MUSSELpdb
Taxonomy
Species & Genus Richness of
freshwater mussels
by Taxonomy
Species & Genus Richness of
freshwater bivalves
by Family
Geography
Species Richness & Endemism of
freshwater mussels
by Geography
Data last updated
Tue 02 June 2026
Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Anodontini | Anodontina
genus Brachyanodon Crosse & Fischer, 1894: cladomics
5 Recent valid species (5 total species)
Cladomics pages are experimental — even more so than the rest of this site!
The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.
The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.
Following the summary, the available published clades representing the taxon are listed.
For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!
Cladomics Summary
number of species = 5 (5 total)
analyzed in 37 cladograms in 24 publications
maximum number of terminals in an analysis = 5 (mean = 1.38, median = 1)
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 6
maximum number of included terminals = 5, (mean = 3.33 median = 3.5)
proportion of cladograms that support basic monophyly = 0.33
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Cladograms
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14
** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.500 | 5 | 8 | 7 | 5 | 627 | 5 | 2 | ** |
+----------------------- Anodonta_anatina | --+ +-------------------- Pseudanodonta | | +--+ +----------------- Brachyanodon_oregonensis__2 ** | | | | +----- Anodonta_cygnea +--+ +--------+ | | | +-- Anodonta_seddonae | | +--+ +--+ +-- Anodonta_exulcerata | | +----------- Brachyanodon_impura ** | | +--+ +-------- Brachyanodon ** | | +--+ +----- Brachyanodon_nuttalliana__1 ** | | +--+----- Brachyanodon_nuttalliana__2 ** | | +-- Brachyanodon_californiensis ** +--+ +-- Brachyanodon_oregonensis__1 **
Chong et al., 2008 [Publ 2668], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.250 | 4 | 18 | 10 | 4 | 14 | 3 | 2 | 23.0 |
+----------------- Anodonta_cygnea | +----------------- Pyganodon | +----------------- Alasminota | +----------------- Anodonta_anatina | +----------------- Unio | | +-- Brachyanodon_nuttalliana ** +--------------+ | +-- Brachyanodon_californiensis ** | --+ +-- Beringiana +--------------+ | +-- Sinanodonta | | +-- Pseudodontoideus | | +--------------+-- Alasmidonta | | | +-- Pterosyna | | +-------------- Pseudanodonta | | | | +----- Strophitus | +--------+ +--+ | +-- Prolasmidonta | +--+ | +-- Platynaias | | +----------- Brachyanodon_oregonensis__4 ** +--+ | +-------- Brachyanodon_oregonensis__3 ** +--+ | +----- Brachyanodon_oregonensis__2 ** +--+ | +-- Brachyanodon_oregonensis__1 ** +--+ +-- Brachyanodon_kennerlyi **
Galal-Khallaf et al., 2024 [Publ 4459], fig. 6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.000 | 4 | 7 | 5 | 4 | 24 | 3 | 1 | 14.0 |
+-- Anodonta_exulcerata
+--------+
| +-- Anodonta_cygnea
|
--+ +-- Brachyanodon_californiensis **
| +-----+
| | +-- Brachyanodon_nuttalliana **
| |
+--+ +-- Brachyanodon_kennerlyi **
| +--+
| | +-- Brachyanodon_oregonensis **
+--+
| +-- Anodonta_anatina
+--+
+-- Utterbackiana
Inoue et al., 2018 [Publ 3858], fig. 1_3a-c
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.000 | 3 | 20 | 0 | 3 | 102 | 0 | 0 | 27.0 |
+-- Alasmidonta_marginata
|
+-- Prolasmidonta
|
+-- Alasmidonta_undulata
|
+-- Beringiana
|
+-- Brachyanodon_kennerlyi **
|
+-- Brachyanodon_nuttalliana **
|
+-- Brachyanodon_oregonensis **
|
+-- Arcidens_confragosus
|
+-- Arcidens_wheeleri
|
+-- Pterosyna
|
+--------------------------------------------------+-- Platynaias_compressa
| |
| +-- Lasmigona
| |
| +-- Platynaias_decorata
| |
| +-- Platynaias_subviridis
| |
| +-- Pyganodon_grandis
| |
| +-- Pyganodon_lacustris
| |
| +-- Pseudodontoideus
| |
| +-- Strophitus
| |
--+ +-- Utterbackia_imbecillis
| |
| +-- Utterbackia_peninsularis__1
| |
| +-- Utterbackia_peninsularis__2
|
| +-------------------------------------------------- Gonidea
| |
| | +----------------------------------------------- Reginaia
| | |
| | +----------------------------------------------- Amblema
| | |
| | | +----- Popenaias
| | | |
| | | | +-- Dromus
| | | | |
| | | | +-- Epioblasma
+--+ | | |
| +-----------------------------------------+ +-- Glebula
| | | |
| | | +-- Lampsilis_ovata
| | | |
| | | +-- Lampsilis_virescens
| | | |
| | | +-- Lemiox
| | +--+
| | +-- Obliquaria
| | |
+--+ +-- Obovaria_olivaria
| |
| +-- Obovaria_unicolor
| |
| +-- Plectomerus
| |
| +-- Potamilus_alatus
| |
| +-- Potamilus_capax
|
| +----- Uniomerus
| |
| | +-- Cyclonaias
| +--------------------------------------+ |
| | | +-- Pustulosa_pustulosa__1
| | | |
| | | +-- Theliderma
| | +--+
| | +-- Pustulosa_pustulosa__2
| | |
+--+ +-- Quadrula_quadrula
| |
| +-- Quadrula_verrucosa
|
| +----------------------------------------- Elliptoideus
| |
| | +-------------------------------------- Parvaspina
| | |
+--+ | +----------------------------------- Plethobasus
| | |
| | +----------------------------------- Eurynia
| | |
| | +----------------------------------- Hemistena
+--+ |
| +----------------------------------- Elliptio
| |
| | +-------------------------------- Pleuronaia
| | |
| | | +-- Pleurobema_cordatum
| | | +--------------------------+
+--+ | | +-- Fusconaia
| | |
| | | +-------------------------- Pleurobema_plenum
| | | |
| | | | +----------------------- Pleurobema_stabile
| | | | |
| | | | | +----- Pleurobema_pyriforme
| | +--+ | +--+
| | | | | | | +-- Pleurobema_georgianum__2
| | | | | | +--+
| | | | | +-----------+ +-- Pleurobema_athearni
| | | | | | |
+--+ | +--+ | | +----- Pleurobema_decisum
| | | | | |
| | | | +--+ +-- Pleurobema_georgianum__1
| | | | | |
| | | | +--+-- Pleurobema_hanleyianum__1
| | | | |
| | | | +-- Pleurobema_hanleyianum__2
| | | |
| | +--+ +-------------- Pleurobema_beadleianum__2
| | | |
| | | +--+ +----------- Pleurobema_strodeanum__2
| | | | | |
| | | | +--+----------- Pleurobema_rubellum
| | | | |
| | | | | +-------- Pleurobema_beadleianum__1
| | | | +--+
+--+ | | | +----- Pleurobema_taitianum
| | | +--+
| | | | +-- Pleurobema_perovatum
| +--+ +--+
| | +-- Pleurobema_strodeanum__1
| |
| | +----- Pleurobema_oviforme__6
| | |
| | +--+----- Pleurobema_clava__3
| | | |
| | | | +-- Pleurobema_oviforme__5
| | | +--+
| +--------+ +-- Pleurobema_clava__2
| |
| | +----- Pleurobema_oviforme__2
| | |
| | +----- Pleurobema_oviforme__3
| +--+
| +----- Pleurobema_oviforme__4
| |
| | +-- Pleurobema_clava__1
| +--+
| +-- Pleurobema_oviforme__1
|
| +----------------- Pleurobema_riddellii
| |
| | +-------------- Pleurobema_sintoxia__34
| | |
| | | +----------- Pleurobema_sintoxia__31
| | | |
+-----------+ | +----------- Pleurobema_sintoxia__32
| | |
| | +----------- Pleurobema_sintoxia__33
| | |
| | | +-------- Pleurobema__4
| | | |
| | | +-------- Pleurobema__5
+--+ | |
| | +-------- Pleurobema_sintoxia__2
| | |
| | +-------- Pleurobema_sintoxia__3
| | |
| | +-------- Pleurobema__8
| | |
| | +-------- Pleurobema__9
| | |
| | +-------- Pleurobema__10
| | |
| | +-------- Pleurobema_sintoxia__5
+--+ |
| +-------- Pleurobema_sintoxia__6
| |
| +-------- Pleurobema_sintoxia__7
| |
| +-------- Pleurobema_sintoxia__8
| |
| +-------- Pleurobema_sintoxia__9
| |
| +-------- Pleurobema_sintoxia__10
| |
| +-------- Pleurobema_sintoxia__11
| |
| +-------- Pleurobema_sintoxia__12
| |
| +-------- Pleurobema_sintoxia__13
| |
| +-------- Pleurobema_sintoxia__14
| |
| +-------- Pleurobema_sintoxia__15
| |
| +-------- Pleurobema_sintoxia__16
+--+
+-------- Pleurobema_sintoxia__17
|
+-------- Pleurobema_sintoxia__18
|
+-------- Pleurobema_sintoxia__19
|
+-------- Pleurobema_sintoxia__20
|
+-------- Pleurobema_sintoxia__21
|
+-------- Pleurobema_sintoxia__22
|
+-------- Pleurobema_sintoxia__23
|
+-------- Pleurobema_sintoxia__24
|
+-------- Pleurobema_sintoxia__25
|
+-------- Pleurobema_sintoxia__26
|
+-------- Pleurobema_sintoxia__27
|
+-------- Pleurobema_sintoxia__28
|
+-------- Pleurobema_sintoxia__29
|
+-------- Pleurobema_sintoxia__30
|
| +-- Pleurobema__6
+-----+
| +-- Pleurobema_sintoxia__1
|
| +-- Pleurobema__7
+-----+
| +-- Pleurobema_sintoxia__4
|
| +----- Pleurobema__3
+--+
| +-- Pleurobema__1
+--+
+-- Pleurobema__2
Gustafson & Iwamoto, 2005 [Publ 2741], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 3 | 0 | 0 | -- |
+----- Beringiana --+ | +-- Brachyanodon_oregonensis ** +--+ +-- Brachyanodon_nuttalliana **
Wu et al., 2026 [Publ 4811], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 15 | 0 | 0 | -- |
+----- Acuticosta --+ | +-- Brachyanodon_californiensis ** +--+ +-- Brachyanodon_nuttalliana **
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.750 | 1 | 1 | 0 | 1 | 80 | 3 | 2 | -- |
+----- Pseudanodonta | --+ +-- Anodonta_anatina | | +--+-- Anodonta_cygnea | +-- Brachyanodon_nuttalliana **
Bolotov et al., 2017 [Publ 3761], fig. S1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.500 | 1 | 1 | 0 | 1 | 138 | 3 | 2 | -- |
+----- Pseudanodonta --+ | +-- Anodonta +--+ +-- Brachyanodon_nuttalliana **
Bolotov et al., 2017 [Publ 3761], fig. S2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.500 | 1 | 1 | 0 | 1 | 138 | 3 | 2 | -- |
+-- Anodonta --+ +-- Brachyanodon_nuttalliana **
Bolotov et al., 2017 [Publ 3764], fig. S1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.500 | 1 | 1 | 0 | 1 | 124 | 3 | 2 | -- |
+-- Alasmidonta | +-- Anodonta_anatina | +-- Anodonta_cygnea | +-- Brachyanodon_nuttalliana ** | +-- Lasmigona | +-- Pseudanodonta | +-- Pyganodon | +-- Simpsonaias | +-- Strophitus --+ +-- Sinanodonta_lauta | +-- Cristaria_plicata | +-- Cristaria_radiata | +-- Pletholophus | +-- Sinanodonta_lucida | +-- Sinanodonta_jourdyi | +-- Lanceolaria_gladiola | +-- Lanceolaria_grayii | +-- Lanceolaria_danzhouensis
Huang et al., 2019 [Publ 3843], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.500 | 1 | 1 | 0 | 1 | 200 | 3 | 2 | -- |
+----- Anodonta_anatina --+ | +-- Anodonta_cygnea +--+ +-- Brachyanodon_nuttalliana **
Huang et al., 2019 [Publ 3843], fig. S3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.500 | 1 | 1 | 0 | 1 | 198 | 3 | 2 | -- |
+----- Anodonta_anatina --+ | +-- Anodonta_cygnea +--+ +-- Brachyanodon_nuttalliana **
Huang et al., 2019 [Publ 3843], fig. S4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.500 | 1 | 1 | 0 | 1 | 200 | 3 | 2 | -- |
+-------- Acuticosta | +-------- Alasmidonta | +-------- Anodonta_anatina | +-------- Anemina | +-------- Anodonta_cygnea | +-------- Cristaria | +-------- Brachyanodon_nuttalliana ** | +-------- Lasmigona | +-------- Lepidodesma | +-------- Pseudanodonta | +-------- Pyganodon | --+-------- Simpsonaias | +-------- Strophitus | +-------- Utterbackia | +-------- Schistodesmus | +-------- Unio | +-------- Scriptolamprotula | +-------- Lanceolaria | | +-- Pletholophus +-----+ | +-- Sinanodonta | | +----- Cuneopsis_celtiformis | | | +----- Tchangsinaia | | +--+----- Cuneopsis_rufescens | +----- Nodularia | | +-- Cuneopsis_szechenyii +--+ +-- Cuneopsis_heudei
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.500 | 1 | 1 | 0 | 1 | 121 | 3 | 2 | -- |
+----- Anodonta_cygnea --+ | +-- Anodonta_anatina +--+ +-- Brachyanodon_nuttalliana **
Riccardi et al., 2020 [Publ 4089], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.500 | 1 | 1 | 0 | 1 | 18 | 4 | 2 | -- |
+-------- Brachyanodon_nuttalliana ** --+ | +----- Platynaias +--+ | +-- Pyganodon +--+ +-- Utterbackia
Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 75 | 3 | 2 | -- |
+----- Pseudanodonta --+ | +-- Brachyanodon_nuttalliana ** +--+ +-- Anodonta
Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.750 | 1 | 1 | 0 | 1 | 75 | 3 | 2 | -- |
+-------- Pseudanodonta --+ | +----- Anodonta_anatina +--+ | +-- Brachyanodon_nuttalliana ** +--+ +-- Anodonta_cygnea
Whelan et al., 2023 [Publ 4287], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.500 | 1 | 1 | 0 | 1 | 40 | 3 | 2 | -- |
+-- Brachyanodon_nuttalliana ** --+ +-- Pseudanodonta
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 94 | 4 | 2 | -- |
+----- Anodonta --+ | +-- Brachyanodon_nuttalliana ** +--+ +-- Pseudanodonta
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.500 | 1 | 1 | 0 | 1 | 94 | 4 | 2 | -- |
+----------------- Brachyanodon_nuttalliana ** | +----------------- Pseudanodonta | --+ +-------------- Anodonta_cygnea | | | | +-- Anodonta_exulcerata | +-----------+ +--+ +-- Anodonta_anatina | | +-- Pyganodon | +--------+ | | +-- Utterbackia +--+ | +-------- Pterosyna +--+ | +----- Prolasmidonta +--+ | +-- Alasmidonta +--+ +-- Platynaias
Wu et al., 2024 [Publ 4350], fig. 8a_S1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.750 | 1 | 1 | 0 | 1 | 90 | 4 | 2 | -- |
+-------------- Brachyanodon_nuttalliana ** | --+ +-- Pyganodon | +--------+ | | +-- Utterbackia +--+ | +-------- Pterosyna +--+ | +----- Prolasmidonta +--+ | +-- Alasmidonta +--+ +-- Platynaias
Wu et al., 2024 [Publ 4350], fig. 8b_S2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.750 | 1 | 1 | 0 | 1 | 90 | 4 | 2 | -- |
+----------------- Brachyanodon_nuttalliana ** | | +-- Pseudanodonta --+ +-----------+ | | +-- Anodonta | | +--+ +-- Pyganodon | +--------+ | | +-- Utterbackia +--+ | +-------- Pterosyna +--+ | +----- Prolasmidonta +--+ | +-- Alasmidonta +--+ +-- Platynaias
Wu et al., 2024 [Publ 4379], fig. 5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.750 | 1 | 1 | 0 | 1 | 52 | 4 | 2 | -- |
+----------------- Brachyanodon_nuttalliana ** | | +-- Pseudanodonta --+ +-----------+ | | +-- Anodonta | | +--+ +-- Pyganodon | +--------+ | | +-- Utterbackia +--+ | +-------- Pterosyna +--+ | +----- Prolasmidonta +--+ | +-- Alasmidonta +--+ +-- Platynaias
Hou et al., 2024 [Publ 4400], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.750 | 1 | 1 | 0 | 1 | 74 | 4 | 2 | -- |
+-------------- Brachyanodon_nuttalliana ** | --+ +-- Pyganodon | +--------+ | | +-- Utterbackia +--+ | +-------- Pterosyna +--+ | +----- Prolasmidonta +--+ | +-- Alasmidonta +--+ +-- Platynaias
Hou et al., 2025 [Publ 4445], fig. 5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.750 | 1 | 1 | 0 | 1 | 51 | 4 | 2 | -- |
+----------------- Brachyanodon_nuttalliana ** | | +-- Pseudanodonta --+ +-----------+ | | +-- Anodonta | | +--+ +-- Pyganodon | +--------+ | | +-- Utterbackia +--+ | +-------- Pterosyna +--+ | +----- Prolasmidonta +--+ | +-- Platynaias +--+ +-- Alasmidonta
Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.500 | 1 | 1 | 0 | 1 | 139 | 4 | 2 | -- |
+-- Brachyanodon_nuttalliana ** --+ +-- Pseudanodonta
Hoeh, 1990 [Publ 2717], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.667 | 1 | 1 | 0 | 1 | 14 | 1 | 1 | -- |
+-- Brachyanodon_kennerlyi ** --+ +-- Utterbackiana
Hoeh, 1990 [Publ 2717], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.667 | 1 | 1 | 0 | 1 | 14 | 1 | 1 | -- |
+----- Pyganodon_fragilis | +----- Utterbackia_imbecillis | +----- Utterbackia_peggyae | +----- Anodonta | +----- Brachyanodon_kennerlyi ** | +----- Utterbackiana_couperiana__1 | --+----- Utterbackiana_couperiana__2 | +----- Utterbackiana_suborbiculata | | +-- Lasmigona +--+ | +-- Utterbackiana_implicata | | +-- Pterosyna | | | +-- Platynaias | | | +-- Pyganodon_cataracta +--+ +-- Pyganodon_gibbosa | +-- Pyganodon_grandis | +-- Pyganodon_lacustris
Hoeh, 1990 [Publ 2717], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.667 | 1 | 1 | 0 | 1 | 14 | 1 | 1 | -- |
+-- Brachyanodon_kennerlyi ** --+ +-- Utterbackiana
Campbell et al., 2005 [Publ 2660], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.667 | 1 | 1 | 0 | 1 | 101 | 1 | 1 | -- |
+----------- Brachyanodon_oregonensis ** | --+ +-- Anodonta | +-----+ | | +-- Alasminota +--+ | +----- Pyganodon +--+ | +-- Lasmigona +--+ +-- Pseudodontoideus
Campbell et al., 2005 [Publ 2660], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 101 | 1 | 1 | -- |
+-- Anodonta --+ +-- Brachyanodon_oregonensis **
Zanatta et al., 2007 [Publ 2698], fig. 1A
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.667 | 1 | 1 | 0 | 1 | 15 | 1 | 1 | -- |
+-- Brachyanodon_kennerlyi ** --+ +-- Utterbackiana
Zanatta et al., 2007 [Publ 2698], fig. 1B
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.667 | 1 | 1 | 0 | 1 | 15 | 1 | 1 | -- |
+-- Brachyanodon_kennerlyi ** --+ +-- Utterbackiana
Cyr et al., 2007 [Publ 2745], fig. 2B
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 4 | 0 | 0 | -- |
+-- Brachyanodon_nuttalliana ** --+ +-- Pyganodon
Lopes-Lima et al., 2019 [Publ 3901], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 27 | 0 | 0 | -- |
+-- Brachyanodon_nuttalliana ** --+ +-- Pyganodon
Lopes-Lima et al., 2019 [Publ 3901], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 28 | 0 | 0 | -- |
+-- Brachyanodon_nuttalliana ** --+ +-- Pyganodon
Lopes-Lima et al., 2019 [Publ 3901], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 25 | 0 | 0 | -- |
+-- Brachyanodon_nuttalliana ** --+ +-- Pyganodon
