Bivalvia | Unionida | Unionoidea | Unionidae | Ambleminae | Popenaiadini | Psoronaias

species Psoronaias crocodilorum (Morelet, 1849): specimens

17 available nominal species synonyms (25 total synonyms)

Recent Taxonomic Opinions

Barynaias pigerrima — Graf & Cummings (2007), Bogan (2010), Graf & Cummings (2021)
Psoronaias semigranosa — Graf & Cummings (2007), Pfeiffer et al. (2019), Graf & Cummings (2021)
Psoronaias crocodilorum — Graf & Cummings (2007), Bogan (2010), Cummings et al. (2025)
Psoronaias martensi — Graf & Cummings (2007), Bogan (2010), Graf & Cummings (2021)
Psoronaias morini — Graf & Cummings (2007), Bogan (2010)
Psoronaias ostreata — Graf & Cummings (2007), Bogan (2010), Graf & Cummings (2021), Czaja et al. (2023)
Psorula rudis — Graf & Cummings (2007), Bogan (2010), Pfeiffer et al. (2019), Inoue et al. (2020), Freeman et al. (2025)
Psorula percompressa — Graf & Cummings (2007), Bogan (2010), Pfeiffer et al. (2019), Inoue et al. (2020)
Psorula profunda — Graf & Cummings (2007), Bogan (2010), Pfeiffer et al. (2019), Inoue et al. (2020), Freeman et al. (2025)
Psorula quadrata — Graf & Cummings (2007), Bogan (2010)
Psorula salinarum — Graf & Cummings (2007), Bogan (2010)
Psorula usumasintae — Graf & Cummings (2007), Bogan (2010), Pfeiffer et al. (2019), Inoue et al. (2020), Freeman et al. (2025)
Psoronaias psorica — Bogan (2010)
Psoronaias testudinarea [sic] — Bogan (2010)
“Psoronaias semigranosa” ** — Pfeiffer et al. (2019), Pfeiffer et al. (2019), Inoue et al. (2020), Smith et al. (2020), Freeman et al. (2025), Campbell & Lydeard (2026)
Psoronaias percompressa — Graf & Cummings (2021), Czaja et al. (2023), Campbell & Lydeard (2026)
Psoronaias profunda — Graf & Cummings (2021), Czaja et al. (2023), Campbell & Lydeard (2026)
Psoronaias quadrata — Graf & Cummings (2021)
Psoronaias salinarum — Graf & Cummings (2021)
Psoronaias usumasintae — Graf & Cummings (2021), Czaja et al. (2023), Campbell & Lydeard (2026)
Barynaias pigerrimus — Czaja et al. (2023)
Psorula percompressus — Freeman et al. (2025)

Range

Endemic to the Usumacinta basin, México and Guatemala (Central America: CA1)

Map of Georeferenced Specimens

specimens map of Psoronaias crocodilorum

MUSSELp Specimen Records

Type Specimens

image of lot USNM_128987
USNM_128987
Usumacinta River, Guatemala
holotype
Quadrula profunda Simpson, 1914

image of lot USNM_128996
USNM_128996
Usumacinta River, Guatemala
lectotype
Quadrula quadrata Simpson, 1914

image of lot USNM_84210
USNM_84210
Rio Taxtunilha, (tributary to the Rio de la Pasion), Guatemala
holotype
Quadrula rudis Simpson, 1900

image of lot SMF_3487
SMF_3487
Rio de las Salinas, N. Guatemala
holotype
Rotundaria salinarum Haas, 1929

image of lot BMNH_1965209
BMNH_1965209
Chiapas, Mexico
holotype
Unio corium Reeve, 1864

image of lot BMNH_1965161-162
BMNH_1965161-162
flumen Usumacinta, Guatemala
paralectotypes
Unio crocodilorum Morelet, 1849

image of lot BMNH_1893-2-4-2030
BMNH_1893-2-4-2030
River Usumacinta, Central America
lectotype
Unio crocodilorum Morelet, 1849

image of lot MNHN_MP_0023
MNHN_MP_0023
Rio Usumacinta, Guatemala
uncatalogued paralectotype
Unio crocodilorum Morelet, 1849

image of lot BMNH_93-2-4-2026
BMNH_93-2-4-2026
Rio Usumacinta, Guatemala
lectotype
Unio morini Morelet, 1851

image of lot BMNH_94-3-22-2
BMNH_94-3-22-2
flumen Usumacinta, Guatemala
lectotype
Unio ostreatus Morelet, 1849

image of lot MNHN_IM-2000-2102
MNHN_IM-2000-2102
Rio Usumacinta superior, Guatemala
paralectotypes
Unio ostreatus Morelet, 1849

image of lot ZMB_39463
ZMB_39463
Rio de las Salinas, Guatemala
lectotype
Unio percompressus Martens, 1887

image of lot ZMB_108849
ZMB_108849
Rio de las Salinas, Guatemala
paralectotypes
Unio percompressus Martens, 1887

image of lot MNHN_IM-2000-1761
MNHN_IM-2000-1761
Mexique
holotype
Unio pigerrimus Crosse & Fischer, 1894

image of lot BMNH_93-2-4-2033
BMNH_93-2-4-2033
Rio Usumacinta, Tabasco
lectotype
Unio psoricus Morelet, 1851

image of lot BMNH_94-3-22-1
BMNH_94-3-22-1
flumen Usumacinta, Guatemala
lectotype
Unio testudineus Morelet, 1849

image of lot ZMB_39466
ZMB_39466
Rio de las Salinas, Guatemala
syntypes
Psoronaias martensi Frierson, 1927

image of lot SMF_3475
SMF_3475
Rio Usumacinta, Guatemala
paralectotypes
Quadrula quadrata Simpson, 1914

image of lot BMNH_1908-12-15-4
BMNH_1908-12-15-4
River Usumacinta, Guatemala
paralectotype
Unio crocodilorum Morelet, 1849

image of lot MNHN_IM-2000-1667
MNHN_IM-2000-1667
Rio Usumacinta, Guatemala
paralectotype
Unio crocodilorum Morelet, 1849

image of lot BMNH_1893-2-4-2031
BMNH_1893-2-4-2031
River Usumacinta, Guatemala
holotype
Unio praestricta Fischer & Crosse, 1894

BMNH_93-2-4-2034. Rio Usumacinta, Tabasco — lectotype [as holotype] Unio psoricus Morelet, 1851

BMNH_1893-2-4-2032. River Usumacinta, Guatemala — holotype Unio semipustulata Fischer & Crosse, 1894

Other Specimens by Collection | show by country

14 Specimens from AMNH

image of lot AMNH_140555
AMNH_140555
1 league [3 miles] above mouth Rio Santa Amelia, Peten, Guatemala

image of lot AMNH_140570
AMNH_140570
Rio de la Pasion, Peten, Guatemala

image of lot AMNH_140571
AMNH_140571
Rio de la Pasion, Santa Amelia, Peten, Guatemala

image of lot AMNH_140558
AMNH_140558
Rio de la Pasion, Santa Amelia, Peten, Guatemala

image of lot AMNH_140557
AMNH_140557
Rio de la Pasion, Santa Amelia, Peten, Guatemala

image of lot AMNH_140561
AMNH_140561
Rio de la Pasion, Santa Amelia, Peten, Guatemala

image of lot AMNH_140560
AMNH_140560
Rio de la Pasion, Santa Amelia, Peten, Guatemala

image of lot AMNH_140559
AMNH_140559
Rio de la Pasion, Santa Amelia, Peten, Guatemala

image of lot AMNH_140556
AMNH_140556
Rio de la Pasion, Santa Amelia, Peten, Guatemala

image of lot AMNH_140562
AMNH_140562
Rio de la Pasion, Santa Amelia, Peten, Guatemala

image of lot AMNH_140564
AMNH_140564
Rio de la Pasion, Santa Amelia, Peten, Guatemala

image of lot AMNH_140565
AMNH_140565
Rio de la Pasion, Santa Amelia, Peten, Guatemala

image of lot AMNH_140566
AMNH_140566
Rio San Pedro, Mactun, 96 km below Paso Caballas, Peten, Guatemala

image of lot AMNH_140563
AMNH_140563
Rio San Pedro, Mactun, 96 km below Paso Caballas, Peten, Guatemala

1 Specimens from AMS

image of lot AMS_102111
AMS_102111
Rio Usumacinta, Emiliano Zapata, Tabasco, Mexico

6 Specimens from ANSP

image of lot ANSP_159973
ANSP_159973
Piedras Negras, Guatemala

image of lot ANSP_41665
ANSP_41665
Rio de la Pasion

image of lot ANSP_185944
ANSP_185944
Rio Pasion, between Sabol and Sayaxche, Guatemala

image of lot ANSP_125582
ANSP_125582
Rio Taxtunilha, (trib. to Rio de la Pasion), Guatemala

image of lot ANSP_138664
ANSP_138664
Hacienda Reforma on the San Pedro River, trib. of the Usumacinta above Balancan, Mexico

image of lot ANSP_62937
ANSP_62937
lower part of Rio Tulija, State Tabasco, Mexico

3 Specimens from BMNH

image of lot BMNH_93-2-4-2045
BMNH_93-2-4-2045
Rio Usumacinta, Tabasco Superieur, Mexico

image of lot BMNH_1907-10-28-181
BMNH_1907-10-28-181
Mexico

image of lot BMNH_196496
BMNH_196496
Mexico

1 Specimens from FMNH

image of lot FMNH_15001
FMNH_15001
[no locality available - Central America]

7 Specimens from INHS

image of lot INHS_24857
INHS_24857
Rio Petexbatum, Peten, Guatemala

image of lot INHS_24858
INHS_24858
Rio Usumacinta, Alter of Sacrifice, Peten, Guatemala

image of lot INHS_91079
INHS_91079
Rio Saguila, Gulf of Mexico - Grijalva - Usumacinta - Chacamax, SE Emiliano Zapata, off Rt. 10, fish camp, Tabasco, Mexico

image of lot INHS_91077
INHS_91077
Rio Saguila, Gulf of Mexico - Grijalva - Usumacinta - Chacamax, SE Emiliano Zapata, off Rt. 10, fish camp, Tabasco, Mexico

image of lot INHS_91078
INHS_91078
Rio Saguila, Gulf of Mexico - Grijalva - Usumacinta - Chacamax, SE Emiliano Zapata, off Rt. 10, fish camp, Tabasco, Mexico

image of lot INHS_24859
INHS_24859
Rio Usumacinta, Boca Lacantum, Chiapas, Mexico

image of lot INHS_24860
INHS_24860
Rio Usumacinta, Boca Lacantum, Chiapas, Mexico

14 Specimens from MCZ

image of lot MCZ_270070
MCZ_270070
3 mi above mouth Rio Santa Amelia, Peten, Guatemala

image of lot MCZ_270078
MCZ_270078
3 mi above mouth Rio Santa Amelia, Peten, Guatemala

image of lot MCZ_270071
MCZ_270071
3 mi above mouth Rio Santa Amelia, Peten, Guatemala

image of lot MCZ_270073
MCZ_270073
3 mi above mouth Rio Santa Amelia, Peten, Guatemala

image of lot MCZ_270074
MCZ_270074
3 mi above mouth Rio Santa Amelia, Peten, Guatemala

image of lot MCZ_270076
MCZ_270076
Rio de la Pasion, Peten, Guatemala

image of lot MCZ_270072
MCZ_270072
Rio de la Pasion, Santa Amelia, Peten, Guatemala

image of lot MCZ_270075
MCZ_270075
Rio de la Pasion, Treasure Island, Peten, Guatemala

image of lot MCZ_270077
MCZ_270077
Rio de la Pasion, Treasure Island, Peten, Guatemala

image of lot MCZ_293588
MCZ_293588
Usumacinta River, Yaxchilan, 106 mi downstream from Sayache, Mexico

image of lot MCZ_293589
MCZ_293589
Usumacinta River, Yaxchilan, 106 mi downstream from Sayache, Mexico

image of lot MCZ_72567
MCZ_72567
Balancan in Usumacinta R., Tabasco, Mexico

image of lot MCZ_288412
MCZ_288412
Rio Usumacinta, Emiliano Zapata, Tabasco, Mexico

MCZ_288418B. small river, 3 mi SE Emiliano Zapata, Tabasco, Mexico

2 Specimens from MNHN

image of lot MNHN_MP_1343
MNHN_MP_1343
Rio Usumacinta, Guatemala

image of lot MNHN_MP_1340
MNHN_MP_1340
Mexico

2 Specimens from OSUM

image of lot OSUM_18868
OSUM_18868
Rio de la Pasion, Peten, Guatemala

image of lot OSUM_18870
OSUM_18870
Usumacinta River, Machtun, 96 km below Paso Caballos [at or very near Mexican border]

9 Specimens from SMF

image of lot SMF_15025
SMF_15025
Cochinchine Jung ?

image of lot SMF_15027
SMF_15027
Rio Cancuen, Nebenfluss d. Rio Usumacinta, Guatemala

image of lot SMF_15030
SMF_15030
Rio de la Pasion, Guatemala (Paso Real)

image of lot SMF_15026
SMF_15026
Rio de la Passion, Guatemala

image of lot SMF_15023
SMF_15023
Rio de las Salinas, N. Guatemala

image of lot SMF_15024
SMF_15024
Rio Usumacinta, Guatemala

image of lot SMF_15028
SMF_15028
Rio Usumacinta, Guatemala

image of lot SMF_15029
SMF_15029
Rio Usumacinta, Guatemala

image of lot SMF_3476
SMF_3476
Rio Usumacinta, Guatemala

19 Specimens from UF

image of lot UF_507905
UF_507905
Rio De La Pasion, 5.5 miles east of Sayaxche, El Peten Department, Guatemala 16.536339, -90.103900

image of lot UF_507906
UF_507906
Rio De La Pasion, 5.5 miles east of Sayaxche, El Peten Department, Guatemala 16.536339, -90.103900

image of lot UF_507907
UF_507907
Rio De La Pasion, 5.5 miles east of Sayaxche, El Peten Department, Guatemala 16.536339, -90.103900

image of lot UF_507908
UF_507908
Rio De La Pasion, 5.5 miles east of Sayaxche, El Peten Department, Guatemala 16.536339, -90.103900

image of lot UF_507909
UF_507909
Rio De La Pasion, 5.5 miles east of Sayaxche, El Peten Department, Guatemala 16.536339, -90.103900

image of lot UF_270399
UF_270399
Chiapas, Mexico

image of lot UF_270520
UF_270520
Mexico

image of lot UF_164057
UF_164057
Rio San Pedro, 3 km N La Palma, Tabasco, Mexico.

image of lot UF_164058
UF_164058
Rio San Pedro, 3 km N La Palma, Tabasco, Mexico.

image of lot UF_164059
UF_164059
Rio San Pedro, 3 km N La Palma, Tabasco, Mexico.

image of lot UF_517941
UF_517941
Rio San Pedro, Bahia Campeche 3 km N of La Palma Tabasco, Mexico

image of lot UF_164065
UF_164065
Rio San Pedro, Cascadas de la Reforma, 7 km N and 7 km E El Aguila, Tabasco, Mexico

image of lot UF_164066
UF_164066
Rio San Pedro, Cascadas de la Reforma, 7 km N and 7 km E El Aguila, Tabasco, Mexico

image of lot UF_164067
UF_164067
Rio San Pedro, Limon, Tabasco, Mexico

image of lot UF_190664
UF_190664
Rio Usumacinta, Emiliano Zapata, Tabasco, Mexico

image of lot UF_378849
UF_378849
Rio Usumacinta, Emilliano Zapata, Tabasco, Mexico 17.7333, -91.8

image of lot UF_270396
UF_270396
Tabasco, Mexico

image of lot UF_507899
UF_507899
Usumacinta River, opposite of Emiliano Zapata. Tabasco, Mexico. 17.749685, -91.766995

UF_529129. Rio San Pedro, Bahia Campeche 3 km N of La Palma Tabasco, Mexico

19 Specimens from UMMZ

image of lot UMMZ_65556
UMMZ_65556
Arroyo Chocop, about 12 mi above Mactun, Dept. Peten, Guatemala

image of lot UMMZ_76912
UMMZ_76912
Guatemala

image of lot UMMZ_65549
UMMZ_65549
Pasion River, about 13 mi above Sayache [=Sayaxche], Dept. Peten, Guatemala

image of lot UMMZ_65551
UMMZ_65551
Pasion River, above mouth of Rio Santa Monica, Dept. Peten, Guatemala

image of lot UMMZ_65545
UMMZ_65545
Pasion River, at Cambio, Dept. Peten, Guatemala

image of lot UMMZ_65543
UMMZ_65543
Pasion River, at Embuido rapids, above Porvenir, Dept. Alta Vera Paz, Guatemala

image of lot UMMZ_65541
UMMZ_65541
Pasion River, at first arroyo below mouth of Rio San Simon, Dept. Alta Vera Paz, Guatemala

image of lot UMMZ_65552
UMMZ_65552
Pasion River, at mouth of Rio Martin, Dept. Peten, Guatemala

image of lot UMMZ_65546
UMMZ_65546
Pasion River, at road above Tres Islas, Dept. Peten, Guatemala

image of lot UMMZ_65542
UMMZ_65542
Pasion River, at rodal de Caresal, above Porvenir, Dept. Alta Vera Paz, Guatemala

image of lot UMMZ_65548
UMMZ_65548
Pasion River, at Santa Amelia, Dept. Peten, Guatemala

image of lot UMMZ_65550
UMMZ_65550
Pasion River, at Sayanche [=Sayaxche], Dept. Peten, Guatemala

image of lot UMMZ_65547
UMMZ_65547
Pasion River, at Tres Islas, Dept. Peten, Guatemala

image of lot UMMZ_65553
UMMZ_65553
Pasion River, below Sayarche [=Sayaxche], Dept. Peten, Guatemala

image of lot UMMZ_65544
UMMZ_65544
Pasion River, Rodal “Grande”, Alta Vera Paz, Guatemala

image of lot UMMZ_65558
UMMZ_65558
San Pedro River, 4-6 mi E Paso Caballo, Dept. Peten, Guatemala

image of lot UMMZ_65557
UMMZ_65557
San Pedro River, Mactun, Dept. Peten, Guatemala

image of lot UMMZ_65554
UMMZ_65554
Subin River, about 2 mi above mouth, Dept. Peten, Guatemala

image of lot UMMZ_65555
UMMZ_65555
Subin River, Santa Teresa, Dept. Peten, Guatemala

28 Specimens from USNM

image of lot USNM_783055
USNM_783055
Rio de la Pasion, Santa Amelia, Guatemala

image of lot USNM_783058
USNM_783058
Rio de la Pasion, Santa Amelia, Guatemala

image of lot USNM_783059
USNM_783059
Rio de la Pasion, Santa Amelia, Guatemala

image of lot USNM_783061
USNM_783061
Rio de la Pasion, Santa Amelia, Guatemala

image of lot USNM_783062
USNM_783062
Rio de la Pasion, Santa Amelia, Guatemala

image of lot USNM_783063
USNM_783063
Rio de la Pasion, Santa Amelia, Guatemala

image of lot USNM_783047
USNM_783047
Rio de la Pasion, Santa Amelia, Peten, Guatemala

image of lot USNM_783048
USNM_783048
Rio de la Pasion, Santa Amelia, Peten, Guatemala

image of lot USNM_783049
USNM_783049
Rio de la Pasion, Santa Amelia, Peten, Guatemala

image of lot USNM_783050
USNM_783050
Rio de la Pasion, Santa Amelia, Peten, Guatemala

image of lot USNM_783051
USNM_783051
Rio de la Pasion, Santa Amelia, Peten, Guatemala

image of lot USNM_783057
USNM_783057
Rio de la Pasion, Santa Amelia, Peten, Guatemala

image of lot USNM_783046
USNM_783046
Rio San Pedro, Mactun, 96 km below Paso Caballas, Peten, Guatemala

image of lot USNM_795111
USNM_795111
Rio San Pedro, Mactun, 96 km below Paso Caballas, Peten, Guatemala

image of lot USNM_128984
USNM_128984
Rio Usumacinta, Guatemala

image of lot USNM_128988
USNM_128988
Rio Usumacinta, Guatemala

image of lot USNM_347187
USNM_347187
Rio Usumacinta, Tabasco, Mexico

image of lot USNM_792459
USNM_792459
Rio Usumasinta, Guatemala

image of lot USNM_128992_01
USNM_128992_01
Usumacinta R., Guatemala

image of lot USNM_84335
USNM_84335
Mexico

image of lot USNM_161475
USNM_161475
Monte Cristo, Tabasco, Mexico

image of lot USNM_359337
USNM_359337
near San Cristobal, Mexico

image of lot USNM_795617
USNM_795617
pond W of Rio Grijalva, about 3 km above Villahermosa, Tabasco, Mexico

image of lot USNM_675261
USNM_675261
Rio Usumacinta, 9.4 mi NNW Emillano Zapata, Mexico

image of lot USNM_267924
USNM_267924
Rio Usumacinta, 9.4 mi NNW Emillano Zapata, Tabasco, Mexico

image of lot USNM_251192
USNM_251192
Rio Usumacinta, near Jonuta, Tabasco, Mexico

image of lot USNM_675260
USNM_675260
Rio Usumacinta, Tabasco, Emillano Zapata, Mexico

image of lot USNM_795616
USNM_795616
trib. of Rio Grijalva, 1 km above Villa Hermosa, Tabasco, Mexico

4 Specimens from ZMB

image of lot ZMB_MP_0486
ZMB_MP_0486
Mexico

image of lot ZMB_MP_0472
ZMB_MP_0472
[no locality available - Central America]

image of lot ZMB_MP_0490
ZMB_MP_0490
Amer Mer.

image of lot ZMB_22255
ZMB_22255
Central America [locality illegible]

4 Specimens from other collections

COMA_2048. Rio Usumacinta, Tabasco, Mexico

COMA_B.1053. Río Candelaria, Campeche, México

COMA_B.0392. Rio González, Laguna del Horizonte, Tabasco, México

COMA_B.0720. Río Usumacinta, Tabasco, México

GenBank Vouchers

Georeferenced GenBank Vouchers

Mexico

UF_507899. (SRA: SRX5278589, go to GenBank) Mexico, Tabasco, Usumacinta River, opposite Emiliano Zapata (17.749685, -91.766995) .

UF_507899 (2015-0722). (16S: MK001775, 28S: MK001809, COI: MK001758, go to GenBank) ditto.

UF_507899 (2015-0726). (16S: MK001776, 28S: MK001810, COI: MK001759, go to GenBank) ditto.

GenBank Vouchers Not Yet Georeferenced

Z_12430 (2015-0509). (16S: MK001783, 28S: MK001817, COI: MK001766, go to GenBank)

Z_12441 (2015-0520). (16S: MK001781, 28S: MK001815, COI: MK001764, go to GenBank)

Z_12451 (2015-0531). (16S: MK001782, 28S: MK001816, COI: MK001765, go to GenBank)

Z_12460 (2015-0540). (16S: MK001780, 28S: MK001814, COI: MK001763, go to GenBank)

Z_12469 (2015-0551). (16S: MK001779, 28S: MK001813, COI: MK001762, go to GenBank)

Z_12471 (2015-0555). (16S: MK001784, 28S: MK001818, COI: MK001767, go to GenBank)