Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Anodontini | Alasmidontina

genus Utterbackiana Frierson, 1927: cladomics

5 Recent valid species (5 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 5 (5 total)
analyzed in 22 cladograms in 14 publications
maximum number of terminals in an analysis = 5 (mean = 2.5, median = 2.5)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 13
maximum number of included terminals = 5, (mean = 3.54 median = 4)
proportion of cladograms that support basic monophyly = 0.92

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Utterbackiana cladogram distances
PCoA Ordination

NJ Tree of Utterbackiana cladogram distances
NJ Tree

Cladograms

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
5 5 3 5 626 31 13 0 
        +-------- Utterbackiana_implicata **    
        |                                    
  +-----+     +-- Utterbackiana_couperiana **   
  |     |  +--+                              
  |     |  |  +-- Utterbackiana_heardi **       
  |     +--+                                 
  |        |  +-- Utterbackiana_hartfieldorum **
--+        +--+                              
  |           +-- Utterbackiana_suborbiculata **
  |                                          
  |  +----------- Simpsonaias                
  |  |                                       
  |  |        +-- Alasminota                 
  +--+  +-----+                              
     |  |     +-- Alasmidonta                
     |  |                                    
     +--+     +-- Prolasmidonta              
        |  +--+                              
        |  |  +-- Platynaias                 
        +--+                                 
           |  +-- Anodontoides               
           +--+                              
              +-- Strophitus

Perkins et al., 2025 [Publ 4447], fig. 2_mtDNA-ITS

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
4 4 2 4 32 29 14 1 
                 +-- Utterbackiana_heardi **       
              +--+                              
              |  +-- Utterbackiana_couperiana **   
     +--------+                                 
     |        |  +-- Utterbackiana_hartfieldorum **
     |        +--+                              
  +--+           +-- Utterbackiana_suborbiculata **
  |  |                                          
  |  |  +----------- Alasminota                 
  |  +--+                                       
  |     |  +-------- Pseudodontoideus           
  |     +--+                                    
  |        |  +----- Lasmigona                  
  |        +--+                                 
  |           |  +-- Pterosyna                  
--+           +--+                              
  |              +-- Arcidens                   
  |                                             
  |              +-- Prolasmidonta              
  |           +--+                              
  |           |  +-- Platynaias                 
  |     +-----+                                 
  |     |     |  +-- Ligodonta                  
  |     |     +--+                              
  |     |        +-- Alasmidonta                
  +-----+                                       
        |        +-- Anodontoides               
        |  +-----+                              
        |  |     +-- Strophitus                 
        +--+                                    
           |  +----- Sulcularia                 
           +--+                                 
              |  +-- Pyganodon                  
              +--+                              
                 +-- Utterbackia

Whelan et al., 2023 [Publ 4287], fig. 2

iTtTbiriSoTroTroSiD
4 4 2 4 37 29 13 1 
              +-- Utterbackia                
  +-----------+                              
  |           +-- Pyganodon                  
  |                                          
  |        +----- Sulcularia                 
  |     +--+                                 
  |     |  |  +-- Anodontoides               
  |     |  +--+                              
  |     |     +-- Strophitus                 
--+  +--+                                    
  |  |  |     +-- Prolasmidonta              
  |  |  |  +--+                              
  |  |  |  |  +-- Platynaias                 
  |  |  +--+                                 
  |  |     |  +-- Alasminota                 
  |  |     +--+                              
  |  |        +-- Alasmidonta                
  +--+                                       
     |        +-- Utterbackiana_couperiana **   
     |     +--+                              
     |     |  +-- Utterbackiana_heardi **       
     |  +--+                                 
     |  |  |  +-- Utterbackiana_suborbiculata **
     |  |  +--+                              
     |  |     +-- Utterbackiana_hartfieldorum **
     +--+                                    
        |     +-- Lasmigona                  
        |  +--+                              
        |  |  +-- Pseudodontoideus           
        +--+                                 
           |  +-- Pterosyna                  
           +--+                              
              +-- Arcidens

Smith et al., 2018 [Publ 3790], fig. 1

iTtTbiriSoTroTroSiD
4 4 2 4 14 13 7 1 
                 +-- Utterbackia                
  +--------------+                              
  |              +-- Pyganodon                  
  |                                             
--+              +-- Anodontoides               
  |  +-----------+                              
  |  |           +-- Strophitus                 
  |  |                                          
  +--+           +-- Alasmidonta                
     |  +--------+                              
     |  |        +-- Alasminota                 
     +--+                                       
        |  +-------- Pseudodontoideus           
        |  |                                    
        +--+     +-- Utterbackiana_couperiana **   
           |  +--+                              
           |  |  +-- Utterbackiana_heardi **       
           +--+                                 
              |  +-- Utterbackiana_hartfieldorum **
              +--+                              
                 +-- Utterbackiana_suborbiculata **

Zanatta et al., 2007 [Publ 2698], fig. 1A

iTtTbiriSoTroTroSiD
4 4 2 4 12 10 5 1 
  +-------- Utterbackiana_implicata **    
--+                                    
  |  +----- Utterbackiana_suborbiculata **
  +--+                                 
     |  +-- Utterbackiana_heardi **       
     +--+                              
        +-- Utterbackiana_couperiana **

Whelan et al., 2023 [Publ 4287], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 4 24 23 11 1 
                    +-- Utterbackiana_heardi **       
                 +--+                              
                 |  +-- Utterbackiana_couperiana **   
  +--------------+                                 
  |              |  +-- Utterbackiana_hartfieldorum **
  |              +--+                              
  |                 +-- Utterbackiana_suborbiculata **
--+                                                
  |                 +-- Utterbackia                
  |  +--------------+                              
  |  |              +-- Pyganodon                  
  |  |                                             
  +--+           +----- Alasmidonta                
     |  +--------+                                 
     |  |        |  +-- Arcidens                   
     |  |        +--+                              
     +--+           +-- Pseudodontoideus           
        |                                          
        |  +----------- Alasminota                 
        |  |                                       
        +--+        +-- Anodontoides               
           |  +-----+                              
           |  |     +-- Strophitus                 
           +--+                                    
              |  +----- Sulcularia                 
              +--+                                 
                 |  +-- Platynaias                 
                 +--+                              
                    +-- Prolasmidonta

Zanatta et al., 2007 [Publ 2698], fig. 1B

iTtTbiriSoTroTroSiD
4 4 2 4 12 10 5 3 
  +-------- Utterbackiana_implicata **    
--+                                    
  |  +----- Utterbackiana_heardi **       
  +--+                                 
     |  +-- Utterbackiana_suborbiculata **
     +--+                              
        +-- Utterbackiana_couperiana **

Whelan et al., 2023 [Publ 4287], fig. 4

iTtTbiriSoTroTroSiD
4 4 1 4 23 21 10 4 
  +-------------------- Sulcularia                 
  |                                                
  |     +-------------- Alasmidonta                
  |  +--+                                          
  |  |  |  +----------- Platynaias                 
--+  |  +--+                                       
  |  |     |  +-------- Strophitus                 
  |  |     +--+                                    
  |  |        |  +----- Pyganodon                  
  |  |        +--+                                 
  +--+           |  +-- Utterbackia                
     |           +--+                              
     |              +-- Anodontoides               
     |                                             
     |           +----- Alasminota                 
     |        +--+                                 
     |        |  |  +-- Arcidens                   
     |        |  +--+                              
     +--------+     +-- Pseudodontoideus           
              |                                    
              |  +----- Utterbackiana_couperiana **   
              |  |                                 
              +--+  +-- Utterbackiana_heardi **       
                 |  |                              
                 +--+-- Utterbackiana_hartfieldorum **
                    |                              
                    +-- Utterbackiana_suborbiculata **

Hoeh, 1990 [Publ 2717], fig. 4

iTtTbiriSoTroTroSiD
3 3 2 4 12 10 5 3 
  +-------- Utterbackiana_implicata **    
--+                                    
  |  +----- Utterbackiana_couperiana__2 **
  +--+                                 
     |  +-- Utterbackiana_suborbiculata **
     +--+                              
        +-- Utterbackiana_couperiana__1 **

Hoeh, 1990 [Publ 2717], fig. 2

iTtTbiriSoTroTroSiD
3 3 2 4 12 10 5 3 
  +-------- Utterbackiana_implicata **    
--+                                    
  |  +----- Utterbackiana_couperiana__2 **
  +--+                                 
     |  +-- Utterbackiana_suborbiculata **
     +--+                              
        +-- Utterbackiana_couperiana__1 **

Hoeh, 1990 [Publ 2717], fig. 3

iTtTbiriSoTroTroSiD
3 15 2 4 12 10 5 16 
  +----- Pyganodon_fragilis         
  |                                 
  +----- Utterbackia_imbecillis     
  |                                 
  +----- Utterbackia_peggyae        
  |                                 
  +----- Anodonta                   
  |                                 
  +----- Brachyanodon               
  |                                 
  +----- Utterbackiana_couperiana__1 **
  |                                 
--+----- Utterbackiana_couperiana__2 **
  |                                 
  +----- Utterbackiana_suborbiculata **
  |                                 
  |  +-- Lasmigona                  
  +--+                              
  |  +-- Utterbackiana_implicata **    
  |                                 
  |  +-- Pterosyna                  
  |  |                              
  |  +-- Platynaias                 
  |  |                              
  |  +-- Pyganodon_cataracta        
  +--+                              
     +-- Pyganodon_gibbosa          
     |                              
     +-- Pyganodon_grandis          
     |                              
     +-- Pyganodon_lacustris

Galal-Khallaf et al., 2024 [Publ 4459], fig. 6

iTtTbiriSoTroTroSiD
2 2 0 2 26 0 0 -- 
  +-- Utterbackiana_couperiana **   
--+                              
  +-- Utterbackiana_hartfieldorum **

Lydeard et al., 1996 [Publ 2656], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 25 4 4 -- 
     +----- Utterbackia             
  +--+                              
  |  |  +-- Utterbackiana_couperiana **
  |  +--+                           
--+     +-- Utterbackiana **           
  |                                 
  |  +----- Strophitus              
  +--+                              
     |  +-- Lasmigona               
     +--+                           
        +-- Alasmidonta

Pfeiffer et al., 2016 [Publ 3660], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 37 0 0 -- 
-- Utterbackiana_suborbiculata **

Turner et al., 2000 [Publ 2722], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 13 0 0 -- 
-- Utterbackiana_couperiana **

Curole & Kocher, 2002 [Publ 2712], fig. 1a-M

iTtTbiriSoTroTroSiD
1 1 0 1 3 0 0 -- 
-- Utterbackiana_implicata **

Curole & Kocher, 2005 [Publ 2682], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 11 1 1 -- 
  +-- Utterbackiana_implicata **
--+                          
  +-- Pyganodon

Huang et al., 2002 [Publ 2657], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 33 4 4 -- 
  +----------- Utterbackiana_couperiana **
--+                                    
  |  +-------- Utterbackia             
  +--+                                 
     |  +----- Strophitus              
     +--+                              
        |  +-- Alasmidonta             
        +--+                           
           +-- Lasmigona

Krebs et al., 2003 [Publ 2691], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 32 8 7 -- 
-- Utterbackiana_couperiana **

Huang et al., 2002 [Publ 2657], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 33 4 4 -- 
  +----- Utterbackia             
  |                              
--+----- Utterbackiana_couperiana **
  |                              
  |  +-- Alasmidonta             
  +--+                           
     +-- Lasmigona

Lydeard et al., 1996 [Publ 2656], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 27 4 4 -- 
-- Utterbackiana_couperiana **

Curole & Kocher, 2002 [Publ 2712], fig. 1b-M

iTtTbiriSoTroTroSiD
1 1 0 1 3 0 0 -- 
-- Utterbackiana_implicata **