Bivalvia | Unionida | Unionoidea | Unionidae | Parreysiinae | Indochinellini

genus Scabies Haas, 1911: cladomics

8 Recent valid species (8 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 8 (8 total)
analyzed in 51 cladograms in 30 publications
maximum number of terminals in an analysis = 8 (mean = 3.94, median = 3)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 30
maximum number of included terminals = 8, (mean = 6 median = 8)
proportion of cladograms that support basic monophyly = 0.93

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Scabies cladogram distances
PCoA Ordination

NJ Tree of Scabies cladogram distances
NJ Tree

Cladograms

Bolotov et al., 2024 [Publ 4369], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cst iT tT bi riS oT roT roS iD
1.000 8 8 6 8 54 21 7 **
Cladomics cladogram Pseudoscabies Indonaia Indochinella Radiatula Scabiellus Harmandia Unionetta Scabies crispata Scabies mandarinus Scabies nucleus Scabies humilis Scabies pilata Scabies phaselus Scabies anceps Scabies scobinatus

Bolotov et al., 2024 [Publ 4369], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 8 8 6 8 55 21 7 6.0
Cladomics cladogram Pseudoscabies Indonaia Indochinella Radiatula Scabiellus Harmandia Unionetta Scabies crispata Scabies mandarinus Scabies nucleus Scabies pilata Scabies humilis Scabies anceps Scabies phaselus Scabies scobinatus

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cst iT tT bi riS oT roT roS iD
0.889 8 8 6 8 625 21 7 10.0
Cladomics cladogram Pseudoscabies Indonaia Indochinella Radiatula Scabiellus Unionetta fabagina 2 Harmandia Unionetta fabagina 1 Scabies crispata Scabies nucleus Scabies mandarinus Scabies pilata Scabies humilis Scabies anceps Scabies scobinatus Scabies phaselus

Konopleva et al., 2026 [Publ 4847], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 8 8 5 8 58 23 7 5.0
Cladomics cladogram Pseudoscabies Indonaia Indochinella Radiatula Scabiellus Unionetta Harmandia Scabies mandarinus Scabies crispata Scabies nucleus Scabies pilata Scabies humilis Scabies anceps Scabies phaselus Scabies scobinatus

Konopleva et al., 2026 [Publ 4847], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 8 8 6 8 58 23 7 0
Cladomics cladogram Pseudoscabies Indonaia Indochinella Radiatula Scabiellus Harmandia Unionetta Scabies crispata Scabies mandarinus Scabies nucleus Scabies humilis Scabies pilata Scabies phaselus Scabies anceps Scabies scobinatus

Konopleva et al., 2026 [Publ 4847], fig. S2

cst iT tT bi riS oT roT roS iD
1.000 8 8 6 8 59 23 7 6.0
Cladomics cladogram Pseudoscabies Indonaia Indochinella Radiatula Scabiellus Harmandia Unionetta Scabies crispata Scabies mandarinus Scabies nucleus Scabies pilata Scabies humilis Scabies anceps Scabies phaselus Scabies scobinatus

Konopleva et al., 2026 [Publ 4847], fig. S3

cst iT tT bi riS oT roT roS iD
1.000 8 8 4 8 59 23 7 6.0
Cladomics cladogram Pseudoscabies Indonaia Indochinella Radiatula Scabiellus Harmandia Unionetta Scabies crispata Scabies mandarinus Scabies humilis Scabies nucleus Scabies pilata Scabies anceps Scabies phaselus Scabies scobinatus

Pfeiffer et al., 2018 [Publ 3827], fig. 2

cst iT tT bi riS oT roT roS iD
0.875 8 8 5 8 20 9 6 1.0
Cladomics cladogram Indonaia Indochinella Radiatula Scabiellus Unionetta fabagina 2 Harmandia Unionetta fabagina 1 Scabies crispata Scabies mandarinus Scabies nucleus Scabies pilata Scabies humilis Scabies phaselus Scabies anceps Scabies scobinatus

Jeratthitikul et al., 2019 [Publ 3922], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 8 8 6 8 15 10 6 8.0
Cladomics cladogram Indonaia Indochinella Radiatula Scabiellus Harmandia Unionetta Scabies crispata Scabies mandarinus Scabies pilata Scabies nucleus Scabies humilis Scabies scobinatus Scabies phaselus Scabies anceps

Bolotov et al., 2020 [Publ 4100], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 8 8 6 8 109 13 6 4.0
Cladomics cladogram Indonaia Indochinella Radiatula Scabiellus Unionetta Harmandia Scabies nucleus Scabies crispata Scabies mandarinus Scabies humilis Scabies pilata Scabies anceps Scabies phaselus Scabies scobinatus

Bolotov et al., 2020 [Publ 4100], fig. S1

cst iT tT bi riS oT roT roS iD
0.875 8 8 5 8 103 12 6 3.0
Cladomics cladogram Indonaia Indochinella Radiatula mouhoti Radiatula myitkyinae Radiatula chaudhurii Radiatula crispisulcata Scabiellus Harmandia Unionetta Scabies mandarinus Scabies crispata Scabies nucleus Scabies humilis Scabies pilata Scabies phaselus Scabies scobinatus Scabies anceps

Bolotov et al., 2022 [Publ 4239], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 8 8 6 8 50 18 6 6.0
Cladomics cladogram Indonaia Indochinella Radiatula Scabiellus Harmandia Unionetta Scabies crispata Scabies mandarinus Scabies nucleus Scabies pilata Scabies humilis Scabies anceps Scabies phaselus Scabies scobinatus

Bolotov et al., 2022 [Publ 4239], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 8 8 6 8 50 18 6 0
Cladomics cladogram Indonaia Indochinella Radiatula Scabiellus Harmandia Unionetta Scabies crispata Scabies mandarinus Scabies nucleus Scabies humilis Scabies pilata Scabies phaselus Scabies anceps Scabies scobinatus

Bolotov et al., 2022 [Publ 4239], fig. S2

cst iT tT bi riS oT roT roS iD
1.000 8 8 6 8 50 18 6 10.0
Cladomics cladogram Indonaia Indochinella Radiatula Scabiellus Harmandia Unionetta Scabies crispata Scabies humilis Scabies mandarinus Scabies nucleus Scabies pilata Scabies anceps Scabies phaselus Scabies scobinatus

Bolotov et al., 2022 [Publ 4252], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 8 8 6 8 53 19 6 0
Cladomics cladogram Indonaia Indochinella Radiatula Scabiellus Harmandia Unionetta Scabies crispata Scabies mandarinus Scabies nucleus Scabies humilis Scabies pilata Scabies phaselus Scabies anceps Scabies scobinatus

Bolotov et al., 2022 [Publ 4252], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 8 8 6 8 53 19 6 10.0
Cladomics cladogram Indonaia Indochinella Radiatula Scabiellus Harmandia Unionetta Scabies crispata Scabies humilis Scabies mandarinus Scabies nucleus Scabies pilata Scabies anceps Scabies phaselus Scabies scobinatus

Konopleva et al., 2023 [Publ 4262], fig. 8

cst iT tT bi riS oT roT roS iD
1.000 8 8 4 8 74 6 6 6.0
Cladomics cladogram Indonaia Indochinella Radiatula Scabiellus Unionetta Harmandia Scabies humilis Scabies pilata Scabies crispata Scabies nucleus Scabies mandarinus Scabies phaselus Scabies anceps Scabies scobinatus

Chen et al., 2026 [Publ 4833], fig. 1

cst iT tT bi riS oT roT roS iD
0.750 8 8 6 8 19 6 6 8.0
Cladomics cladogram Harmandia Chenunionetta Unionetta fabagina 2 Unionetta fabagina 1 Scabies crispata Scabies mandarinus Scabies humilis Scabies pilata Scabies nucleus Scabies scobinatus Scabies anceps Scabies phaselus

Bolotov et al., 2017 [Publ 3761], fig. S1

cst iT tT bi riS oT roT roS iD
0.800 4 7 5 4 135 7 3 9.0
Cladomics cladogram Indonaia Indochinella Scabies crispata Radiatula Scabies mandarinus Scabies humilis Scabies phaselus

Bolotov et al., 2017 [Publ 3761], fig. S2

cst iT tT bi riS oT roT roS iD
0.800 4 7 5 4 135 7 3 9.0
Cladomics cladogram Indonaia Indochinella Scabies crispata Radiatula Scabies mandarinus Scabies humilis Scabies phaselus

Bolotov et al., 2017 [Publ 3764], fig. S1

cst iT tT bi riS oT roT roS iD
0.600 3 3 0 3 122 9 3 6.0
Cladomics cladogram Indochinella Radiatula Scabies humilis Scabies phaselus Scabies mandarinus

Bolotov et al., 2018 [Publ 3826], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 79 8 3 5.0
Cladomics cladogram Indonaia Indochinella Radiatula Scabies mandarinus Scabies humilis Scabies phaselus

Huang et al., 2019 [Publ 3843], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 198 9 3 7.0
Cladomics cladogram Indonaia Indochinella Radiatula Scabies humilis Scabies phaselus Scabies mandarinus

Huang et al., 2019 [Publ 3843], fig. S3

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 196 9 3 7.0
Cladomics cladogram Indonaia Indochinella Radiatula Scabies phaselus Scabies humilis Scabies mandarinus

Huang et al., 2019 [Publ 3843], fig. S4

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 198 9 3 7.0
Cladomics cladogram Indonaia Indochinella Radiatula Scabies humilis Scabies phaselus Scabies mandarinus

Konopleva et al., 2019 [Publ 3900], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 0 3 48 9 3 6.0
Cladomics cladogram Indonaia Indochinella Radiatula Scabies humilis Scabies mandarinus Scabies phaselus

Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 1

cst iT tT bi riS oT roT roS iD
0.800 3 3 0 3 33 5 3 6.0
Cladomics cladogram Indonaia Indochinella Radiatula mouhoti Radiatula myitkyinae Radiatula chaudhurii Scabies humilis Scabies phaselus Scabies mandarinus

Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 2

cst iT tT bi riS oT roT roS iD
0.800 3 3 0 3 33 5 3 6.0
Cladomics cladogram Radiatula mouhoti Radiatula myitkyinae Radiatula chaudhurii Indochinella Indonaia Scabies humilis Scabies phaselus Scabies mandarinus

Dai et al., 2024 [Publ 4386], fig. 5-BI

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 20 0 0 --
Cladomics cladogram Trapezidens Scabies scobinatus Scabies crispata

Dai et al., 2024 [Publ 4386], fig. 5-ML

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 20 0 0 --
Cladomics cladogram Trapezidens Scabies scobinatus Scabies crispata

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 56 3 3 --
Cladomics cladogram Indonaia Indochinella Scabies phaselus Harmandia

Pfeiffer et al., 2019 [Publ 3937], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 56 3 3 --
Cladomics cladogram Indonaia Indochinella Scabies phaselus Harmandia

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 39 2 2 --
Cladomics cladogram Indochinella Radiatula Scabies mandarinus

Konopleva et al., 2017 [Publ 3807], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 2 2 --
Cladomics cladogram Indochinella Radiatula Scabies mandarinus

Wu et al., 2021 [Publ 4366], fig. 2_I

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 4 2 --
Cladomics cladogram Indonaia Scabies crispata Radiatula

Wu et al., 2021 [Publ 4366], fig. 6

cst iT tT bi riS oT roT roS iD
0.750 1 1 0 1 90 4 2 --
Cladomics cladogram Indonaia Radiatula myitkyinae Scabies crispata Radiatula mouhoti Radiatula chaudhurii

Combosch et al., 2017 [Publ 3806], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 212 0 0 --
Cladomics cladogram Lampsilis Scabies crispata

Combosch et al., 2017 [Publ 3806], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 213 0 0 --
Cladomics cladogram Scabies crispata Lampsilis

Huang et al., 2018 [Publ 3828], fig. 3_5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 28 0 0 --
Cladomics cladogram Lampsilis Scabies crispata

Jeratthitikul et al., 2019 [Publ 3911], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 8 0 0 --
Cladomics cladogram Pilsbryoconcha Scabies crispata

Wu et al., 2020 [Publ 4121], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 33 0 0 --
Cladomics cladogram Lampsilis Scabies crispata

Wu et al., 2020 [Publ 4121], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 33 0 0 --
Cladomics cladogram Lampsilis Scabies crispata

Sano et al., 2020 [Publ 4138], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 68 0 0 --
Cladomics cladogram Scabies crispata Lampsilis Sinohyriopsis Inversidens Ptychorhynchus Obovalis Lamprotula Pronodularia Potomida Anemina Buldowskia Pletholophus Beringiana Sinanodonta Cristaria Anodonta Pyganodon Utterbackia Alasmidonta Prolasmidonta Platynaias Lanceolaria Lepidodesma Scriptolamprotula Unio Inversiunio Nodularia Tchangsinaia Schistodesmus

Sano et al., 2020 [Publ 4138], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 68 0 0 --
Cladomics cladogram Scabies crispata Sinohyriopsis Lampsilis Lamprotula Potomida Pronodularia Inversidens Obovalis Ptychorhynchus Anemina Buldowskia Sinanodonta Pletholophus Beringiana Cristaria Anodonta Pyganodon Utterbackia Alasmidonta Platynaias Prolasmidonta Lanceolaria Lepidodesma Scriptolamprotula Unio Inversiunio Nodularia Schistodesmus Tchangsinaia

Sano et al., 2020 [Publ 4138], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 68 0 0 --
Cladomics cladogram Scabies crispata Lampsilis Sinohyriopsis Inversidens Obovalis Ptychorhynchus Lamprotula Pronodularia Potomida Anemina Buldowskia Sinanodonta Pletholophus Beringiana Cristaria Anodonta Pyganodon Utterbackia Alasmidonta Prolasmidonta Platynaias Lepidodesma Lanceolaria Scriptolamprotula Unio Nodularia Inversiunio Schistodesmus Tchangsinaia

Smith, 2021 [Publ 4331], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 19 0 0 --
Cladomics cladogram Scabies phaselus Cristaria Pyganodon Utterbackia

Dai et al., 2024 [Publ 4385], fig. 3-BI

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 0 0 --
Cladomics cladogram Trapezidens Scabies crispata

Dai et al., 2024 [Publ 4385], fig. 3-ML

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 0 0 --
Cladomics cladogram Trapezidens Scabies crispata

Dai et al., 2024 [Publ 4386], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 0 0 --
Cladomics cladogram Scabies scobinatus Unio Scriptolamprotula Anodonta Lepidodesma Middendorffinaia Inversiunio Cuneopsis Nodularia

Xiang et al., 2025 [Publ 4433], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 27 0 0 --
Cladomics cladogram Scabies crispata Pletholophus Beringiana Mimo Sinanodonta Cristaria Anemina Amuranodonta Buldowskia

Dai et al., 2026 [Publ 4757], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 33 0 0 --
Cladomics cladogram Scabies crispata Trapezidens