Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Anodontini | Anodontina

genus Brachyanodon Crosse & Fischer, 1894: cladomics

5 Recent valid species (5 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 5 (5 total)
analyzed in 37 cladograms in 24 publications
maximum number of terminals in an analysis = 5 (mean = 1.38, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 6
maximum number of included terminals = 5, (mean = 3.33 median = 3.5)
proportion of cladograms that support basic monophyly = 0.33

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Brachyanodon cladogram distances
PCoA Ordination

NJ Tree of Brachyanodon cladogram distances
NJ Tree

Cladograms

Chong et al., 2008 [Publ 2668], fig. 3

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
4 18 10 7 14 3 2 0 
  +----------------- Anodonta_cygnea            
  |                                             
  +----------------- Pyganodon                  
  |                                             
  +----------------- Alasminota                 
  |                                             
  +----------------- Anodonta_anatina           
  |                                             
  +----------------- Unio                       
  |                                             
  |              +-- Brachyanodon_nuttalliana **   
  +--------------+                              
  |              +-- Brachyanodon_californiensis **
  |                                             
--+              +-- Beringiana                 
  +--------------+                              
  |              +-- Sinanodonta                
  |                                             
  |              +-- Pseudodontoideus           
  |              |                              
  +--------------+-- Alasmidonta                
  |              |                              
  |              +-- Pterosyna                  
  |                                             
  |  +-------------- Pseudanodonta              
  |  |                                          
  |  |        +----- Strophitus                 
  |  +--------+                                 
  +--+        |  +-- Prolasmidonta              
     |        +--+                              
     |           +-- Platynaias                 
     |                                          
     |  +----------- Brachyanodon_oregonensis__4 **
     +--+                                       
        |  +-------- Brachyanodon_oregonensis__3 **
        +--+                                    
           |  +----- Brachyanodon_oregonensis__2 **
           +--+                                 
              |  +-- Brachyanodon_oregonensis__1 **
              +--+                              
                 +-- Brachyanodon_kennerlyi **

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
5 8 7 7 627 5 2 23 
  +----------------------- Anodonta_anatina           
  |                                                   
--+  +-------------------- Pseudanodonta              
  |  |                                                
  +--+  +----------------- Brachyanodon_oregonensis__2 **
     |  |                                             
     |  |           +----- Anodonta_cygnea            
     +--+  +--------+                                 
        |  |        |  +-- Anodonta_seddonae          
        |  |        +--+                              
        +--+           +-- Anodonta_exulcerata        
           |                                          
           |  +----------- Brachyanodon_impura **        
           |  |                                       
           +--+  +-------- Brachyanodon **               
              |  |                                    
              +--+  +----- Brachyanodon_nuttalliana__1 **
                 |  |                                 
                 +--+----- Brachyanodon_nuttalliana__2 **
                    |                                 
                    |  +-- Brachyanodon_californiensis **
                    +--+                              
                       +-- Brachyanodon_oregonensis__1 **

Galal-Khallaf et al., 2024 [Publ 4459], fig. 6

iTtTbiriSoTroTroSiD
4 7 5 4 24 3 1 20 
           +-- Anodonta_exulcerata        
  +--------+                              
  |        +-- Anodonta_cygnea            
  |                                       
--+        +-- Brachyanodon_californiensis **
  |  +-----+                              
  |  |     +-- Brachyanodon_nuttalliana **   
  |  |                                    
  +--+     +-- Brachyanodon_kennerlyi **     
     |  +--+                              
     |  |  +-- Brachyanodon_oregonensis **   
     +--+                                 
        |  +-- Anodonta_anatina           
        +--+                              
           +-- Utterbackiana

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

iTtTbiriSoTroTroSiD
3 20 0 3 102 0 0 23 
  +-- Alasmidonta_marginata      
  |                              
  +-- Prolasmidonta              
  |                              
  +-- Alasmidonta_undulata       
  |                              
  +-- Beringiana                 
  |                              
  +-- Brachyanodon_kennerlyi **     
  |                              
  +-- Brachyanodon_nuttalliana **   
  |                              
  +-- Brachyanodon_oregonensis **   
  |                              
  +-- Arcidens_confragosus       
  |                              
  +-- Arcidens_wheeleri          
  |                              
  +-- Pterosyna                  
  |                              
--+-- Platynaias_compressa       
  |                              
  +-- Lasmigona                  
  |                              
  +-- Platynaias_decorata        
  |                              
  +-- Platynaias_subviridis      
  |                              
  +-- Pyganodon_grandis          
  |                              
  +-- Pyganodon_lacustris        
  |                              
  +-- Pseudodontoideus           
  |                              
  +-- Strophitus                 
  |                              
  +-- Utterbackia_imbecillis     
  |                              
  +-- Utterbackia_peninsularis__1
  |                              
  +-- Utterbackia_peninsularis__2

Wu et al., 2026 [Publ 4811], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 15 0 0 -- 
  +-- Brachyanodon_californiensis **
--+                              
  +-- Brachyanodon_nuttalliana **

Gustafson & Iwamoto, 2005 [Publ 2741], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 3 0 0 -- 
  +-- Brachyanodon_oregonensis **
--+                           
  +-- Brachyanodon_nuttalliana **

Zanatta et al., 2007 [Publ 2698], fig. 1B

iTtTbiriSoTroTroSiD
1 1 0 1 15 1 1 -- 
-- Brachyanodon_kennerlyi **

Campbell et al., 2005 [Publ 2660], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 101 1 1 -- 
  +-- Anodonta                
--+                           
  +-- Brachyanodon_oregonensis **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
1 1 0 1 121 3 2 -- 
  +----- Anodonta_cygnea         
--+                              
  |  +-- Anodonta_anatina        
  +--+                           
     +-- Brachyanodon_nuttalliana **

Hoeh, 1990 [Publ 2717], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 14 1 1 -- 
-- Brachyanodon_kennerlyi **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
1 1 0 1 91 4 2 -- 
-- Brachyanodon_nuttalliana **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
1 1 0 1 140 4 2 -- 
  +-- Brachyanodon_nuttalliana **
--+                           
  +-- Pseudanodonta

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
1 1 0 1 95 4 2 -- 
  +----- Anodonta                
--+                              
  |  +-- Brachyanodon_nuttalliana **
  +--+                           
     +-- Pseudanodonta

Cyr et al., 2007 [Publ 2745], fig. 2B

iTtTbiriSoTroTroSiD
1 1 0 1 4 0 0 -- 
-- Brachyanodon_nuttalliana **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
1 1 0 1 137 3 2 -- 
  +-- Anodonta                
--+                           
  +-- Brachyanodon_nuttalliana **

Lopes-Lima et al., 2019 [Publ 3901], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 27 0 0 -- 
-- Brachyanodon_nuttalliana **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
1 1 0 1 137 3 2 -- 
  +----- Pseudanodonta           
--+                              
  |  +-- Anodonta                
  +--+                           
     +-- Brachyanodon_nuttalliana **

Hoeh, 1990 [Publ 2717], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 14 1 1 -- 
-- Brachyanodon_kennerlyi **

Hoeh, 1990 [Publ 2717], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 14 1 1 -- 
-- Brachyanodon_kennerlyi **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
1 1 0 1 95 4 2 -- 
-- Brachyanodon_nuttalliana **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 199 3 2 -- 
  +----- Anodonta_anatina        
--+                              
  |  +-- Anodonta_cygnea         
  +--+                           
     +-- Brachyanodon_nuttalliana **

Zanatta et al., 2007 [Publ 2698], fig. 1A

iTtTbiriSoTroTroSiD
1 1 0 1 15 1 1 -- 
-- Brachyanodon_kennerlyi **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
1 1 0 1 91 4 2 -- 
-- Brachyanodon_nuttalliana **

Lopes-Lima et al., 2019 [Publ 3901], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 28 0 0 -- 
-- Brachyanodon_nuttalliana **

Lopes-Lima et al., 2019 [Publ 3901], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 25 0 0 -- 
-- Brachyanodon_nuttalliana **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
1 1 0 1 75 3 2 -- 
  +----- Pseudanodonta           
--+                              
  |  +-- Brachyanodon_nuttalliana **
  +--+                           
     +-- Anodonta

Riccardi et al., 2020 [Publ 4089], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 18 4 2 -- 
-- Brachyanodon_nuttalliana **

Wu et al., 2024 [Publ 4379], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 53 4 2 -- 
-- Brachyanodon_nuttalliana **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
1 1 0 1 79 3 2 -- 
  +----- Pseudanodonta           
  |                              
--+  +-- Anodonta_anatina        
  |  |                           
  +--+-- Anodonta_cygnea         
     |                           
     +-- Brachyanodon_nuttalliana **

Hou et al., 2025 [Publ 4445], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 52 4 2 -- 
-- Brachyanodon_nuttalliana **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
1 1 0 1 197 3 2 -- 
  +----- Anodonta_anatina        
--+                              
  |  +-- Anodonta_cygnea         
  +--+                           
     +-- Brachyanodon_nuttalliana **

Whelan et al., 2023 [Publ 4287], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 3 2 -- 
  +-- Brachyanodon_nuttalliana **
--+                           
  +-- Pseudanodonta

Hou et al., 2024 [Publ 4400], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 75 4 2 -- 
-- Brachyanodon_nuttalliana **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
1 1 0 1 75 3 2 -- 
  +-------- Pseudanodonta           
--+                                 
  |  +----- Anodonta_anatina        
  +--+                              
     |  +-- Brachyanodon_nuttalliana **
     +--+                           
        +-- Anodonta_cygnea

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
1 1 0 1 123 3 2 -- 
-- Brachyanodon_nuttalliana **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
1 1 0 1 199 3 2 -- 
-- Brachyanodon_nuttalliana **

Campbell et al., 2005 [Publ 2660], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 101 1 1 -- 
-- Brachyanodon_oregonensis **