio Usumacinta, near Jonuta, Tabasco, Mexico	BMNH_93-2-4-2033 Rio Usumacinta, Tabasco	BMNH_93-2-4-2033 Rio Usumacinta, Tabasco
USNM_675260 Rio Usumacinta, Tabasco, Emillano Zapata, Mexico	USNM_675260 Rio Usumacinta, Tabasco, Emillano Zapata, Mexico	UF_270396 Tabasco, Mexico
USNM_795616 trib. of Rio Grijalva, 1 km above Villa Hermosa, Tabasco, Mexico	USNM_795616 trib. of Rio Grijalva, 1 km above Villa Hermosa, Tabasco, Mexico	UF_507899 Usumacinta River, opposite of Emiliano Zapata. Tabasco, Mexico. 17.749685, -91.766995

COMA_B.1053. Río Candelaria, Campeche, México

COMA_B.0392. Rio González, Laguna del Horizonte, Tabasco, México

UF_529129. Rio San Pedro, Bahia Campeche 3 km N of La Palma Tabasco, Mexico

BMNH_93-2-4-2034. Rio Usumacinta, Tabasco

COMA_B.0720. Río Usumacinta, Tabasco, México

MCZ_288418B. small river, 3 mi SE Emiliano Zapata, Tabasco, Mexico

unknown

FMNH_15001 [no locality available - Central America]	ZMB_MP_0472 [no locality available - Central America]	ZMB_MP_0472 [no locality available - Central America]
ZMB_MP_0472 [no locality available - Central America]	ZMB_MP_0490 Amer Mer.	ZMB_22255 Central America [locality illegible]
ZMB_22255 Central America [locality illegible]	ZMB_22255 Central America [locality illegible]

Georeferenced Genbank Vouchers

Mexico

UF_507899. (SRA: SRR8473037, go to Genbank) Mexico, Tabasco, Usumacinta River, opposite Emiliano Zapata (17.749685, -91.766995) .

UF_507899 (2015-0722). (16S: MK001775, 28S: MK001809, COI: MK001758, go to Genbank) ditto.

UF_507899 (2015-0726). (16S: MK001776, 28S: MK001810, COI: MK001759, go to Genbank) ditto.