Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Anodontini | Alasmidontina

genus Pyganodon Crosse & Fischer, 1894: cladomics

6 Recent valid species (6 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 6 (6 total)
analyzed in 184 cladograms in 111 publications
maximum number of terminals in an analysis = 5 (mean = 1.34, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 33
maximum number of included terminals = 5, (mean = 2.88 median = 2)
proportion of cladograms that support basic monophyly = 0.82

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Pyganodon cladogram distances
PCoA Ordination

NJ Tree of Pyganodon cladogram distances
NJ Tree

Cladograms

Cyr et al., 2007 [Publ 2745], fig. 2C

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
4 4 3 8 1 0 0 0 
        +-- Pyganodon_grandis **     
  +-----+                         
  |     +-- Pyganodon__1 **          
  |                               
--+  +----- Pyganodon__3 **          
  |  |                            
  |  |  +-- Pyganodon_cataracta__1 **
  +--+  |                         
     |  +-- Pyganodon_fragilis__1 ** 
     |  |                         
     +--+-- Pyganodon__2 **          
        |                         
        +-- Pyganodon_cataracta__2 **
        |                         
        +-- Pyganodon_fragilis__2 **

Perkins et al., 2025 [Publ 4447], fig. 2_mtDNA-ITS

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
1 1 0 1 35 32 14 -- 
     +-------------- Utterbackiana    
  +--+                                
  |  |  +----------- Alasminota       
  |  +--+                             
  |     |  +-------- Pseudodontoideus 
  |     +--+                          
  |        |  +----- Lasmigona        
  |        +--+                       
  |           |  +-- Pterosyna        
  |           +--+                    
--+              +-- Arcidens         
  |                                   
  |              +-- Prolasmidonta    
  |           +--+                    
  |           |  +-- Platynaias       
  |     +-----+                       
  |     |     |  +-- Ligodonta        
  |     |     +--+                    
  |     |        +-- Alasmidonta      
  +-----+                             
        |        +-- Anodontoides     
        |  +-----+                    
        |  |     +-- Strophitus       
        +--+                          
           |  +----- Sulcularia       
           +--+                       
              |  +-- Pyganodon_grandis **
              +--+                    
                 +-- Utterbackia

Hoeh, 1990 [Publ 2717], fig. 4

iTtTbiriSoTroTroSiD
5 5 3 5 10 8 5 9 
  +----------- Utterbackia        
  |                               
--+  +-------- Pyganodon_fragilis ** 
  |  |                            
  +--+     +-- Pyganodon_cataracta **
     |  +--+                      
     |  |  +-- Pyganodon_gibbosa **  
     +--+                         
        |  +-- Pyganodon_lacustris **
        +--+                      
           +-- Pyganodon_grandis **

Zanatta et al., 2007 [Publ 2698], fig. 1A

iTtTbiriSoTroTroSiD
5 5 0 5 11 9 5 8 
  +----- Utterbackia        
  |                         
  |  +-- Pyganodon_gibbosa **  
--+  |                      
  |  +-- Pyganodon_cataracta **
  |  |                      
  +--+-- Pyganodon_fragilis ** 
     |                      
     +-- Pyganodon_grandis **  
     |                      
     +-- Pyganodon_lacustris **

Hoeh, 1990 [Publ 2717], fig. 3

iTtTbiriSoTroTroSiD
5 15 2 5 10 8 5 18 
  +----- Pyganodon_fragilis **         
  |                                 
  +----- Utterbackia_imbecillis     
  |                                 
  +----- Utterbackia_peggyae        
  |                                 
  +----- Anodonta_cygnea            
  |                                 
  +----- Anodonta_kennerlyi         
  |                                 
  +----- Utterbackiana_couperiana__1
  |                                 
--+----- Utterbackiana_couperiana__2
  |                                 
  +----- Utterbackiana_suborbiculata
  |                                 
  |  +-- Lasmigona                  
  +--+                              
  |  +-- Utterbackiana_implicata    
  |                                 
  |  +-- Pterosyna                  
  |  |                              
  |  +-- Platynaias                 
  |  |                              
  |  +-- Pyganodon_cataracta **        
  +--+                              
     +-- Pyganodon_gibbosa **          
     |                              
     +-- Pyganodon_grandis **          
     |                              
     +-- Pyganodon_lacustris **

Hoeh, 1990 [Publ 2717], fig. 2

iTtTbiriSoTroTroSiD
5 5 3 5 10 8 5 7 
  +-------------- Utterbackia        
--+                                  
  |  +----------- Pyganodon_lacustris **
  +--+                               
     |  +-------- Pyganodon_grandis **  
     +--+                            
        |  +----- Pyganodon_fragilis ** 
        +--+                         
           |  +-- Pyganodon_cataracta **
           +--+                      
              +-- Pyganodon_gibbosa **

Zanatta et al., 2007 [Publ 2698], fig. 1B

iTtTbiriSoTroTroSiD
5 5 3 5 11 9 5 7 
  +-------------- Utterbackia        
--+                                  
  |  +----------- Pyganodon_lacustris **
  +--+                               
     |  +-------- Pyganodon_grandis **  
     +--+                            
        |  +----- Pyganodon_fragilis ** 
        +--+                         
           |  +-- Pyganodon_gibbosa **  
           +--+                      
              +-- Pyganodon_cataracta **

Hoeh et al., 1995 [Publ 3391], fig. 2

iTtTbiriSoTroTroSiD
4 4 1 4 3 3 1 6 
  +-------- Utterbackia        
  |                            
--+  +----- Pyganodon_lacustris **
  |  |                         
  +--+----- Pyganodon_grandis **  
     |                         
     |  +-- Pyganodon_cataracta **
     +--+                      
        +-- Pyganodon_fragilis **

Inoue et al., 2014 [Publ 3635], fig. 3B

iTtTbiriSoTroTroSiD
4 7 2 4 4 3 2 8 
  +----- Pyganodon_grandis **  
  |                         
  +----- Pyganodon_cataracta **
  |                         
--+----- Pyganodon_fragilis ** 
  |                         
  +----- Pyganodon **          
  |                         
  |  +-- Platynaias         
  +--+                      
     +-- Arcidens

Cyr et al., 2007 [Publ 2745], fig. 2D

iTtTbiriSoTroTroSiD
4 4 2 5 0 0 0 5 
  +-------- Pyganodon_grandis **  
  |                            
--+-------- Pyganodon__2 **       
  |                            
  |  +----- Pyganodon__1 **       
  +--+                         
     |  +-- Pyganodon_cataracta **
     +--+                      
        +-- Pyganodon_fragilis **

Cyr et al., 2007 [Publ 2745], fig. 2A

iTtTbiriSoTroTroSiD
4 4 2 5 1 0 0 5 
  +-------- Pyganodon_grandis **  
  |                            
--+-------- Pyganodon__2 **       
  |                            
  |  +----- Pyganodon__1 **       
  +--+                         
     |  +-- Pyganodon_cataracta **
     +--+                      
        +-- Pyganodon_fragilis **

Cyr et al., 2007 [Publ 2745], fig. 2B

iTtTbiriSoTroTroSiD
4 4 3 5 1 0 0 5 
        +-- Pyganodon_grandis **  
  +-----+                      
  |     +-- Pyganodon__1 **       
--+                            
  |  +----- Pyganodon__2 **       
  +--+                         
     |  +-- Pyganodon_cataracta **
     +--+                      
        +-- Pyganodon_fragilis **

Stewart et al., 2021 [Publ 4451], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 13 7 5 7 
              +-- Alasmidonta        
  +-----------+                      
  |           +-- Pterosyna          
  |                                  
--+           +-- Strophitus         
  |  +--------+                      
  |  |        +-- Platynaias         
  +--+                               
     |  +-------- Utterbackia        
     +--+                            
        |  +----- Pyganodon_grandis **  
        +--+                         
           |  +-- Pyganodon_cataracta **
           +--+                      
              +-- Pyganodon_lacustris **

Krebs et al., 2019 [Publ 4092], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 3 0 0 0 5 
  +----- Pyganodon_grandis **  
--+                         
  |  +-- Pyganodon_fragilis ** 
  +--+                      
     +-- Pyganodon_cataracta **

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

iTtTbiriSoTroTroSiD
2 20 0 2 102 14 9 23 
  +-- Alasmidonta_marginata      
  |                              
  +-- Prolasmidonta              
  |                              
  +-- Alasmidonta_undulata       
  |                              
  +-- Beringiana                 
  |                              
  +-- Anodonta_kennerlyi         
  |                              
  +-- Anodonta_nuttalliana       
  |                              
  +-- Anodonta_oregonensis       
  |                              
  +-- Arcidens_confragosus       
  |                              
  +-- Arcidens_wheeleri          
  |                              
  +-- Pterosyna                  
  |                              
--+-- Platynaias_compressa       
  |                              
  +-- Lasmigona                  
  |                              
  +-- Platynaias_decorata        
  |                              
  +-- Platynaias_subviridis      
  |                              
  +-- Pyganodon_grandis **          
  |                              
  +-- Pyganodon_lacustris **        
  |                              
  +-- Pseudodontoideus           
  |                              
  +-- Strophitus                 
  |                              
  +-- Utterbackia_imbecillis     
  |                              
  +-- Utterbackia_peninsularis__1
  |                              
  +-- Utterbackia_peninsularis__2

Rosenberg et al., 1997 [Publ 2720], fig. 2B

iTtTbiriSoTroTroSiD
2 17 0 2 20 1 1 18 
  +-- Gonidea            
  |                      
  +-- Pyganodon_grandis **  
  |                      
  +-- Utterbackia        
  |                      
  +-- Pyganodon_cataracta **
  |                      
  +-- Lampsilis_straminea
  |                      
  +-- Lampsilis_teres    
  |                      
  +-- Megalonaias        
  |                      
  +-- Obliquaria         
  |                      
--+-- Theliderma         
  |                      
  +-- Quadrula           
  |                      
  +-- Uniomerus          
  |                      
  +-- Amblema            
  |                      
  +-- Elliptio           
  |                      
  +-- Fusconaia          
  |                      
  +-- Plectomerus        
  |                      
  +-- Aculamprotula      
  |                      
  +-- Unio

Walker et al., 2006 [Publ 2666], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 13 0 0 -- 
  +-- Pyganodon_fragilis **
--+                     
  +-- Pyganodon_grandis **

Chase et al., 2018 [Publ 3847], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 36 9 6 -- 
        +----- Utterbackia        
  +-----+                         
  |     |  +-- Pyganodon_lacustris **
  |     +--+                      
  |        +-- Pyganodon_grandis **  
--+                               
  |        +-- Lasmigona          
  |  +-----+                      
  |  |     +-- Pterosyna          
  +--+                            
     |  +----- Alasmidonta        
     +--+                         
        |  +-- Strophitus         
        +--+                      
           +-- Platynaias

Walker et al., 2006 [Publ 2666], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 13 0 0 -- 
  +-- Pyganodon_fragilis **
--+                     
  +-- Pyganodon_grandis **

Whelan et al., 2023 [Publ 4287], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 25 23 10 -- 
  +-------------------- Sulcularia         
  |                                        
  |           +-------- Utterbackiana      
--+  +--------+                            
  |  |        |  +----- Alasminota         
  |  |        +--+                         
  |  |           |  +-- Arcidens           
  +--+           +--+                      
     |              +-- Pseudodontoideus   
     |                                     
     |  +-------------- Alasmidonta        
     |  |                                  
     +--+  +----------- Platynaias         
        |  |                               
        +--+  +-------- Strophitus         
           |  |                            
           +--+     +-- Pyganodon_grandis **  
              |  +--+                      
              |  |  +-- Pyganodon_cataracta **
              +--+                         
                 |  +-- Utterbackia        
                 +--+                      
                    +-- Anodontoides

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

iTtTbiriSoTroTroSiD
2 2 0 2 28 3 3 -- 
        +-- Alasmidonta       
  +-----+                     
  |     +-- Lasmigona         
--+                           
  |  +----- Strophitus        
  +--+                        
     |  +-- Pyganodon_grandis ** 
     +--+                     
        +-- Pyganodon_fragilis **

Whelan et al., 2023 [Publ 4287], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 3 39 31 13 -- 
        +-------- Utterbackia         
  +-----+                             
  |     |  +----- Pyganodon_grandis__2 **
  |     +--+                          
  |        |  +-- Pyganodon_grandis__1 **
  |        +--+                       
  |           +-- Pyganodon_lacustris ** 
  |                                   
--+     +-------- Utterbackiana       
  |     |                             
  |  +--+     +-- Lasmigona           
  |  |  |  +--+                       
  |  |  |  |  +-- Pseudodontoideus    
  |  |  +--+                          
  |  |     |  +-- Pterosyna           
  |  |     +--+                       
  +--+        +-- Arcidens            
     |                                
     |     +----- Sulcularia          
     |  +--+                          
     |  |  |  +-- Anodontoides        
     |  |  +--+                       
     |  |     +-- Strophitus          
     +--+                             
        |     +-- Prolasmidonta       
        |  +--+                       
        |  |  +-- Platynaias          
        +--+                          
           |  +-- Alasminota          
           +--+                       
              +-- Alasmidonta

Wu et al., 2018 [Publ 3836], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 51 3 3 -- 
  +-------- Strophitus         
  |                            
--+  +----- Platynaias         
  |  |                         
  +--+----- Alasmidonta        
     |                         
     |  +-- Pyganodon_cataracta **
     +--+                      
        +-- Pyganodon_grandis **

Rosenberg et al., 1994 [Publ 2719], fig. 3

iTtTbiriSoTroTroSiD
2 10 0 2 17 1 1 11 
  +-- Pyganodon_cataracta **
  |                      
  +-- Pyganodon_grandis **  
  |                      
  +-- Utterbackia        
  |                      
  +-- Lampsilis_teres    
  |                      
  +-- Lampsilis_straminea
--+                      
  +-- Megalonaias        
  |                      
  +-- Obliquaria         
  |                      
  +-- Theliderma         
  |                      
  +-- Quadrula           
  |                      
  +-- Uniomerus

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

iTtTbiriSoTroTroSiD
2 2 0 2 28 3 3 -- 
        +-- Alasmidonta       
  +-----+                     
  |     +-- Lasmigona         
--+                           
  |  +----- Strophitus        
  +--+                        
     |  +-- Pyganodon_grandis ** 
     +--+                     
        +-- Pyganodon_fragilis **

Hoeh et al., 2002 [Publ 2673], fig. 1-F

iTtTbiriSoTroTroSiD
2 2 0 2 7 0 0 -- 
  +-- Pyganodon_fragilis **
--+                     
  +-- Pyganodon_grandis **

Walker et al., 2006 [Publ 2666], fig. 6-M

iTtTbiriSoTroTroSiD
2 2 0 2 12 0 0 -- 
  +-- Pyganodon_fragilis **
--+                     
  +-- Pyganodon_grandis **

Hoeh et al., 2002 [Publ 2673], fig. 1-M

iTtTbiriSoTroTroSiD
2 2 0 2 7 0 0 -- 
  +-- Pyganodon_fragilis **
--+                     
  +-- Pyganodon_grandis **

King et al., 1999 [Publ 2699], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 7 7 5 -- 
           +-- Pyganodon_grandis ** 
  +--------+                     
  |        +-- Pyganodon_fragilis **
  |                              
--+        +-- Lasmigona         
  |  +-----+                     
  |  |     +-- Pterosyna         
  +--+                           
     |  +----- Strophitus        
     +--+                        
        |  +-- Prolasmidonta     
        +--+                     
           +-- Platynaias

Walker et al., 2006 [Publ 2666], fig. 6

iTtTbiriSoTroTroSiD
2 2 0 2 34 0 0 -- 
  +-- Pyganodon_fragilis **
--+                     
  +-- Pyganodon_grandis **

Hoeh et al., 1996 [Publ 2694], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 2 0 0 -- 
  +-- Pyganodon_fragilis **
--+                     
  +-- Pyganodon_grandis **

Walker et al., 2006 [Publ 2666], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 13 0 0 -- 
  +-- Pyganodon_fragilis **
--+                     
  +-- Pyganodon_grandis **

Breton et al., 2011 [Publ 3376], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 35 9 6 -- 
              +-- Pterosyna          
  +-----------+                      
  |           +-- Lasmigona          
--+                                  
  |  +----------- Strophitus         
  |  |                               
  +--+        +-- Platynaias         
     |  +-----+                      
     |  |     +-- Alasmidonta        
     +--+                            
        |  +----- Utterbackia        
        +--+                         
           |  +-- Pyganodon_lacustris **
           +--+                      
              +-- Pyganodon_grandis **

Rosenberg et al., 1997 [Publ 2720], fig. 2A

iTtTbiriSoTroTroSiD
2 17 0 2 20 1 1 18 
  +-- Gonidea            
  |                      
  +-- Pyganodon_grandis **  
  |                      
  +-- Utterbackia        
  |                      
  +-- Pyganodon_cataracta **
  |                      
  +-- Lampsilis_straminea
  |                      
  +-- Lampsilis_teres    
  |                      
  +-- Megalonaias        
  |                      
  +-- Obliquaria         
  |                      
--+-- Theliderma         
  |                      
  +-- Quadrula           
  |                      
  +-- Uniomerus          
  |                      
  +-- Amblema            
  |                      
  +-- Elliptio           
  |                      
  +-- Fusconaia          
  |                      
  +-- Plectomerus        
  |                      
  +-- Pleurobema         
  |                      
  +-- Unio

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 40 4 2 -- 
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia

Graf & Cummings, 2006 [Publ 1968], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 46 3 3 -- 
  +-------- Lasmigona        
--+                          
  |  +----- Alasmidonta      
  +--+                       
     |  +-- Pyganodon_grandis **
     +--+                    
        +-- Strophitus

Guerra et al., 2017 [Publ 3772], fig. 2a

iTtTbiriSoTroTroSiD
1 1 0 1 24 4 2 -- 
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia

Krebs et al., 2003 [Publ 2691], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 32 8 7 -- 
-- Pyganodon_grandis **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
1 1 0 1 124 4 4 -- 
-- Pyganodon_grandis **

Campbell et al., 2005 [Publ 2660], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 101 3 3 -- 
  +----- Pyganodon_grandis **
--+                       
  |  +-- Lasmigona        
  +--+                    
     +-- Pseudodontoideus

Graf, 2000 [Publ 2647], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 16 0 0 -- 
-- Pyganodon **

Burzynski et al., 2017 [Publ 3774], fig. 1-top

iTtTbiriSoTroTroSiD
1 1 0 1 12 1 1 -- 
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iTtTbiriSoTroTroSiD
1 1 0 1 30 1 1 -- 
-- Pyganodon_grandis **

Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 6 1 1 -- 
  +-- Pyganodon_grandis **
--+                    
  +-- Utterbackia

Wang et al., 2018 [Publ 3865], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 1 1 -- 
  +-- Pyganodon_grandis **
--+                    
  +-- Platynaias

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
1 1 0 1 95 6 5 -- 
           +-- Pyganodon_grandis **
  +--------+                    
  |        +-- Utterbackia      
--+                             
  |  +-------- Pterosyna        
  +--+                          
     |  +----- Prolasmidonta    
     +--+                       
        |  +-- Alasmidonta      
        +--+                    
           +-- Platynaias

Huang et al., 2019 [Publ 3843], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 22 1 1 -- 
  +-- Pyganodon_grandis **
--+                    
  +-- Utterbackia

Huang et al., 2018 [Publ 3828], fig. 6-M

iTtTbiriSoTroTroSiD
1 1 0 1 16 1 1 -- 
  +-- Pyganodon_grandis **
--+                    
  +-- Utterbackia

Amaro et al., 2017 [Publ 3791], fig. 3B-M

iTtTbiriSoTroTroSiD
1 1 0 1 6 1 1 -- 
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Hoeh et al., 2009 [Publ 2716], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 45 3 3 -- 
  +-------- Lasmigona        
--+                          
  |  +----- Alasmidonta      
  +--+                       
     |  +-- Strophitus       
     +--+                    
        +-- Pyganodon_grandis **

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 43 3 2 -- 
  +----- Pterosyna        
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia

Lopes-Lima et al., 2019 [Publ 3901], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 28 0 0 -- 
-- Pyganodon_grandis **

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 87 3 2 -- 
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia

Froufe et al., 2019 [Publ 3936], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 18 1 1 -- 
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Chase et al., 2018 [Publ 3861], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 10 1 1 -- 
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 52 2 2 -- 
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia

Guerra et al., 2017 [Publ 3772], fig. 2b-M

iTtTbiriSoTroTroSiD
1 1 0 1 7 1 1 -- 
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Wu et al., 2016 [Publ 3711], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 20 2 1 -- 
  +-- Pyganodon_grandis **
--+                    
  +-- Utterbackia

Araujo et al., 2018 [Publ 3779], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 17 1 1 -- 
  +-- Pyganodon_grandis **
--+                    
  +-- Alasmidonta

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 33 1 1 -- 
  +-- Strophitus       
--+                    
  +-- Pyganodon_grandis **

Guerra et al., 2017 [Publ 3772], fig. 2b

iTtTbiriSoTroTroSiD
1 1 0 1 24 4 2 -- 
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia

Smith et al., 2018 [Publ 3790], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 17 16 7 -- 
           +-- Utterbackia      
  +--------+                    
  |        +-- Pyganodon_grandis **
  |                             
--+        +-- Anodontoides     
  |  +-----+                    
  |  |     +-- Strophitus       
  |  |                          
  +--+     +-- Alasmidonta      
     |  +--+                    
     |  |  +-- Alasminota       
     +--+                       
        |  +-- Pseudodontoideus 
        +--+                    
           +-- Utterbackiana

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 -- 
-- Pyganodon_grandis **

Liu et al., 2023 [Publ 4310], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 23 2 2 -- 
  +----- Lasmigona        
--+                       
  |  +-- Strophitus       
  +--+                    
     +-- Pyganodon_grandis **

Hou et al., 2024 [Publ 4400], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 75 6 5 -- 
           +-- Pyganodon_grandis **
  +--------+                    
  |        +-- Utterbackia      
--+                             
  |  +-------- Pterosyna        
  +--+                          
     |  +----- Prolasmidonta    
     +--+                       
        |  +-- Alasmidonta      
        +--+                    
           +-- Platynaias

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 90 2 2 -- 
  +-- Pyganodon_grandis **
--+                    
  +-- Strophitus

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 67 5 4 -- 
        +-- Pyganodon_grandis **
  +-----+                    
  |     +-- Utterbackia      
--+                          
  |  +----- Alasmidonta      
  +--+                       
     |  +-- Prolasmidonta    
     +--+                    
        +-- Platynaias

Hoeh et al., 2002 [Publ 2653], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 1 1 -- 
  +-- Pyganodon_grandis **
--+                    
  +-- Strophitus

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
1 1 0 1 35 1 1 -- 
  +-- Pyganodon_grandis **
--+                    
  +-- Lasmigona

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
1 1 0 1 91 6 5 -- 
           +-- Pyganodon_grandis **
  +--------+                    
  |        +-- Utterbackia      
--+                             
  |  +-------- Pterosyna        
  +--+                          
     |  +----- Prolasmidonta    
     +--+                       
        |  +-- Alasmidonta      
        +--+                    
           +-- Platynaias

Amaro et al., 2017 [Publ 3791], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 15 4 3 -- 
     +-- Lasmigona        
  +--+                    
  |  +-- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 85 3 2 -- 
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia

Wang et al., 2018 [Publ 3846], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 1 1 -- 
  +-- Pyganodon_grandis **
--+                    
  +-- Platynaias

Lopes-Lima et al., 2019 [Publ 3901], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 27 0 0 -- 
-- Pyganodon_grandis **

Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M

iTtTbiriSoTroTroSiD
1 1 0 1 9 1 1 -- 
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Whelan et al., 2023 [Publ 4287], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 27 26 11 -- 
  +-------------------- Utterbackiana    
  |                                      
  |                 +-- Utterbackia      
--+  +--------------+                    
  |  |              +-- Pyganodon_grandis **
  |  |                                   
  +--+           +----- Alasmidonta      
     |  +--------+                       
     |  |        |  +-- Arcidens         
     |  |        +--+                    
     +--+           +-- Pseudodontoideus 
        |                                
        |  +----------- Alasminota       
        |  |                             
        +--+        +-- Anodontoides     
           |  +-----+                    
           |  |     +-- Strophitus       
           +--+                          
              |  +----- Sulcularia       
              +--+                       
                 |  +-- Platynaias       
                 +--+                    
                    +-- Prolasmidonta

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iTtTbiriSoTroTroSiD
1 1 0 1 30 1 1 -- 
  +-- Strophitus       
--+                    
  +-- Pyganodon_grandis **

Curole & Kocher, 2005 [Publ 2682], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 -- 
-- Pyganodon_grandis **

Huang et al., 2013 [Publ 3629], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 15 3 2 -- 
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
1 1 0 1 75 4 4 -- 
        +-- Pyganodon_grandis **
  +-----+                    
  |     +-- Simpsonaias      
--+                          
  |  +----- Strophitus       
  +--+                       
     |  +-- Alasmidonta      
     +--+                    
        +-- Lasmigona

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 45 3 2 -- 
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 68 5 4 -- 
        +-- Pyganodon_grandis **
  +-----+                    
  |     +-- Utterbackia      
--+                          
  |  +----- Prolasmidonta    
  +--+                       
     |  +-- Platynaias       
     +--+                    
        +-- Alasmidonta

Soroka, 2008 [Publ 3695], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 3 0 0 -- 
-- Pyganodon_grandis **

Hewitt et al., 2018 [Publ 3758], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 8 5 5 -- 
  +-------- Pyganodon_grandis **
  |                          
  |     +-- Platynaias       
--+-----+                    
  |     +-- Strophitus       
  |                          
  |  +----- Lasmigona        
  +--+                       
     |  +-- Arcidens         
     +--+                    
        +-- Pterosyna

Manendo et al., 2008 [Publ 2669], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 12 1 1 -- 
  +-- Platynaias       
--+                    
  +-- Pyganodon_grandis **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
1 1 0 1 90 2 2 -- 
  +-- Pyganodon_grandis **
--+                    
  +-- Strophitus

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 73 3 2 -- 
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia

Huang et al., 2018 [Publ 3828], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 41 3 2 -- 
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia

Burzynski et al., 2017 [Publ 3774], fig. 1-bottom

iTtTbiriSoTroTroSiD
1 1 0 1 12 1 1 -- 
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Zanatta & Murphy, 2006 [Publ 2672], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 56 0 0 -- 
-- Pyganodon_grandis **

Riccardi et al., 2020 [Publ 4089], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 18 3 2 -- 
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
1 1 0 1 91 6 5 -- 
           +-- Pyganodon_grandis **
  +--------+                    
  |        +-- Utterbackia      
--+                             
  |  +-------- Pterosyna        
  +--+                          
     |  +----- Prolasmidonta    
     +--+                       
        |  +-- Alasmidonta      
        +--+                    
           +-- Platynaias

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 22 3 2 -- 
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia

Wang et al., 2025 [Publ 4488], fig. 3B

iTtTbiriSoTroTroSiD
1 1 0 1 64 6 5 -- 
           +-- Pyganodon_grandis **
  +--------+                    
  |        +-- Utterbackia      
--+                             
  |  +-------- Pterosyna        
  +--+                          
     |  +----- Prolasmidonta    
     +--+                       
        |  +-- Alasmidonta      
        +--+                    
           +-- Platynaias

Sheng et al., 2014 [Publ 3704], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 7 0 0 -- 
-- Pyganodon_grandis **

Amaro et al., 2017 [Publ 3791], fig. 3A

iTtTbiriSoTroTroSiD
1 1 0 1 11 3 2 -- 
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 48 3 2 -- 
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 55 3 2 -- 
  +----- Lasmigona        
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia

Chong et al., 2008 [Publ 2668], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 17 7 7 -- 
-- Pyganodon_fragilis **

Fonseca et al., 2016 [Publ 3775], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 30 3 2 -- 
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iTtTbiriSoTroTroSiD
1 1 0 1 19 1 1 -- 
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 56 0 0 -- 
-- Pyganodon_grandis **

Dai et al., 2024 [Publ 4386], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 41 1 1 -- 
  +-- Pyganodon_grandis **
--+                    
  +-- Utterbackia

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 67 5 4 -- 
        +-- Pyganodon_grandis **
  +-----+                    
  |     +-- Utterbackia      
--+                          
  |  +----- Alasmidonta      
  +--+                       
     |  +-- Platynaias       
     +--+                    
        +-- Prolasmidonta

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 52 2 2 -- 
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia

Graf, 2002 [Publ 2651], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 17 1 1 -- 
  +-- Alasmidonta      
--+                    
  +-- Pyganodon_grandis **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
1 1 0 1 198 6 5 -- 
  +-- Pyganodon_grandis **
--+                    
  +-- Utterbackia

Hoeh et al., 2009 [Publ 2716], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 45 3 3 -- 
  +-------- Lasmigona        
--+                          
  |  +----- Alasmidonta      
  +--+                       
     |  +-- Strophitus       
     +--+                    
        +-- Pyganodon_grandis **

Amaro et al., 2017 [Publ 3791], fig. 3A-M

iTtTbiriSoTroTroSiD
1 1 0 1 6 1 1 -- 
  +-- Pyganodon_grandis **
--+                    
  +-- Utterbackia

Zhou et al., 2016 [Publ 3710], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 19 3 2 -- 
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iTtTbiriSoTroTroSiD
1 1 0 1 30 1 1 -- 
  +-- Pyganodon_grandis **
--+                    
  +-- Strophitus

Boyer et al., 2011 [Publ 2718], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 37 8 8 -- 
  +-------------- Strophitus       
  |                                
  |        +----- Pterosyna        
--+  +-----+                       
  |  |     |  +-- Lasmigona        
  |  |     +--+                    
  +--+        +-- Arcidens         
     |                             
     |  +-------- Utterbackia      
     |  |                          
     +--+-------- Alasmidonta      
        |                          
        |  +----- Pyganodon_grandis **
        +--+                       
           |  +-- Anodontoides     
           +--+                    
              +-- Platynaias

Wu et al., 2023 [Publ 4296], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 22 2 2 -- 
  +----- Lasmigona        
--+                       
  |  +-- Strophitus       
  +--+                    
     +-- Pyganodon_grandis **

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
1 1 0 1 38 0 0 -- 
-- Pyganodon_grandis **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 200 6 5 -- 
  +-- Pyganodon_grandis **
--+                    
  +-- Utterbackia

Froufe et al., 2015 [Publ 3664], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 7 1 1 -- 
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
1 1 0 1 35 1 1 -- 
  +-- Pyganodon_grandis **
--+                    
  +-- Lasmigona

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 53 3 3 -- 
     +-- Pyganodon_grandis **
  +--+                    
  |  +-- Utterbackia      
--+                       
  |  +-- Alasmidonta      
  +--+                    
     +-- Prolasmidonta

Hewitt et al., 2019 [Publ 3962], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 9 0 0 -- 
-- Pyganodon_grandis **

Plazzi et al., 2011 [Publ 3676], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 17 0 0 -- 
-- Pyganodon_grandis **

Marshall et al., 2014 [Publ 3593], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 13 0 0 -- 
-- Pyganodon_grandis **

Burzynski & Soroka, 2018 [Publ 3862], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 3 1 1 -- 
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Wen et al., 2017 [Publ 3795], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 5 1 1 -- 
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 67 5 4 -- 
        +-- Pyganodon_grandis **
  +-----+                    
  |     +-- Utterbackia      
--+                          
  |  +----- Alasmidonta      
  +--+                       
     |  +-- Prolasmidonta    
     +--+                    
        +-- Platynaias

Guerra et al., 2019 [Publ 4168], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 21 1 1 -- 
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Wei et al., 2016 [Publ 3665], fig. 0

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Pyganodon_grandis **

Krebs, 2004 [Publ 2692], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 4 1 1 -- 
  +-- Lasmigona        
--+                    
  +-- Pyganodon_grandis **

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
1 1 0 1 47 3 2 -- 
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia

Wu et al., 2022 [Publ 4238], fig. 2e

iTtTbiriSoTroTroSiD
1 1 0 1 17 1 1 -- 
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Chen et al., 2012 [Publ 3624], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 6 0 0 -- 
-- Pyganodon_grandis **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
1 1 0 1 138 4 4 -- 
  +----- Pyganodon_grandis **
  |                       
  +----- Simpsonaias      
--+                       
  +----- Strophitus       
  |                       
  |  +-- Alasmidonta      
  +--+                    
     +-- Lasmigona

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

iTtTbiriSoTroTroSiD
1 1 0 1 22 1 1 -- 
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Zanatta & Harris, 2013 [Publ 3530], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 4 0 0 -- 
-- Pyganodon_grandis **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
1 1 0 1 73 6 4 -- 
        +-- Pyganodon_grandis **
  +-----+                    
  |     +-- Utterbackia      
--+                          
  |  +----- Prolasmidonta    
  +--+                       
     |  +-- Alasmidonta      
     +--+                    
        +-- Platynaias

Chase et al., 2018 [Publ 3861], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 10 1 1 -- 
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Cha et al., 2018 [Publ 3916], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 37 3 2 -- 
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia

Amaro et al., 2017 [Publ 3791], fig. 2-M

iTtTbiriSoTroTroSiD
1 1 0 1 9 2 2 -- 
-- Pyganodon_grandis **

Soroka & Burzynski, 2018 [Publ 3762], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 6 1 1 -- 
  +-- Pyganodon_grandis **
--+                    
  +-- Utterbackia

Aunins et al., 2018 [Publ 3863], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 39 4 3 -- 
     +-- Prolasmidonta    
  +--+                    
  |  +-- Alasmidonta      
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
1 1 0 1 95 6 5 -- 
        +-- Pyganodon_grandis **
  +-----+                    
  |     +-- Utterbackia      
  |                          
--+     +-- Alasmidonta      
  |  +--+                    
  |  |  +-- Platynaias       
  +--+                       
     |  +-- Prolasmidonta    
     +--+                    
        +-- Pterosyna

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 4 2 -- 
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iTtTbiriSoTroTroSiD
1 1 0 1 30 1 1 -- 
-- Pyganodon_grandis **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
1 1 0 1 79 4 4 -- 
        +-- Pyganodon_grandis **
  +-----+                    
  |     +-- Simpsonaias      
--+                          
  |  +----- Strophitus       
  +--+                       
     |  +-- Alasmidonta      
     +--+                    
        +-- Lasmigona

Zanatta & Murphy, 2006 [Publ 2672], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 55 0 0 -- 
-- Pyganodon_grandis **

Wu et al., 2020 [Publ 4147], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 29 2 1 -- 
  +-- Pyganodon_grandis **
--+                    
  +-- Utterbackia

Fonseca et al., 2016 [Publ 3775], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 8 1 1 -- 
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Huang et al., 2013 [Publ 3629], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 5 1 1 -- 
  +-- Pyganodon_grandis **
--+                    
  +-- Utterbackia

Inoue et al., 2013 [Publ 3579], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 8 0 0 -- 
-- Pyganodon_grandis **

Feng et al., 2016 [Publ 3800], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 13 1 1 -- 
-- Pyganodon_grandis **

Campbell et al., 2005 [Publ 2660], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 101 3 3 -- 
-- Pyganodon_grandis **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
1 1 0 1 75 4 4 -- 
        +-- Alasmidonta      
  +-----+                    
  |     +-- Lasmigona        
--+                          
  |  +----- Pyganodon_grandis **
  +--+                       
     |  +-- Strophitus       
     +--+                    
        +-- Simpsonaias

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

iTtTbiriSoTroTroSiD
1 1 0 1 24 3 3 -- 
  +----- Lasmigona        
  |                       
--+  +-- Strophitus       
  |  |                    
  +--+-- Alasmidonta      
     |                    
     +-- Pyganodon_grandis **

Wu et al., 2022 [Publ 4238], fig. 2d

iTtTbiriSoTroTroSiD
1 1 0 1 38 3 2 -- 
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia

Hoeh et al., 2009 [Publ 2716], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 46 3 3 -- 
  +-------- Lasmigona        
--+                          
  |  +----- Alasmidonta      
  +--+                       
     |  +-- Strophitus       
     +--+                    
        +-- Pyganodon_grandis **

Inoue et al., 2013 [Publ 3528], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 16 0 0 -- 
-- Pyganodon_grandis **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
1 1 0 1 56 0 0 -- 
-- Pyganodon_grandis **

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 82 4 4 -- 
  +----------- Pyganodon_grandis **
--+                             
  |  +-------- Simpsonaias      
  +--+                          
     |  +----- Strophitus       
     +--+                       
        |  +-- Alasmidonta      
        +--+                    
           +-- Lasmigona

Soroka, 2008 [Publ 3695], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 3 0 0 -- 
-- Pyganodon_grandis **

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
1 1 0 1 73 6 4 -- 
        +-- Pyganodon_grandis **
  +-----+                    
  |     +-- Utterbackia      
--+                          
  |  +----- Prolasmidonta    
  +--+                       
     |  +-- Alasmidonta      
     +--+                    
        +-- Platynaias

Burzynski & Soroka, 2018 [Publ 3862], fig. 5b

iTtTbiriSoTroTroSiD
1 1 0 1 9 1 1 -- 
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Boyer et al., 2011 [Publ 2718], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 34 5 5 -- 
  +----- Alasmidonta      
  |                       
  +----- Pyganodon_grandis **
  |                       
--+  +-- Pterosyna        
  +--+                    
  |  +-- Lasmigona        
  |                       
  |  +-- Platynaias       
  +--+                    
     +-- Strophitus

Smith, 2021 [Publ 4331], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 19 1 1 -- 
  +-- Pyganodon_grandis **
--+                    
  +-- Utterbackia

Chen et al., 2020 [Publ 4170], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 33 5 4 -- 
  +----- Platynaias       
--+                       
  |  +-- Utterbackia      
  +--+                    
     +-- Pyganodon_grandis **

Guerra et al., 2019 [Publ 4168], fig. 2-M

iTtTbiriSoTroTroSiD
1 1 0 1 21 1 1 -- 
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Amaro et al., 2017 [Publ 3791], fig. 3B

iTtTbiriSoTroTroSiD
1 1 0 1 11 3 2 -- 
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia

Curole & Kocher, 2005 [Publ 2682], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 11 1 1 -- 
  +-- Utterbackiana    
--+                    
  +-- Pyganodon_grandis **

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 43 3 2 -- 
  +----- Pterosyna        
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia

Froufe et al., 2019 [Publ 3936], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 18 1 1 -- 
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Inoue et al., 2014 [Publ 3635], fig. 3A

iTtTbiriSoTroTroSiD
1 1 0 1 19 13 8 -- 
     +-------- Pseudodontoideus 
  +--+                          
  |  |  +----- Lasmigona        
  |  +--+                       
  |     |  +-- Pterosyna        
--+     +--+                    
  |        +-- Arcidens         
  |                             
  |  +-------- Strophitus       
  |  |                          
  +--+     +-- Pyganodon_grandis **
     |  +--+                    
     |  |  +-- Utterbackia      
     +--+                       
        |  +-- Platynaias       
        +--+                    
           +-- Alasmidonta

Liu et al., 2023 [Publ 4310], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 23 2 2 -- 
  +-- Pyganodon_grandis **
--+                    
  +-- Strophitus

Guerra et al., 2017 [Publ 3772], fig. 2a-M

iTtTbiriSoTroTroSiD
1 1 0 1 7 1 1 -- 
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 31 3 2 -- 
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 39 1 1 -- 
  +-- Pyganodon_grandis **
--+                    
  +-- Lasmigona

Lopes-Lima et al., 2019 [Publ 3901], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 25 0 0 -- 
-- Pyganodon_grandis **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
1 1 0 1 200 6 5 -- 
-- Pyganodon_grandis **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
1 1 0 1 138 4 4 -- 
        +-- Lasmigona        
  +-----+                    
  |     +-- Alasmidonta      
--+                          
  |  +----- Simpsonaias      
  +--+                       
     |  +-- Pyganodon_grandis **
     +--+                    
        +-- Strophitus

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
1 1 0 1 121 4 4 -- 
  +----- Pyganodon_grandis **
  |                       
  +----- Simpsonaias      
--+                       
  +----- Strophitus       
  |                       
  |  +-- Alasmidonta      
  +--+                    
     +-- Lasmigona

Krebs, 2004 [Publ 2692], fig. 2-M

iTtTbiriSoTroTroSiD
1 1 0 1 3 1 1 -- 
  +-- Lasmigona        
--+                    
  +-- Pyganodon_grandis **

Wu et al., 2024 [Publ 4379], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 53 6 5 -- 
           +-- Pyganodon_grandis **
  +--------+                    
  |        +-- Utterbackia      
--+                             
  |  +-------- Pterosyna        
  +--+                          
     |  +----- Prolasmidonta    
     +--+                       
        |  +-- Alasmidonta      
        +--+                    
           +-- Platynaias

Hou et al., 2025 [Publ 4445], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 52 6 5 -- 
           +-- Pyganodon_grandis **
  +--------+                    
  |        +-- Utterbackia      
--+                             
  |  +-------- Pterosyna        
  +--+                          
     |  +----- Prolasmidonta    
     +--+                       
        |  +-- Platynaias       
        +--+                    
           +-- Alasmidonta

Froufe et al., 2015 [Publ 3664], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 11 1 1 -- 
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Wu et al., 2018 [Publ 4143], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 21 2 1 -- 
  +-- Pyganodon_grandis **
--+                    
  +-- Utterbackia

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 2 2 -- 
  +----- Strophitus       
--+                       
  |  +-- Platynaias       
  +--+                    
     +-- Pyganodon_grandis **

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 36 0 0 -- 
-- Pyganodon_grandis **