Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Anodontini | Alasmidontina

genus Pyganodon Crosse & Fischer, 1894: cladomics

6 Recent valid species (6 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 6 (6 total)
analyzed in 191 cladograms in 115 publications
maximum number of terminals in an analysis = 5 (mean = 1.34, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 34
maximum number of included terminals = 5, (mean = 2.91 median = 2)
proportion of cladograms that support basic monophyly = 0.82


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Pyganodon cladogram distances
PCoA Ordination

NJ Tree of Pyganodon cladogram distances
NJ Tree


Cladograms

Hoeh, 1990 [Publ 2717], fig. 2

** Top Monophyly Cladogram **

cstiTtTbiriSoTroTroSiD
0.857 5 5 3 5 10 8 5 **
  +-------------- Utterbackia        
--+                                  
  |  +----------- Pyganodon_lacustris **
  +--+                               
     |  +-------- Pyganodon_grandis **  
     +--+                            
        |  +----- Pyganodon_fragilis ** 
        +--+                         
           |  +-- Pyganodon_cataracta **
           +--+                      
              +-- Pyganodon_gibbosa **  

Perkins et al., 2025 [Publ 4447], fig. 2_mtDNA-ITS

** Top Sister-Group Cladogram **

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 35 32 14 --
     +-------------- Utterbackiana    
  +--+                                
  |  |  +----------- Alasminota       
  |  +--+                             
  |     |  +-------- Pseudodontoideus 
  |     +--+                          
  |        |  +----- Lasmigona        
  |        +--+                       
  |           |  +-- Pterosyna        
  |           +--+                    
--+              +-- Arcidens         
  |                                   
  |              +-- Prolasmidonta    
  |           +--+                    
  |           |  +-- Platynaias       
  |     +-----+                       
  |     |     |  +-- Ligodonta        
  |     |     +--+                    
  |     |        +-- Alasmidonta      
  +-----+                             
        |        +-- Anodontoides     
        |  +-----+                    
        |  |     +-- Strophitus       
        +--+                          
           |  +----- Sulcularia       
           +--+                       
              |  +-- Pyganodon_grandis **
              +--+                    
                 +-- Utterbackia      

Hoeh, 1990 [Publ 2717], fig. 3

cstiTtTbiriSoTroTroSiD
0.429 5 15 2 5 10 8 5 15.0
  +----- Pyganodon_fragilis **         
  |                                 
  +----- Utterbackia_imbecillis     
  |                                 
  +----- Utterbackia_peggyae        
  |                                 
  +----- Anodonta                   
  |                                 
  +----- Brachyanodon               
  |                                 
  +----- Utterbackiana_couperiana__1
  |                                 
--+----- Utterbackiana_couperiana__2
  |                                 
  +----- Utterbackiana_suborbiculata
  |                                 
  |  +-- Lasmigona                  
  +--+                              
  |  +-- Utterbackiana_implicata    
  |                                 
  |  +-- Pterosyna                  
  |  |                              
  |  +-- Platynaias                 
  |  |                              
  |  +-- Pyganodon_cataracta **        
  +--+                              
     +-- Pyganodon_gibbosa **          
     |                              
     +-- Pyganodon_grandis **          
     |                              
     +-- Pyganodon_lacustris **        

Hoeh, 1990 [Publ 2717], fig. 4

cstiTtTbiriSoTroTroSiD
0.857 5 5 3 5 10 8 5 4.0
  +----------- Utterbackia        
  |                               
--+  +-------- Pyganodon_fragilis ** 
  |  |                            
  +--+     +-- Pyganodon_cataracta **
     |  +--+                      
     |  |  +-- Pyganodon_gibbosa **  
     +--+                         
        |  +-- Pyganodon_lacustris **
        +--+                      
           +-- Pyganodon_grandis **  

Zanatta et al., 2007 [Publ 2698], fig. 1A

cstiTtTbiriSoTroTroSiD
0.857 5 5 0 5 11 9 5 3.0
  +----- Utterbackia        
  |                         
  |  +-- Pyganodon_gibbosa **  
--+  |                      
  |  +-- Pyganodon_cataracta **
  |  |                      
  +--+-- Pyganodon_fragilis ** 
     |                      
     +-- Pyganodon_grandis **  
     |                      
     +-- Pyganodon_lacustris **

Zanatta et al., 2007 [Publ 2698], fig. 1B

cstiTtTbiriSoTroTroSiD
0.857 5 5 3 5 11 9 5 0.0
  +-------------- Utterbackia        
--+                                  
  |  +----------- Pyganodon_lacustris **
  +--+                               
     |  +-------- Pyganodon_grandis **  
     +--+                            
        |  +----- Pyganodon_fragilis ** 
        +--+                         
           |  +-- Pyganodon_gibbosa **  
           +--+                      
              +-- Pyganodon_cataracta **

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cstiTtTbiriSoTroTroSiD
0.933 4 4 3 4 629 33 13 4.0
     +-------------- Utterbackia           
  +--+                                     
  |  |  +----------- Pyganodon_grandis **     
  |  +--+                                  
  |     |  +-------- Pyganodon_lacustris **   
  |     +--+                               
  |        |  +----- Pyganodon_cataracta__2 **
--+        +--+                            
  |           |  +-- Pyganodon_cataracta__1 **
  |           +--+                         
  |              +-- Pyganodon_fragilis **    
  |                                        
  |  +-------------- Utterbackiana         
  |  |                                     
  +--+  +----------- Simpsonaias           
     |  |                                  
     |  |        +-- Alasminota            
     +--+  +-----+                         
        |  |     +-- Alasmidonta           
        |  |                               
        +--+     +-- Prolasmidonta         
           |  +--+                         
           |  |  +-- Platynaias            
           +--+                            
              |  +-- Anodontoides          
              +--+                         
                 +-- Strophitus            

Inoue et al., 2014 [Publ 3635], fig. 3B

cstiTtTbiriSoTroTroSiD
0.750 4 7 2 4 4 3 2 7.0
  +-------- Amblema            
  |                            
  |  +----- Pyganodon_grandis **  
--+  |                         
  |  +----- Pyganodon_cataracta **
  |  |                         
  +--+----- Pyganodon_fragilis ** 
     |                         
     +----- Pyganodon **          
     |                         
     |  +-- Platynaias         
     +--+                      
        +-- Arcidens           

Hoeh et al., 1995 [Publ 3391], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 4 4 1 4 3 3 1 2.0
  +-------- Utterbackia        
  |                            
--+  +----- Pyganodon_lacustris **
  |  |                         
  +--+----- Pyganodon_grandis **  
     |                         
     |  +-- Pyganodon_cataracta **
     +--+                      
        +-- Pyganodon_fragilis ** 

Cyr et al., 2007 [Publ 2745], fig. 2A

cstiTtTbiriSoTroTroSiD
1.000 4 4 2 4 1 0 0 3.0
  +----------- Lampsilis          
  |                               
--+  +-------- Pyganodon_grandis **  
  |  |                            
  +--+-------- Pyganodon__2 **       
     |                            
     |  +----- Pyganodon__1 **       
     +--+                         
        |  +-- Pyganodon_cataracta **
        +--+                      
           +-- Pyganodon_fragilis ** 

Cyr et al., 2007 [Publ 2745], fig. 2B

cstiTtTbiriSoTroTroSiD
1.000 4 4 3 4 1 0 0 3.0
  +----------- Brachyanodon       
  |                               
--+        +-- Pyganodon_grandis **  
  |  +-----+                      
  |  |     +-- Pyganodon__1 **       
  +--+                            
     |  +----- Pyganodon__2 **       
     +--+                         
        |  +-- Pyganodon_cataracta **
        +--+                      
           +-- Pyganodon_fragilis ** 

Cyr et al., 2007 [Publ 2745], fig. 2C

cstiTtTbiriSoTroTroSiD
1.000 4 4 3 4 1 0 0 7.0
  +----------- Pseudanodonta         
  |                                  
  |        +-- Pyganodon_grandis **     
--+  +-----+                         
  |  |     +-- Pyganodon__1 **          
  |  |                               
  +--+  +----- Pyganodon__3 **          
     |  |                            
     |  |  +-- Pyganodon_cataracta__1 **
     +--+  |                         
        |  +-- Pyganodon_fragilis__1 ** 
        |  |                         
        +--+-- Pyganodon__2 **          
           |                         
           +-- Pyganodon_cataracta__2 **
           |                         
           +-- Pyganodon_fragilis__2 ** 

Cyr et al., 2007 [Publ 2745], fig. 2D

cstiTtTbiriSoTroTroSiD
1.000 4 4 2 4 0 0 0 3.0
  +-------- Pyganodon_grandis **  
  |                            
--+-------- Pyganodon__2 **       
  |                            
  |  +----- Pyganodon__1 **       
  +--+                         
     |  +-- Pyganodon_cataracta **
     +--+                      
        +-- Pyganodon_fragilis ** 

Stewart et al., 2021 [Publ 4451], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 3 3 1 3 13 7 5 4.0
              +-- Alasmidonta        
  +-----------+                      
  |           +-- Pterosyna          
  |                                  
--+           +-- Strophitus         
  |  +--------+                      
  |  |        +-- Platynaias         
  +--+                               
     |  +-------- Utterbackia        
     +--+                            
        |  +----- Pyganodon_grandis **  
        +--+                         
           |  +-- Pyganodon_cataracta **
           +--+                      
              +-- Pyganodon_lacustris **

Krebs et al., 2019 [Publ 4092], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 3 3 1 3 0 0 0 2.0
  +----- Pyganodon_grandis **  
--+                         
  |  +-- Pyganodon_fragilis ** 
  +--+                      
     +-- Pyganodon_cataracta **

Whelan et al., 2023 [Publ 4287], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 2 2 1 2 39 31 13 --
        +-------- Utterbackia         
  +-----+                             
  |     |  +----- Pyganodon_grandis__2 **
  |     +--+                          
  |        |  +-- Pyganodon_grandis__1 **
  |        +--+                       
  |           +-- Pyganodon_lacustris ** 
  |                                   
--+     +-------- Utterbackiana       
  |     |                             
  |  +--+     +-- Lasmigona           
  |  |  |  +--+                       
  |  |  |  |  +-- Pseudodontoideus    
  |  |  +--+                          
  |  |     |  +-- Pterosyna           
  |  |     +--+                       
  +--+        +-- Arcidens            
     |                                
     |     +----- Sulcularia          
     |  +--+                          
     |  |  |  +-- Anodontoides        
     |  |  +--+                       
     |  |     +-- Strophitus          
     +--+                             
        |     +-- Prolasmidonta       
        |  +--+                       
        |  |  +-- Platynaias          
        +--+                          
           |  +-- Alasminota          
           +--+                       
              +-- Alasmidonta         

Whelan et al., 2023 [Publ 4287], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 25 23 10 --
  +-------------------- Sulcularia         
  |                                        
  |           +-------- Utterbackiana      
--+  +--------+                            
  |  |        |  +----- Alasminota         
  |  |        +--+                         
  |  |           |  +-- Arcidens           
  +--+           +--+                      
     |              +-- Pseudodontoideus   
     |                                     
     |  +-------------- Alasmidonta        
     |  |                                  
     +--+  +----------- Platynaias         
        |  |                               
        +--+  +-------- Strophitus         
           |  |                            
           +--+     +-- Pyganodon_grandis **  
              |  +--+                      
              |  |  +-- Pyganodon_cataracta **
              +--+                         
                 |  +-- Utterbackia        
                 +--+                      
                    +-- Anodontoides       

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

cstiTtTbiriSoTroTroSiD
0.455 2 20 0 2 103 14 9 22.0
                                                     +-- Alasmidonta_marginata      
                                                     |                              
                                                     +-- Prolasmidonta              
                                                     |                              
                                                     +-- Alasmidonta_undulata       
                                                     |                              
                                                     +-- Beringiana                 
                                                     |                              
                                                     +-- Brachyanodon_kennerlyi     
                                                     |                              
                                                     +-- Brachyanodon_nuttalliana   
                                                     |                              
                                                     +-- Brachyanodon_oregonensis   
                                                     |                              
                                                     +-- Arcidens_confragosus       
                                                     |                              
                                                     +-- Arcidens_wheeleri          
                                                     |                              
                                                     +-- Pterosyna                  
                                                     |                              
  +--------------------------------------------------+-- Platynaias_compressa       
  |                                                  |                              
  |                                                  +-- Lasmigona                  
  |                                                  |                              
  |                                                  +-- Platynaias_decorata        
  |                                                  |                              
  |                                                  +-- Platynaias_subviridis      
  |                                                  |                              
  |                                                  +-- Pyganodon_grandis **          
  |                                                  |                              
  |                                                  +-- Pyganodon_lacustris **        
  |                                                  |                              
  |                                                  +-- Pseudodontoideus           
  |                                                  |                              
  |                                                  +-- Strophitus                 
  |                                                  |                              
--+                                                  +-- Utterbackia_imbecillis     
  |                                                  |                              
  |                                                  +-- Utterbackia_peninsularis__1
  |                                                  |                              
  |                                                  +-- Utterbackia_peninsularis__2
  |                                                                                 
  |  +-------------------------------------------------- Gonidea                    
  |  |                                                                              
  |  |  +----------------------------------------------- Reginaia                   
  |  |  |                                                                           
  |  |  +----------------------------------------------- Amblema                    
  |  |  |                                                                           
  |  |  |                                         +----- Popenaias                  
  |  |  |                                         |                                 
  |  |  |                                         |  +-- Dromus                     
  |  |  |                                         |  |                              
  |  |  |                                         |  +-- Epioblasma                 
  +--+  |                                         |  |                              
     |  +-----------------------------------------+  +-- Glebula                    
     |  |                                         |  |                              
     |  |                                         |  +-- Lampsilis_ovata            
     |  |                                         |  |                              
     |  |                                         |  +-- Lampsilis_virescens        
     |  |                                         |  |                              
     |  |                                         |  +-- Lemiox                     
     |  |                                         +--+                              
     |  |                                            +-- Obliquaria                 
     |  |                                            |                              
     +--+                                            +-- Obovaria_olivaria          
        |                                            |                              
        |                                            +-- Obovaria_unicolor          
        |                                            |                              
        |                                            +-- Plectomerus                
        |                                            |                              
        |                                            +-- Potamilus_alatus           
        |                                            |                              
        |                                            +-- Potamilus_capax            
        |                                                                           
        |                                         +----- Uniomerus                  
        |                                         |                                 
        |                                         |  +-- Cyclonaias                 
        |  +--------------------------------------+  |                              
        |  |                                      |  +-- Pustulosa_pustulosa__1     
        |  |                                      |  |                              
        |  |                                      |  +-- Theliderma                 
        |  |                                      +--+                              
        |  |                                         +-- Pustulosa_pustulosa__2     
        |  |                                         |                              
        +--+                                         +-- Quadrula_quadrula          
           |                                         |                              
           |                                         +-- Quadrula_verrucosa         
           |                                                                        
           |  +----------------------------------------- Elliptoideus               
           |  |                                                                     
           |  |  +-------------------------------------- Parvaspina                 
           |  |  |                                                                  
           +--+  |  +----------------------------------- Plethobasus                
              |  |  |                                                               
              |  |  +----------------------------------- Eurynia                    
              |  |  |                                                               
              |  |  +----------------------------------- Hemistena                  
              +--+  |                                                               
                 |  +----------------------------------- Elliptio                   
                 |  |                                                               
                 |  |  +-------------------------------- Pleuronaia                 
                 |  |  |                                                            
                 |  |  |                             +-- Pleurobema_cordatum        
                 |  |  |  +--------------------------+                              
                 +--+  |  |                          +-- Fusconaia                  
                    |  |  |                                                         
                    |  |  |  +-------------------------- Pleurobema_plenum          
                    |  |  |  |                                                      
                    |  |  |  |  +----------------------- Pleurobema_stabile         
                    |  |  |  |  |                                                   
                    |  |  |  |  |                 +----- Pleurobema_pyriforme       
                    |  |  +--+  |              +--+                                 
                    |  |  |  |  |              |  |  +-- Pleurobema_georgianum__2   
                    |  |  |  |  |              |  +--+                              
                    |  |  |  |  |  +-----------+     +-- Pleurobema_athearni        
                    |  |  |  |  |  |           |                                    
                    +--+  |  +--+  |           |  +----- Pleurobema_decisum         
                       |  |     |  |           |  |                                 
                       |  |     |  |           +--+  +-- Pleurobema_georgianum__1   
                       |  |     |  |              |  |                              
                       |  |     |  |              +--+-- Pleurobema_hanleyianum__1  
                       |  |     |  |                 |                              
                       |  |     |  |                 +-- Pleurobema_hanleyianum__2  
                       |  |     |  |                                                
                       |  |     +--+     +-------------- Pleurobema_beadleianum__2  
                       |  |        |     |                                          
                       |  |        |  +--+  +----------- Pleurobema_strodeanum__2   
                       |  |        |  |  |  |                                       
                       |  |        |  |  +--+----------- Pleurobema_rubellum        
                       |  |        |  |     |                                       
                       |  |        |  |     |  +-------- Pleurobema_beadleianum__1  
                       |  |        |  |     +--+                                    
                       +--+        |  |        |  +----- Pleurobema_taitianum       
                          |        |  |        +--+                                 
                          |        |  |           |  +-- Pleurobema_perovatum       
                          |        +--+           +--+                              
                          |           |              +-- Pleurobema_strodeanum__1   
                          |           |                                             
                          |           |           +----- Pleurobema_oviforme__6     
                          |           |           |                                 
                          |           |        +--+----- Pleurobema_clava__3        
                          |           |        |  |                                 
                          |           |        |  |  +-- Pleurobema_oviforme__5     
                          |           |        |  +--+                              
                          |           +--------+     +-- Pleurobema_clava__2        
                          |                    |                                    
                          |                    |  +----- Pleurobema_oviforme__2     
                          |                    |  |                                 
                          |                    |  +----- Pleurobema_oviforme__3     
                          |                    +--+                                 
                          |                       +----- Pleurobema_oviforme__4     
                          |                       |                                 
                          |                       |  +-- Pleurobema_clava__1        
                          |                       +--+                              
                          |                          +-- Pleurobema_oviforme__1     
                          |                                                         
                          |           +----------------- Pleurobema_riddellii       
                          |           |                                             
                          |           |  +-------------- Pleurobema_sintoxia__34    
                          |           |  |                                          
                          |           |  |  +----------- Pleurobema_sintoxia__31    
                          |           |  |  |                                       
                          +-----------+  |  +----------- Pleurobema_sintoxia__32    
                                      |  |  |                                       
                                      |  |  +----------- Pleurobema_sintoxia__33    
                                      |  |  |                                       
                                      |  |  |  +-------- Pleurobema__4              
                                      |  |  |  |                                    
                                      |  |  |  +-------- Pleurobema__5              
                                      +--+  |  |                                    
                                         |  |  +-------- Pleurobema_sintoxia__2     
                                         |  |  |                                    
                                         |  |  +-------- Pleurobema_sintoxia__3     
                                         |  |  |                                    
                                         |  |  +-------- Pleurobema__8              
                                         |  |  |                                    
                                         |  |  +-------- Pleurobema__9              
                                         |  |  |                                    
                                         |  |  +-------- Pleurobema__10             
                                         |  |  |                                    
                                         |  |  +-------- Pleurobema_sintoxia__5     
                                         +--+  |                                    
                                            |  +-------- Pleurobema_sintoxia__6     
                                            |  |                                    
                                            |  +-------- Pleurobema_sintoxia__7     
                                            |  |                                    
                                            |  +-------- Pleurobema_sintoxia__8     
                                            |  |                                    
                                            |  +-------- Pleurobema_sintoxia__9     
                                            |  |                                    
                                            |  +-------- Pleurobema_sintoxia__10    
                                            |  |                                    
                                            |  +-------- Pleurobema_sintoxia__11    
                                            |  |                                    
                                            |  +-------- Pleurobema_sintoxia__12    
                                            |  |                                    
                                            |  +-------- Pleurobema_sintoxia__13    
                                            |  |                                    
                                            |  +-------- Pleurobema_sintoxia__14    
                                            |  |                                    
                                            |  +-------- Pleurobema_sintoxia__15    
                                            |  |                                    
                                            |  +-------- Pleurobema_sintoxia__16    
                                            +--+                                    
                                               +-------- Pleurobema_sintoxia__17    
                                               |                                    
                                               +-------- Pleurobema_sintoxia__18    
                                               |                                    
                                               +-------- Pleurobema_sintoxia__19    
                                               |                                    
                                               +-------- Pleurobema_sintoxia__20    
                                               |                                    
                                               +-------- Pleurobema_sintoxia__21    
                                               |                                    
                                               +-------- Pleurobema_sintoxia__22    
                                               |                                    
                                               +-------- Pleurobema_sintoxia__23    
                                               |                                    
                                               +-------- Pleurobema_sintoxia__24    
                                               |                                    
                                               +-------- Pleurobema_sintoxia__25    
                                               |                                    
                                               +-------- Pleurobema_sintoxia__26    
                                               |                                    
                                               +-------- Pleurobema_sintoxia__27    
                                               |                                    
                                               +-------- Pleurobema_sintoxia__28    
                                               |                                    
                                               +-------- Pleurobema_sintoxia__29    
                                               |                                    
                                               +-------- Pleurobema_sintoxia__30    
                                               |                                    
                                               |     +-- Pleurobema__6              
                                               +-----+                              
                                               |     +-- Pleurobema_sintoxia__1     
                                               |                                    
                                               |     +-- Pleurobema__7              
                                               +-----+                              
                                               |     +-- Pleurobema_sintoxia__4     
                                               |                                    
                                               |  +----- Pleurobema__3              
                                               +--+                                 
                                                  |  +-- Pleurobema__1              
                                                  +--+                              
                                                     +-- Pleurobema__2              

Breton et al., 2011 [Publ 3376], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 36 9 6 --
              +-- Pterosyna          
  +-----------+                      
  |           +-- Lasmigona          
--+                                  
  |  +----------- Strophitus         
  |  |                               
  +--+        +-- Platynaias         
     |  +-----+                      
     |  |     +-- Alasmidonta        
     +--+                            
        |  +----- Utterbackia        
        +--+                         
           |  +-- Pyganodon_lacustris **
           +--+                      
              +-- Pyganodon_grandis **  

Chase et al., 2018 [Publ 3847], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 37 9 6 --
        +----- Utterbackia        
  +-----+                         
  |     |  +-- Pyganodon_lacustris **
  |     +--+                      
  |        +-- Pyganodon_grandis **  
--+                               
  |        +-- Lasmigona          
  |  +-----+                      
  |  |     +-- Pterosyna          
  +--+                            
     |  +----- Alasmidonta        
     +--+                         
        |  +-- Strophitus         
        +--+                      
           +-- Platynaias         

King et al., 1999 [Publ 2699], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 7 7 5 --
           +-- Pyganodon_grandis ** 
  +--------+                     
  |        +-- Pyganodon_fragilis **
  |                              
--+        +-- Lasmigona         
  |  +-----+                     
  |  |     +-- Pterosyna         
  +--+                           
     |  +----- Strophitus        
     +--+                        
        |  +-- Prolasmidonta     
        +--+                     
           +-- Platynaias        

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 28 3 3 --
        +-- Alasmidonta       
  +-----+                     
  |     +-- Lasmigona         
--+                           
  |  +----- Strophitus        
  +--+                        
     |  +-- Pyganodon_grandis ** 
     +--+                     
        +-- Pyganodon_fragilis **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 28 3 3 --
        +-- Alasmidonta       
  +-----+                     
  |     +-- Lasmigona         
--+                           
  |  +----- Strophitus        
  +--+                        
     |  +-- Pyganodon_grandis ** 
     +--+                     
        +-- Pyganodon_fragilis **

Wu et al., 2018 [Publ 3836], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 51 3 3 --
  +-------- Strophitus         
  |                            
--+  +----- Platynaias         
  |  |                         
  +--+----- Alasmidonta        
     |                         
     |  +-- Pyganodon_cataracta **
     +--+                      
        +-- Pyganodon_grandis **  

Rosenberg et al., 1994 [Publ 2719], fig. 3

cstiTtTbiriSoTroTroSiD
0.333 2 10 0 2 17 1 1 11.0
     +-- Elliptio           
     |                      
     +-- Fusconaia          
     |                      
  +--+-- Plectomerus        
  |  |                      
  |  +-- Pleurobema         
  |  |                      
  |  +-- Unio               
  |                         
  |  +-- Pyganodon_cataracta **
--+  |                      
  |  +-- Pyganodon_grandis **  
  |  |                      
  |  +-- Utterbackia        
  |  |                      
  |  +-- Lampsilis_teres    
  |  |                      
  |  +-- Lampsilis_straminea
  +--+                      
     +-- Megalonaias        
     |                      
     +-- Obliquaria         
     |                      
     +-- Theliderma         
     |                      
     +-- Quadrula           
     |                      
     +-- Uniomerus          

Rosenberg et al., 1997 [Publ 2720], fig. 2A

cstiTtTbiriSoTroTroSiD
0.333 2 17 0 2 20 1 1 18.0
  +----- Anodonta           
  |                         
  |  +-- Gonidea            
  |  |                      
  |  +-- Pyganodon_grandis **  
  |  |                      
  |  +-- Utterbackia        
  |  |                      
  |  +-- Pyganodon_cataracta **
--+  |                      
  |  +-- Lampsilis_straminea
  |  |                      
  |  +-- Lampsilis_teres    
  |  |                      
  |  +-- Megalonaias        
  |  |                      
  |  +-- Obliquaria         
  |  |                      
  +--+-- Theliderma         
     |                      
     +-- Quadrula           
     |                      
     +-- Uniomerus          
     |                      
     +-- Amblema            
     |                      
     +-- Elliptio           
     |                      
     +-- Fusconaia          
     |                      
     +-- Plectomerus        
     |                      
     +-- Pleurobema         
     |                      
     +-- Unio               

Rosenberg et al., 1997 [Publ 2720], fig. 2B

cstiTtTbiriSoTroTroSiD
0.333 2 17 0 2 20 1 1 18.0
  +----- Anodonta           
  |                         
  |  +-- Gonidea            
  |  |                      
  |  +-- Pyganodon_grandis **  
  |  |                      
  |  +-- Utterbackia        
  |  |                      
  |  +-- Pyganodon_cataracta **
--+  |                      
  |  +-- Lampsilis_straminea
  |  |                      
  |  +-- Lampsilis_teres    
  |  |                      
  |  +-- Megalonaias        
  |  |                      
  |  +-- Obliquaria         
  |  |                      
  +--+-- Theliderma         
     |                      
     +-- Quadrula           
     |                      
     +-- Uniomerus          
     |                      
     +-- Amblema            
     |                      
     +-- Elliptio           
     |                      
     +-- Fusconaia          
     |                      
     +-- Plectomerus        
     |                      
     +-- Scriptolamprotula  
     |                      
     +-- Unio               

Hoeh et al., 1996 [Publ 2694], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 2 0 0 --
  +----- Fusconaia         
--+                        
  |  +-- Pyganodon_fragilis **
  +--+                     
     +-- Pyganodon_grandis ** 

Hoeh et al., 2002 [Publ 2673], fig. 1-F

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 7 0 0 --
                 +-- Pyganodon_fragilis **
  +--------------+                     
  |              +-- Pyganodon_grandis ** 
--+                                    
  |  +-------------- Fusconaia         
  +--+                                 
     |  +----------- Cyrtonaias        
     +--+                              
        |  +-------- Ortmanniana       
        +--+                           
           |  +----- Lampsilis         
           +--+                        
              |  +-- Potamilus         
              +--+                     
                 +-- Ligumia           

Hoeh et al., 2002 [Publ 2673], fig. 1-M

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 7 0 0 --
                 +-- Pyganodon_fragilis **
  +--------------+                     
  |              +-- Pyganodon_grandis ** 
--+                                    
  |  +-------------- Fusconaia         
  +--+                                 
     |  +----------- Potamilus         
     +--+                              
        |  +-------- Cyrtonaias        
        +--+                           
           |  +----- Ligumia           
           +--+                        
              |  +-- Lampsilis         
              +--+                     
                 +-- Ortmanniana       

Walker et al., 2006 [Publ 2666], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 13 0 0 --
  +----- Anodonta          
--+                        
  |  +-- Pyganodon_fragilis **
  +--+                     
     +-- Pyganodon_grandis ** 

Walker et al., 2006 [Publ 2666], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 13 0 0 --
  +----- Anodonta          
--+                        
  |  +-- Pyganodon_fragilis **
  +--+                     
     +-- Pyganodon_grandis ** 

Walker et al., 2006 [Publ 2666], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 13 0 0 --
  +----- Anodonta          
--+                        
  |  +-- Pyganodon_fragilis **
  +--+                     
     +-- Pyganodon_grandis ** 

Walker et al., 2006 [Publ 2666], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 34 0 0 --
                    +-- Pyganodon_fragilis **
  +-----------------+                     
  |                 +-- Pyganodon_grandis ** 
  |                                       
  |              +----- Cumberlandia      
--+  +-----------+                        
  |  |           |  +-- Pseudunio         
  |  |           +--+                     
  |  |              +-- Margaritifera     
  +--+                                    
     |              +-- Gonidea           
     |  +-----------+                     
     |  |           +-- Sundadontina      
     +--+                                 
        |  +----------- Cyrtonaias        
        |  |                              
        +--+----------- Fusconaia         
           |                              
           |  +-------- Potamilus         
           +--+                           
              |  +----- Lampsilis         
              +--+                        
                 |  +-- Ligumia           
                 +--+                     
                    +-- Ortmanniana       

Walker et al., 2006 [Publ 2666], fig. 6-M

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 12 0 0 --
                 +-- Pyganodon_fragilis **
  +--------------+                     
  |              +-- Pyganodon_grandis ** 
--+                                    
  |  +-------------- Fusconaia         
  +--+                                 
     |  +----------- Potamilus         
     +--+                              
        |  +-------- Cyrtonaias        
        +--+                           
           |  +----- Ligumia           
           +--+                        
              |  +-- Lampsilis         
              +--+                     
                 +-- Ortmanniana       

Whelan et al., 2023 [Publ 4287], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 27 26 11 --
  +-------------------- Utterbackiana    
  |                                      
  |                 +-- Utterbackia      
--+  +--------------+                    
  |  |              +-- Pyganodon_grandis **
  |  |                                   
  +--+           +----- Alasmidonta      
     |  +--------+                       
     |  |        |  +-- Arcidens         
     |  |        +--+                    
     +--+           +-- Pseudodontoideus 
        |                                
        |  +----------- Alasminota       
        |  |                             
        +--+        +-- Anodontoides     
           |  +-----+                    
           |  |     +-- Strophitus       
           +--+                          
              |  +----- Sulcularia       
              +--+                       
                 |  +-- Platynaias       
                 +--+                    
                    +-- Prolasmidonta    

Boyer et al., 2011 [Publ 2718], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 37 8 8 --
  +-------------- Strophitus       
  |                                
  |        +----- Pterosyna        
--+  +-----+                       
  |  |     |  +-- Lasmigona        
  |  |     +--+                    
  +--+        +-- Arcidens         
     |                             
     |  +-------- Utterbackia      
     |  |                          
     +--+-------- Alasmidonta      
        |                          
        |  +----- Pyganodon_grandis **
        +--+                       
           |  +-- Anodontoides     
           +--+                    
              +-- Platynaias       

Inoue et al., 2014 [Publ 3635], fig. 3A

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 19 13 8 --
     +-------- Pseudodontoideus 
  +--+                          
  |  |  +----- Lasmigona        
  |  +--+                       
  |     |  +-- Pterosyna        
--+     +--+                    
  |        +-- Arcidens         
  |                             
  |  +-------- Strophitus       
  |  |                          
  +--+     +-- Pyganodon_grandis **
     |  +--+                    
     |  |  +-- Utterbackia      
     +--+                       
        |  +-- Platynaias       
        +--+                    
           +-- Alasmidonta      

Krebs et al., 2003 [Publ 2691], fig. 3

cstiTtTbiriSoTroTroSiD
0.889 1 1 0 1 32 8 7 --
  +-- Pyganodon_grandis **
--+                    
  +-- Anodonta         

Chong et al., 2008 [Publ 2668], fig. 3

cstiTtTbiriSoTroTroSiD
0.889 1 1 0 1 17 7 7 --
  +----------------- Anodonta_cygnea            
  |                                             
  +----------------- Pyganodon_fragilis **         
  |                                             
  +----------------- Alasminota                 
  |                                             
  +----------------- Anodonta_anatina           
  |                                             
  +----------------- Unio                       
  |                                             
  |              +-- Brachyanodon_nuttalliana   
  +--------------+                              
  |              +-- Brachyanodon_californiensis
  |                                             
--+              +-- Beringiana                 
  +--------------+                              
  |              +-- Sinanodonta                
  |                                             
  |              +-- Pseudodontoideus           
  |              |                              
  +--------------+-- Alasmidonta                
  |              |                              
  |              +-- Pterosyna                  
  |                                             
  |  +-------------- Pseudanodonta              
  |  |                                          
  |  |        +----- Strophitus                 
  |  +--------+                                 
  +--+        |  +-- Prolasmidonta              
     |        +--+                              
     |           +-- Platynaias                 
     |                                          
     |  +----------- Brachyanodon_oregonensis__4
     +--+                                       
        |  +-------- Brachyanodon_oregonensis__3
        +--+                                    
           |  +----- Brachyanodon_oregonensis__2
           +--+                                 
              |  +-- Brachyanodon_oregonensis__1
              +--+                              
                 +-- Brachyanodon_kennerlyi     

Smith et al., 2018 [Publ 3790], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 17 16 7 --
           +-- Utterbackia      
  +--------+                    
  |        +-- Pyganodon_grandis **
  |                             
--+        +-- Anodontoides     
  |  +-----+                    
  |  |     +-- Strophitus       
  |  |                          
  +--+     +-- Alasmidonta      
     |  +--+                    
     |  |  +-- Alasminota       
     +--+                       
        |  +-- Pseudodontoideus 
        +--+                    
           +-- Utterbackiana    

Boyer et al., 2011 [Publ 2718], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 34 5 5 --
  +----- Alasmidonta      
  |                       
  +----- Pyganodon_grandis **
  |                       
--+  +-- Pterosyna        
  +--+                    
  |  +-- Lasmigona        
  |                       
  |  +-- Platynaias       
  +--+                    
     +-- Strophitus       

Hewitt et al., 2018 [Publ 3758], fig. 5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 8 5 5 --
  +-------- Pyganodon_grandis **
  |                          
  |     +-- Platynaias       
--+-----+                    
  |     +-- Strophitus       
  |                          
  |  +----- Lasmigona        
  +--+                       
     |  +-- Arcidens         
     +--+                    
        +-- Pterosyna        

Huang et al., 2019 [Publ 3843], fig. 2

cstiTtTbiriSoTroTroSiD
0.857 1 1 0 1 200 6 5 --
  +-- Pyganodon_grandis **
--+                    
  +-- Utterbackia      

Huang et al., 2019 [Publ 3843], fig. S3

cstiTtTbiriSoTroTroSiD
0.857 1 1 0 1 198 6 5 --
  +-- Pyganodon_grandis **
--+                    
  +-- Utterbackia      

Huang et al., 2019 [Publ 3843], fig. S4

cstiTtTbiriSoTroTroSiD
0.857 1 1 0 1 200 6 5 --
  +-------- Acuticosta           
  |                              
  +-------- Alasmidonta          
  |                              
  +-------- Anodonta_anatina     
  |                              
  +-------- Anemina              
  |                              
  +-------- Anodonta_cygnea      
  |                              
  +-------- Cristaria            
  |                              
  +-------- Brachyanodon         
  |                              
  +-------- Lasmigona            
  |                              
  +-------- Lepidodesma          
  |                              
  +-------- Pseudanodonta        
  |                              
  +-------- Pyganodon_grandis **    
  |                              
--+-------- Simpsonaias          
  |                              
  +-------- Strophitus           
  |                              
  +-------- Utterbackia          
  |                              
  +-------- Schistodesmus        
  |                              
  +-------- Unio                 
  |                              
  +-------- Scriptolamprotula    
  |                              
  +-------- Lanceolaria          
  |                              
  |     +-- Pletholophus         
  +-----+                        
  |     +-- Sinanodonta          
  |                              
  |  +----- Cuneopsis_celtiformis
  |  |                           
  |  +----- Tchangsinaia         
  |  |                           
  +--+----- Cuneopsis_rufescens  
     |                           
     +----- Nodularia            
     |                           
     |  +-- Cuneopsis_szechenyii 
     +--+                        
        +-- Cuneopsis_heudei     

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 94 6 5 --
        +-- Pyganodon_grandis **
  +-----+                    
  |     +-- Utterbackia      
  |                          
--+     +-- Alasmidonta      
  |  +--+                    
  |  |  +-- Platynaias       
  +--+                       
     |  +-- Prolasmidonta    
     +--+                    
        +-- Pterosyna        

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 94 6 5 --
           +-- Pyganodon_grandis **
  +--------+                    
  |        +-- Utterbackia      
--+                             
  |  +-------- Pterosyna        
  +--+                          
     |  +----- Prolasmidonta    
     +--+                       
        |  +-- Alasmidonta      
        +--+                    
           +-- Platynaias       

Wu et al., 2024 [Publ 4350], fig. 8a_S1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 90 6 5 --
           +-- Pyganodon_grandis **
  +--------+                    
  |        +-- Utterbackia      
--+                             
  |  +-------- Pterosyna        
  +--+                          
     |  +----- Prolasmidonta    
     +--+                       
        |  +-- Alasmidonta      
        +--+                    
           +-- Platynaias       

Wu et al., 2024 [Publ 4350], fig. 8b_S2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 90 6 5 --
           +-- Pyganodon_grandis **
  +--------+                    
  |        +-- Utterbackia      
--+                             
  |  +-------- Pterosyna        
  +--+                          
     |  +----- Prolasmidonta    
     +--+                       
        |  +-- Alasmidonta      
        +--+                    
           +-- Platynaias       

Wu et al., 2024 [Publ 4379], fig. 5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 52 6 5 --
           +-- Pyganodon_grandis **
  +--------+                    
  |        +-- Utterbackia      
--+                             
  |  +-------- Pterosyna        
  +--+                          
     |  +----- Prolasmidonta    
     +--+                       
        |  +-- Alasmidonta      
        +--+                    
           +-- Platynaias       

Hou et al., 2024 [Publ 4400], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 74 6 5 --
           +-- Pyganodon_grandis **
  +--------+                    
  |        +-- Utterbackia      
--+                             
  |  +-------- Pterosyna        
  +--+                          
     |  +----- Prolasmidonta    
     +--+                       
        |  +-- Alasmidonta      
        +--+                    
           +-- Platynaias       

Hou et al., 2025 [Publ 4445], fig. 5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 51 6 5 --
           +-- Pyganodon_grandis **
  +--------+                    
  |        +-- Utterbackia      
--+                             
  |  +-------- Pterosyna        
  +--+                          
     |  +----- Prolasmidonta    
     +--+                       
        |  +-- Platynaias       
        +--+                    
           +-- Alasmidonta      

Wang et al., 2025 [Publ 4488], fig. 3B

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 63 6 5 --
           +-- Pyganodon_grandis **
  +--------+                    
  |        +-- Utterbackia      
--+                             
  |  +-------- Pterosyna        
  +--+                          
     |  +----- Prolasmidonta    
     +--+                       
        |  +-- Alasmidonta      
        +--+                    
           +-- Platynaias       

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 139 7 5 --
        +-- Pyganodon_grandis **
  +-----+                    
  |     +-- Utterbackia      
  |                          
--+     +-- Pterosyna        
  |  +--+                    
  |  |  +-- Prolasmidonta    
  +--+                       
     |  +-- Alasmidonta      
     +--+                    
        +-- Platynaias       

Wu et al., 2026 [Publ 4811], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 94 6 5 --
           +-- Pyganodon_grandis **
  +--------+                    
  |        +-- Utterbackia      
--+                             
  |  +-------- Pterosyna        
  +--+                          
     |  +----- Prolasmidonta    
     +--+                       
        |  +-- Alasmidonta      
        +--+                    
           +-- Platynaias       

Wu et al., 2026 [Publ 4825], fig. 6B

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 64 6 5 --
           +-- Pyganodon_grandis **
  +--------+                    
  |        +-- Utterbackia      
--+                             
  |  +-------- Pterosyna        
  +--+                          
     |  +----- Prolasmidonta    
     +--+                       
        |  +-- Alasmidonta      
        +--+                    
           +-- Platynaias       

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 80 4 4 --
        +-- Pyganodon_grandis **
  +-----+                    
  |     +-- Simpsonaias      
--+                          
  |  +----- Strophitus       
  +--+                       
     |  +-- Alasmidonta      
     +--+                    
        +-- Lasmigona        

Bolotov et al., 2017 [Publ 3761], fig. S1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 138 4 4 --
        +-- Lasmigona        
  +-----+                    
  |     +-- Alasmidonta      
--+                          
  |  +----- Simpsonaias      
  +--+                       
     |  +-- Pyganodon_grandis **
     +--+                    
        +-- Strophitus       

Bolotov et al., 2017 [Publ 3761], fig. S2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 138 4 4 --
  +----- Pyganodon_grandis **
  |                       
  +----- Simpsonaias      
--+                       
  +----- Strophitus       
  |                       
  |  +-- Alasmidonta      
  +--+                    
     +-- Lasmigona        

Bolotov et al., 2017 [Publ 3764], fig. S1

cstiTtTbiriSoTroTroSiD
0.833 1 1 0 1 124 4 4 --
  +-- Alasmidonta             
  |                           
  +-- Anodonta_anatina        
  |                           
  +-- Anodonta_cygnea         
  |                           
  +-- Brachyanodon            
  |                           
  +-- Lasmigona               
  |                           
  +-- Pseudanodonta           
  |                           
  +-- Pyganodon_grandis **       
  |                           
  +-- Simpsonaias             
  |                           
  +-- Strophitus              
--+                           
  +-- Sinanodonta_lauta       
  |                           
  +-- Cristaria_plicata       
  |                           
  +-- Cristaria_radiata       
  |                           
  +-- Pletholophus            
  |                           
  +-- Sinanodonta_lucida      
  |                           
  +-- Sinanodonta_jourdyi     
  |                           
  +-- Lanceolaria_gladiola    
  |                           
  +-- Lanceolaria_grayii      
  |                           
  +-- Lanceolaria_danzhouensis

Bolotov et al., 2018 [Publ 3826], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 81 4 4 --
  +----------- Pyganodon_grandis **
--+                             
  |  +-------- Simpsonaias      
  +--+                          
     |  +----- Strophitus       
     +--+                       
        |  +-- Alasmidonta      
        +--+                    
           +-- Lasmigona        

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 121 4 4 --
  +----- Pyganodon_grandis **
  |                       
  +----- Simpsonaias      
--+                       
  +----- Strophitus       
  |                       
  |  +-- Alasmidonta      
  +--+                    
     +-- Lasmigona        

Sano et al., 2020 [Publ 4138], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 68 5 4 --
        +-- Pyganodon_grandis **
  +-----+                    
  |     +-- Utterbackia      
--+                          
  |  +----- Alasmidonta      
  +--+                       
     |  +-- Prolasmidonta    
     +--+                    
        +-- Platynaias       

Sano et al., 2020 [Publ 4138], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 68 5 4 --
        +-- Pyganodon_grandis **
  +-----+                    
  |     +-- Utterbackia      
--+                          
  |  +----- Alasmidonta      
  +--+                       
     |  +-- Platynaias       
     +--+                    
        +-- Prolasmidonta    

Sano et al., 2020 [Publ 4138], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 68 5 4 --
        +-- Pyganodon_grandis **
  +-----+                    
  |     +-- Utterbackia      
--+                          
  |  +----- Alasmidonta      
  +--+                       
     |  +-- Prolasmidonta    
     +--+                    
        +-- Platynaias       

Chen et al., 2020 [Publ 4170], fig. 1

cstiTtTbiriSoTroTroSiD
0.667 1 1 0 1 33 5 4 --
  +----- Platynaias       
--+                       
  |  +-- Utterbackia      
  +--+                    
     +-- Pyganodon_grandis **

Wu et al., 2021 [Publ 4366], fig. 3-BI

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 72 6 4 --
        +-- Pyganodon_grandis **
  +-----+                    
  |     +-- Utterbackia      
--+                          
  |  +----- Prolasmidonta    
  +--+                       
     |  +-- Alasmidonta      
     +--+                    
        +-- Platynaias       

Wu et al., 2021 [Publ 4366], fig. 3-ML

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 72 6 4 --
        +-- Pyganodon_grandis **
  +-----+                    
  |     +-- Utterbackia      
--+                          
  |  +----- Prolasmidonta    
  +--+                       
     |  +-- Alasmidonta      
     +--+                    
        +-- Platynaias       

Wu et al., 2021 [Publ 4366], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 67 5 4 --
        +-- Pyganodon_grandis **
  +-----+                    
  |     +-- Utterbackia      
--+                          
  |  +----- Prolasmidonta    
  +--+                       
     |  +-- Platynaias       
     +--+                    
        +-- Alasmidonta      

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 75 4 4 --
        +-- Pyganodon_grandis **
  +-----+                    
  |     +-- Simpsonaias      
--+                          
  |  +----- Strophitus       
  +--+                       
     |  +-- Alasmidonta      
     +--+                    
        +-- Lasmigona        

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 75 4 4 --
        +-- Alasmidonta      
  +-----+                    
  |     +-- Lasmigona        
--+                          
  |  +----- Pyganodon_grandis **
  +--+                       
     |  +-- Strophitus       
     +--+                    
        +-- Simpsonaias      

Campbell et al., 2005 [Publ 2660], fig. 1

cstiTtTbiriSoTroTroSiD
0.800 1 1 0 1 101 3 3 --
  +----- Pyganodon_grandis **
--+                       
  |  +-- Lasmigona        
  +--+                    
     +-- Pseudodontoideus 

Campbell et al., 2005 [Publ 2660], fig. 2

cstiTtTbiriSoTroTroSiD
0.800 1 1 0 1 101 3 3 --
  +-------- Pyganodon_grandis **
  |                          
  |     +-- Lasmigona        
--+-----+                    
  |     +-- Pseudodontoideus 
  |                          
  |  +----- Alasminota       
  +--+                       
     |  +-- Anodonta         
     +--+                    
        +-- Brachyanodon     

Graf & Cummings, 2006 [Publ 1968], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 46 3 3 --
  +-------- Lasmigona        
--+                          
  |  +----- Alasmidonta      
  +--+                       
     |  +-- Pyganodon_grandis **
     +--+                    
        +-- Strophitus       

Hoeh et al., 2009 [Publ 2716], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 46 3 3 --
  +-------- Lasmigona        
--+                          
  |  +----- Alasmidonta      
  +--+                       
     |  +-- Strophitus       
     +--+                    
        +-- Pyganodon_grandis **

Hoeh et al., 2009 [Publ 2716], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 45 3 3 --
  +-------- Lasmigona        
--+                          
  |  +----- Alasmidonta      
  +--+                       
     |  +-- Strophitus       
     +--+                    
        +-- Pyganodon_grandis **

Hoeh et al., 2009 [Publ 2716], fig. 5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 45 3 3 --
  +-------- Lasmigona        
--+                          
  |  +----- Alasmidonta      
  +--+                       
     |  +-- Strophitus       
     +--+                    
        +-- Pyganodon_grandis **

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 24 3 3 --
  +----- Lasmigona        
  |                       
--+  +-- Strophitus       
  |  |                    
  +--+-- Alasmidonta      
     |                    
     +-- Pyganodon_grandis **

Amaro et al., 2017 [Publ 3791], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 15 4 3 --
     +-- Lasmigona        
  +--+                    
  |  +-- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia      

Aunins et al., 2018 [Publ 3863], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 39 4 3 --
     +-- Prolasmidonta    
  +--+                    
  |  +-- Alasmidonta      
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia      

Fukata & Iigo, 2020 [Publ 4106], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 53 3 3 --
     +-- Pyganodon_grandis **
  +--+                    
  |  +-- Utterbackia      
--+                       
  |  +-- Alasmidonta      
  +--+                    
     +-- Prolasmidonta    

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 30 2 2 --
  +----- Strophitus       
--+                       
  |  +-- Platynaias       
  +--+                    
     +-- Pyganodon_grandis **

Huang et al., 2013 [Publ 3629], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 15 3 2 --
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia      

Zhou et al., 2016 [Publ 3710], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 19 3 2 --
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia      

Fonseca et al., 2016 [Publ 3775], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 30 3 2 --
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia      

Guerra et al., 2017 [Publ 3772], fig. 2a

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 24 4 2 --
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia      

Guerra et al., 2017 [Publ 3772], fig. 2b

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 24 4 2 --
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia      

Amaro et al., 2017 [Publ 3791], fig. 2-M

cstiTtTbiriSoTroTroSiD
0.750 1 1 0 1 9 2 2 --
  +----- Pyganodon_grandis **
  |                       
--+  +-- Lasmigona        
  |  |                    
  +--+-- Sinanodonta      
     |                    
     +-- Utterbackia      

Amaro et al., 2017 [Publ 3791], fig. 3A

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 11 3 2 --
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia      

Amaro et al., 2017 [Publ 3791], fig. 3B

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 11 3 2 --
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia      

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 31 3 2 --
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia      

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 22 3 2 --
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia      

Huang et al., 2018 [Publ 3828], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 41 3 2 --
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia      

Cha et al., 2018 [Publ 3916], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 37 3 2 --
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia      

Huang et al., 2019 [Publ 3843], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 55 3 2 --
  +----- Lasmigona        
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia      

Wu et al., 2019 [Publ 3864], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 45 3 2 --
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia      

Froufe et al., 2019 [Publ 3936], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 43 3 2 --
  +----- Pterosyna        
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia      

Froufe et al., 2019 [Publ 3936], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 43 3 2 --
  +----- Pterosyna        
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia      

Guerra et al., 2019 [Publ 4168], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 40 4 2 --
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia      

Guerra et al., 2019 [Publ 4168], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 40 4 2 --
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia      

Riccardi et al., 2020 [Publ 4089], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 18 3 2 --
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia      

Wu et al., 2020 [Publ 4121], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 47 3 2 --
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia      

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 47 3 2 --
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia      

Wu et al., 2021 [Publ 4366], fig. 2_I

cstiTtTbiriSoTroTroSiD
0.750 1 1 0 1 90 2 2 --
  +-- Pyganodon_grandis **
--+                    
  +-- Strophitus       

Wu et al., 2021 [Publ 4366], fig. 6

cstiTtTbiriSoTroTroSiD
0.750 1 1 0 1 90 2 2 --
  +-- Pyganodon_grandis **
--+                    
  +-- Strophitus       

Wu et al., 2022 [Publ 4238], fig. 2d

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 37 3 2 --
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia      

Wu et al., 2022 [Publ 4238], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 72 3 2 --
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia      

Dai et al., 2023 [Publ 4285], fig. 5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 52 2 2 --
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia      

Wu et al., 2023 [Publ 4296], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 22 2 2 --
  +----- Lasmigona        
--+                       
  |  +-- Strophitus       
  +--+                    
     +-- Pyganodon_grandis **

Liu et al., 2023 [Publ 4310], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 23 2 2 --
  +----- Lasmigona        
--+                       
  |  +-- Strophitus       
  +--+                    
     +-- Pyganodon_grandis **

Liu et al., 2023 [Publ 4310], fig. 5

cstiTtTbiriSoTroTroSiD
0.750 1 1 0 1 23 2 2 --
  +-- Pyganodon_grandis **
--+                    
  +-- Strophitus       

Dai et al., 2024 [Publ 4358], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 52 2 2 --
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia      

Dai et al., 2025 [Publ 4440], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 86 3 2 --
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia      

Chen et al., 2025 [Publ 4448], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 84 3 2 --
  +----- Platynaias       
--+                       
  |  +-- Pyganodon_grandis **
  +--+                    
     +-- Utterbackia      

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 33 1 1 --
  +-- Strophitus       
--+                    
  +-- Pyganodon_grandis **

Hoeh et al., 2001 [Publ 2652], fig. 14f2

cstiTtTbiriSoTroTroSiD
0.667 1 1 0 1 30 1 1 --
  +----- Pyganodon_grandis **
--+                       
  |  +-- Anodonta         
  +--+                    
     +-- Strophitus       

Hoeh et al., 2001 [Publ 2652], fig. 14f4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 30 1 1 --
  +-- Strophitus       
--+                    
  +-- Pyganodon_grandis **

Graf, 2002 [Publ 2651], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 17 1 1 --
  +-- Alasmidonta      
--+                    
  +-- Pyganodon_grandis **

Hoeh et al., 2002 [Publ 2653], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 30 1 1 --
  +-- Pyganodon_grandis **
--+                    
  +-- Strophitus       

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

cstiTtTbiriSoTroTroSiD
0.667 1 1 0 1 30 1 1 --
  +----- Pyganodon_grandis **
--+                       
  |  +-- Anodonta         
  +--+                    
     +-- Strophitus       

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 30 1 1 --
  +-- Pyganodon_grandis **
--+                    
  +-- Strophitus       

Krebs, 2004 [Publ 2692], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 4 1 1 --
  +-- Lasmigona        
--+                    
  +-- Pyganodon_grandis **

Krebs, 2004 [Publ 2692], fig. 2-M

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 3 1 1 --
  +-- Lasmigona        
--+                    
  +-- Pyganodon_grandis **

Curole & Kocher, 2005 [Publ 2682], fig. 1-M

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 11 1 1 --
  +-- Utterbackiana    
--+                    
  +-- Pyganodon_grandis **

Manendo et al., 2008 [Publ 2669], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 12 1 1 --
  +-- Platynaias       
--+                    
  +-- Pyganodon_grandis **

Whelan et al., 2011 [Publ 3380], fig. 1-ML

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 35 1 1 --
  +-- Pyganodon_grandis **
--+                    
  +-- Lasmigona        

Whelan et al., 2011 [Publ 3380], fig. 1-MP

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 35 1 1 --
  +-- Pyganodon_grandis **
--+                    
  +-- Lasmigona        

Huang et al., 2013 [Publ 3629], fig. 3-M

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 5 1 1 --
  +-- Pyganodon_grandis **
--+                    
  +-- Utterbackia      

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 39 1 1 --
  +-- Pyganodon_grandis **
--+                    
  +-- Lasmigona        

Froufe et al., 2015 [Publ 3664], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 11 1 1 --
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Froufe et al., 2015 [Publ 3664], fig. 1-M

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 7 1 1 --
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Wu et al., 2016 [Publ 3711], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 20 2 1 --
  +-- Pyganodon_grandis **
--+                    
  +-- Utterbackia      

Fonseca et al., 2016 [Publ 3775], fig. 1-M

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 8 1 1 --
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Feng et al., 2016 [Publ 3800], fig. 2

cstiTtTbiriSoTroTroSiD
0.667 1 1 0 1 13 1 1 --
  +----- Pyganodon_grandis **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta      

Guerra et al., 2017 [Publ 3772], fig. 2a-M

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 7 1 1 --
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Guerra et al., 2017 [Publ 3772], fig. 2b-M

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 7 1 1 --
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Burzynski et al., 2017 [Publ 3774], fig. 1-bottom

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 12 1 1 --
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Burzynski et al., 2017 [Publ 3774], fig. 1-top

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 12 1 1 --
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Amaro et al., 2017 [Publ 3791], fig. 3A-M

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 6 1 1 --
  +-- Pyganodon_grandis **
--+                    
  +-- Utterbackia      

Amaro et al., 2017 [Publ 3791], fig. 3B-M

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 6 1 1 --
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 9 1 1 --
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Wen et al., 2017 [Publ 3795], fig. 3-M

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 5 1 1 --
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Soroka & Burzynski, 2018 [Publ 3762], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 6 1 1 --
  +-- Pyganodon_grandis **
--+                    
  +-- Utterbackia      

Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 6 1 1 --
  +-- Pyganodon_grandis **
--+                    
  +-- Utterbackia      

Araujo et al., 2018 [Publ 3779], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 17 1 1 --
  +-- Pyganodon_grandis **
--+                    
  +-- Alasmidonta      

Huang et al., 2018 [Publ 3828], fig. 6-M

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 16 1 1 --
  +-- Pyganodon_grandis **
--+                    
  +-- Utterbackia      

Wang et al., 2018 [Publ 3846], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 30 1 1 --
  +-- Pyganodon_grandis **
--+                    
  +-- Platynaias       

Chase et al., 2018 [Publ 3861], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 10 1 1 --
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Chase et al., 2018 [Publ 3861], fig. 1-M

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 10 1 1 --
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Burzynski & Soroka, 2018 [Publ 3862], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 3 1 1 --
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 19 1 1 --
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5b

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 9 1 1 --
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Wang et al., 2018 [Publ 3865], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 30 1 1 --
  +-- Pyganodon_grandis **
--+                    
  +-- Platynaias       

Wu et al., 2018 [Publ 4143], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 21 2 1 --
  +-- Pyganodon_grandis **
--+                    
  +-- Utterbackia      

Huang et al., 2019 [Publ 3843], fig. 3-M

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 22 1 1 --
  +-- Pyganodon_grandis **
--+                    
  +-- Utterbackia      

Froufe et al., 2019 [Publ 3936], fig. 3-M

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 18 1 1 --
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Froufe et al., 2019 [Publ 3936], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 18 1 1 --
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Guerra et al., 2019 [Publ 4168], fig. 1-M

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 21 1 1 --
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Guerra et al., 2019 [Publ 4168], fig. 2-M

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 21 1 1 --
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Wu et al., 2020 [Publ 4147], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 29 2 1 --
  +-- Pyganodon_grandis **
--+                    
  +-- Utterbackia      

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 22 1 1 --
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Smith, 2021 [Publ 4331], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 19 1 1 --
  +-- Pyganodon_grandis **
--+                    
  +-- Utterbackia      

Wu et al., 2022 [Publ 4238], fig. 2e

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 17 1 1 --
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Dai et al., 2024 [Publ 4386], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 40 1 1 --
  +-- Pyganodon_grandis **
--+                    
  +-- Utterbackia      

Klymus et al., 2025 [Publ 4814], fig. 3A

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 35 1 1 --
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Klymus et al., 2025 [Publ 4814], fig. 3B

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 35 1 1 --
  +-- Utterbackia      
--+                    
  +-- Pyganodon_grandis **

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 46 1 1 --
  +-- Pyganodon_grandis **
--+                    
  +-- Utterbackia      

Graf, 2000 [Publ 2647], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 16 0 0 --
  +-- Pyganodon **
--+            
  +-- Lampsilis

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 10 0 0 --
  +-- Unio             
--+                    
  +-- Pyganodon_grandis **

Curole & Kocher, 2005 [Publ 2682], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 10 0 0 --
  +-- Pyganodon_grandis **
--+                    
  +-- Margaritifera    

Zanatta & Murphy, 2006 [Publ 2672], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 55 0 0 --
  +----------------------------------------- Pyganodon_grandis **      
  |                                                                 
  |                                      +-- Amblema                
  |                                   +--+                          
  |                                   |  +-- Popenaias              
  |                                   |                             
--+  +--------------------------------+  +-- Pleurobema             
  |  |                                |  |                          
  |  |                                |  +-- Eurynia                
  |  |                                +--+                          
  |  |                                   +-- Pleuronaia             
  |  |                                   |                          
  +--+                                   +-- Fusconaia              
     |                                                              
     |                                   +-- Cyrtonaias             
     |  +--------------------------------+                          
     |  |                                +-- Glebula                
     |  |                                                           
     +--+  +-------------------------------- Paetulunio             
        |  |                                                        
        |  |                             +-- Obliquaria             
        +--+  +--------------------------+                          
           |  |                          +-- Toxolasma              
           |  |                                                     
           |  |  +-------------------------- Sagittunio             
           +--+  |                                                  
              |  |              +----------- Potamilus              
              |  |              |                                   
              |  |  +-----------+        +-- Medionidus_conradicus  
              |  |  |           |  +-----+                          
              +--+  |           |  |     +-- Ptychobranchus         
                 |  |           +--+                                
                 |  |              |  +----- Medionidus_acutissimus 
                 |  |              |  |                             
                 |  |              +--+----- Lemiox                 
                 |  |                 |                             
                 |  |                 |  +-- Dromus                 
                 +--+                 +--+                          
                    |                    +-- Cyprogenia             
                    |                                               
                    |                 +----- Lampsilis_teres        
                    |  +--------------+                             
                    |  |              |  +-- Truncilla              
                    |  |              +--+                          
                    |  |                 +-- Ellipsaria             
                    |  |                                            
                    +--+  +----------------- Leaunio                
                       |  |                                         
                       |  |     +----------- Hamiota                
                       |  |     |                                   
                       |  |  +--+        +-- Lampsilis_fasciola     
                       +--+  |  |  +-----+                          
                          |  |  |  |     +-- Ortmanniana_pectorosa  
                          |  |  +--+                                
                          |  |     |  +----- Lampsilis_ornata       
                          |  |     +--+                             
                          |  |        |  +-- Lampsilis_ovata        
                          +--+        +--+                          
                             |           +-- Lampsilis_cardium      
                             |                                      
                             |        +----- Venustaconcha          
                             |  +-----+                             
                             |  |     |  +-- Obovaria               
                             |  |     +--+                          
                             +--+        +-- Epioblasma             
                                |                                   
                                |  +-------- Lampsilis_siliquoidea  
                                +--+                                
                                   |  +----- Ortmanniana_ligamentina
                                   +--+                             
                                      |  +-- Cambarunio             
                                      +--+                          
                                         +-- Ligumia                

Zanatta & Murphy, 2006 [Publ 2672], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 56 0 0 --
  +----------------------------------------------- Pyganodon_grandis **      
  |                                                                       
  +----------------------------------------------- Cumberlandia           
  |                                                                       
  |                                      +-------- Pleuronaia             
  |                                      |                                
--+  +-----------------------------------+-------- Pleurobema             
  |  |                                   |                                
  |  |                                   |  +----- Fusconaia              
  |  |                                   +--+                             
  |  |                                      |  +-- Hemistena              
  |  |                                      +--+                          
  +--+                                         +-- Eurynia                
     |                                                                    
     |  +----------------------------------------- Amblema                
     |  |                                                                 
     |  +----------------------------------------- Popenaias              
     |  |                                                                 
     |  |                                      +-- Paetulunio             
     +--+  +-----------------------------------+                          
        |  |                                   +-- Toxolasma              
        |  |                                                              
        |  |                                +----- Obliquaria             
        |  |  +-----------------------------+                             
        +--+  |                             |  +-- Glebula                
           |  |                             +--+                          
           |  |                                +-- Cyrtonaias             
           |  |                                                           
           |  |                             +----- Ptychobranchus         
           |  |                          +--+                             
           +--+                          |  |  +-- Dromus                 
              |                          |  +--+                          
              |  +-----------------------+     +-- Cyprogenia             
              |  |                       |                                
              |  |                       |  +----- Medionidus_conradicus  
              |  |                       +--+                             
              |  |                          |  +-- Medionidus_acutissimus 
              |  |                          +--+                          
              +--+                             +-- Lemiox                 
                 |                                                        
                 |  +----------------------------- Sagittunio             
                 |  |                                                     
                 |  +----------------------------- Potamilus              
                 |  |                                                     
                 |  |                          +-- Truncilla              
                 +--+  +-----------------------+                          
                    |  |                       +-- Ellipsaria             
                    |  |                                                  
                    |  |                       +-- Ortmanniana_pectorosa  
                    |  |                 +-----+                          
                    |  |                 |     +-- Lampsilis_fasciola     
                    +--+  +--------------+                                
                       |  |              |  +----- Lampsilis_ornata       
                       |  |              +--+                             
                       |  |                 |  +-- Lampsilis_ovata        
                       |  |                 +--+                          
                       |  |                    +-- Lampsilis_cardium      
                       +--+                                               
                          |                    +-- Ligumia                
                          |                 +--+                          
                          |                 |  +-- Cambarunio             
                          |  +--------------+                             
                          |  |              |  +-- Lampsilis_teres        
                          |  |              +--+                          
                          +--+                 +-- Hamiota                
                             |                                            
                             |  +----------------- Leaunio                
                             |  |                                         
                             +--+  +-------------- Ortmanniana_ligamentina
                                |  |                                      
                                +--+  +----------- Lampsilis_siliquoidea  
                                   |  |                                   
                                   +--+  +-------- Obovaria_olivaria      
                                      |  |                                
                                      |  +-------- Venustaconcha          
                                      +--+                                
                                         +-------- Epioblasma             
                                         |                                
                                         |  +----- Obovaria_unicolor      
                                         +--+                             
                                            |  +-- Obovaria_subrotunda    
                                            +--+                          
                                               +-- Obovaria_arkansasensis 

Soroka, 2008 [Publ 3695], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 3 0 0 --
  +-- Anodonta         
--+                    
  +-- Pyganodon_grandis **

Soroka, 2008 [Publ 3695], fig. 1-M

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 3 0 0 --
  +-- Anodonta         
--+                    
  +-- Pyganodon_grandis **

Plazzi et al., 2011 [Publ 3676], fig. 5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 17 0 0 --
  +-- Pyganodon_grandis **
--+                    
  +-- Pseudanodonta    

Chen et al., 2012 [Publ 3624], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 6 0 0 --
  +-- Pyganodon_grandis **
--+                    
  +-- Cristaria        

Inoue et al., 2013 [Publ 3528], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 16 0 0 --
  +----------------- Gonidea          
  |                                   
  +----------------- Pyganodon_grandis **
  |                                   
--+              +-- Eurynia          
  |  +-----------+                    
  |  |           +-- Quadrula         
  |  |                                
  +--+        +----- Paetulunio       
     |  +-----+                       
     |  |     |  +-- Amblema          
     |  |     +--+                    
     +--+        +-- Reginaia         
        |                             
        |  +-------- Lampsilis        
        |  |                          
        +--+  +----- Leaunio          
           |  |                       
           +--+----- Obovaria         
              |                       
              |  +-- Villosa          
              +--+                    
                 +-- Cambarunio       

Zanatta & Harris, 2013 [Publ 3530], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 4 0 0 --
  +-------- Pyganodon_grandis **
--+                          
  |  +----- Lampsilis        
  +--+                       
     |  +-- Epioblasma       
     +--+                    
        +-- Venustaconcha    

Inoue et al., 2013 [Publ 3579], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 8 0 0 --
  +-------- Pyganodon_grandis **
--+                          
  |  +----- Cumberlandia     
  +--+                       
     |  +-- Pseudunio        
     +--+                    
        +-- Margaritifera    

Marshall et al., 2014 [Publ 3593], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 13 0 0 --
  +-- Pyganodon_grandis **
--+                    
  +-- Unio             

Sheng et al., 2014 [Publ 3704], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 7 0 0 --
  +----------- Pyganodon_grandis **
  |                             
--+  +-------- Lampsilis        
  |  |                          
  +--+     +-- Venustaconcha    
     |  +--+                    
     |  |  +-- Quadrula         
     +--+                       
        |  +-- Lamprotula       
        +--+                    
           +-- Sinohyriopsis    

Wei et al., 2016 [Publ 3665], fig. 0

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 5 0 0 --
  +-- Cristaria        
  |                    
  +-- Lampsilis        
--+                    
  +-- Pyganodon_grandis **
  |                    
  +-- Quadrula         

Lopes-Lima et al., 2019 [Publ 3901], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 27 0 0 --
  +-- Brachyanodon     
--+                    
  +-- Pyganodon_grandis **

Lopes-Lima et al., 2019 [Publ 3901], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 28 0 0 --
  +-- Brachyanodon     
--+                    
  +-- Pyganodon_grandis **

Lopes-Lima et al., 2019 [Publ 3901], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 25 0 0 --
  +-- Brachyanodon     
--+                    
  +-- Pyganodon_grandis **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 56 0 0 --
  +-- Anodonta         
--+                    
  +-- Pyganodon_grandis **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 56 0 0 --
  +-- Anodonta         
--+                    
  +-- Pyganodon_grandis **

Hewitt et al., 2019 [Publ 3962], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 9 0 0 --
  +-------------------------- Pyganodon_grandis **    
  |                                                
--+  +----------------------- Fusconaia            
  |  |                                             
  +--+  +-------------------- Lampsilis_siliquoidea
     |  |                                          
     |  |  +----------------- Lampsilis_fasciola   
     +--+  |                                       
        |  |              +-- Epioblasma           
        |  +--------------+                        
        +--+              +-- Lampsilis_radiata    
           |                                       
           |  +-------------- Lampsilis_ornata     
           |  |                                    
           +--+           +-- Lampsilis_cardium__1 
              |  +--------+                        
              |  |        +-- Lampsilis_ovata__1   
              +--+                                 
                 |  +-------- Lampsilis_cardium__6 
                 |  |                              
                 +--+  +----- Lampsilis_cardium__5 
                    |  |                           
                    +--+  +-- Lampsilis_cardium__2 
                       |  |                        
                       |  +-- Lampsilis_ovata__2   
                       +--+                        
                          +-- Lampsilis_cardium__3 
                          |                        
                          +-- Lampsilis_cardium__4 

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 36 0 0 --
  +-- Anodonta         
--+                    
  +-- Pyganodon_grandis **

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 38 0 0 --
  +-- Anodonta         
--+                    
  +-- Pyganodon_grandis **