Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Anodontini | Alasmidontina

genus Strophitus Rafinesque, 1820: cladomics

5 Recent valid species (8 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 5 (8 total)
analyzed in 54 cladograms in 37 publications
maximum number of terminals in an analysis = 5 (mean = 1.48, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 8
maximum number of included terminals = 5, (mean = 4.25 median = 4.5)
proportion of cladograms that support basic monophyly = 1

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Strophitus cladogram distances
PCoA Ordination

NJ Tree of Strophitus cladogram distances
NJ Tree

Cladograms

Perkins et al., 2025 [Publ 4447], fig. 2_mtDNA-ITS

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cst iT tT bi riS oT roT roS iD
1.000 5 5 5 5 31 28 14 **
Cladomics cladogram Utterbackiana Alasminota Pseudodontoideus Lasmigona Pterosyna Arcidens Prolasmidonta Platynaias Ligodonta Alasmidonta Sulcularia Pyganodon Utterbackia Anodontoides Strophitus undulatus 2 Strophitus howellsi Strophitus undulatus 1 Strophitus pascagoulaensis 2 Strophitus pascagoulaensis 1 Strophitus williamsi Strophitus radiatus

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cst iT tT bi riS oT roT roS iD
0.933 5 5 5 5 628 32 13 4.0
Cladomics cladogram Utterbackia Pyganodon Utterbackiana Simpsonaias Alasminota Alasmidonta Prolasmidonta Platynaias Anodontoides Strophitus undulatus 2 Strophitus howellsi Strophitus undulatus 1 Strophitus pascagoulaensis 2 Strophitus radiatus Strophitus pascagoulaensis 1 Strophitus williamsi

Smith et al., 2018 [Publ 3790], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 5 5 4 5 13 12 7 3.0
Cladomics cladogram Utterbackia Pyganodon Alasmidonta Alasminota Pseudodontoideus Utterbackiana Anodontoides Strophitus pascagoulaensis Strophitus radiatus Strophitus williamsi Strophitus howellsi 2 Strophitus undulatus Strophitus howellsi 1

Smith et al., 2023 [Publ 4291], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 5 2 2 1 2.0
Cladomics cladogram Anodontoides Strophitus undulatus Strophitus howellsi Strophitus pascagoulaensis Strophitus radiatus Strophitus williamsi

Whelan et al., 2023 [Publ 4287], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 4 4 3 4 37 29 13 5.0
Cladomics cladogram Utterbackia Pyganodon Utterbackiana Lasmigona Pseudodontoideus Pterosyna Arcidens Prolasmidonta Platynaias Alasminota Alasmidonta Sulcularia Anodontoides Strophitus undulatus Strophitus pascagoulaensis 2 Strophitus radiatus Strophitus pascagoulaensis 1 Strophitus williamsi

Whelan et al., 2023 [Publ 4287], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 4 4 3 4 24 23 11 3.0
Cladomics cladogram Utterbackiana Utterbackia Pyganodon Alasmidonta Arcidens Pseudodontoideus Alasminota Sulcularia Platynaias Prolasmidonta Anodontoides Strophitus undulatus Strophitus pascagoulaensis 2 Strophitus pascagoulaensis 1 Strophitus radiatus Strophitus williamsi

Whelan et al., 2023 [Publ 4287], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 23 21 10 5.0
Cladomics cladogram Sulcularia Utterbackiana Alasminota Arcidens Pseudodontoideus Alasmidonta Platynaias Pyganodon Utterbackia Anodontoides Strophitus pascagoulaensis Strophitus undulatus Strophitus radiatus Strophitus williamsi

Krebs et al., 2003 [Publ 2691], fig. 3

cst iT tT bi riS oT roT roS iD
0.889 2 2 0 2 31 7 7 --
Cladomics cladogram Strophitus radiatus Strophitus undulatus Utterbackia Pyganodon Anodonta

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

cst iT tT bi riS oT roT roS iD
0.455 1 1 0 1 104 15 9 --
Cladomics cladogram Alasmidonta marginata Prolasmidonta Alasmidonta undulata Beringiana Brachyanodon kennerlyi Brachyanodon nuttalliana Brachyanodon oregonensis Arcidens confragosus Arcidens wheeleri Pterosyna Platynaias compressa Lasmigona Platynaias decorata Platynaias subviridis Pyganodon grandis Pyganodon lacustris Pseudodontoideus Strophitus undulatus Utterbackia imbecillis Utterbackia peninsularis 1 Utterbackia peninsularis 2

Boyer et al., 2011 [Publ 2718], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 37 8 8 --
Cladomics cladogram Strophitus undulatus Pterosyna Lasmigona Arcidens Utterbackia Alasmidonta Pyganodon Anodontoides Platynaias

Inoue et al., 2014 [Publ 3635], fig. 3A

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 19 13 8 --
Cladomics cladogram Pseudodontoideus Lasmigona Pterosyna Arcidens Strophitus undulatus Pyganodon Utterbackia Platynaias Alasmidonta

Chong et al., 2008 [Publ 2668], fig. 3

cst iT tT bi riS oT roT roS iD
0.889 1 1 0 1 17 7 7 --
Cladomics cladogram Strophitus undulatus Prolasmidonta Platynaias

Breton et al., 2011 [Publ 3376], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 37 10 6 --
Cladomics cladogram Pterosyna Lasmigona Strophitus undulatus Platynaias Alasmidonta Pyganodon Utterbackia

Chase et al., 2018 [Publ 3847], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 38 10 6 --
Cladomics cladogram Pyganodon Utterbackia Lasmigona Pterosyna Alasmidonta Strophitus undulatus Platynaias

King et al., 1999 [Publ 2699], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 8 8 5 --
Cladomics cladogram Pyganodon Lasmigona Pterosyna Strophitus undulatus Prolasmidonta Platynaias

Boyer et al., 2011 [Publ 2718], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 34 5 5 --
Cladomics cladogram Alasmidonta Pyganodon Pterosyna Lasmigona Platynaias Strophitus undulatus

Hewitt et al., 2018 [Publ 3758], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 8 5 5 --
Cladomics cladogram Pyganodon Platynaias Strophitus undulatus Lasmigona Arcidens Pterosyna

Huang et al., 2019 [Publ 3843], fig. 2

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 200 6 5 --
Cladomics cladogram Alasmidonta Lasmigona Pseudanodonta Simpsonaias Strophitus undulatus Pyganodon Utterbackia Anodonta anatina Anodonta cygnea Brachyanodon

Huang et al., 2019 [Publ 3843], fig. S3

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 198 6 5 --
Cladomics cladogram Simpsonaias Strophitus undulatus

Huang et al., 2019 [Publ 3843], fig. S4

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 200 6 5 --
Cladomics cladogram Acuticosta Alasmidonta Anodonta anatina Anemina Anodonta cygnea Cristaria Brachyanodon Lasmigona Lepidodesma Pseudanodonta Pyganodon Simpsonaias Strophitus undulatus Utterbackia Schistodesmus Unio Scriptolamprotula Lanceolaria Pletholophus Sinanodonta Cuneopsis celtiformis Tchangsinaia Cuneopsis rufescens Nodularia Cuneopsis szechenyii Cuneopsis heudei

Stewart et al., 2021 [Publ 4451], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 15 9 5 --
Cladomics cladogram Alasmidonta Pterosyna Strophitus undulatus Platynaias Pyganodon Utterbackia

Lydeard et al., 1996 [Publ 2656], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 26 5 4 --
Cladomics cladogram Utterbackia Utterbackiana Strophitus radiatus Lasmigona Alasmidonta

Lydeard et al., 1996 [Publ 2656], fig. 4

cst iT tT bi riS oT roT roS iD
0.833 1 1 0 1 27 4 4 --
Cladomics cladogram Strophitus radiatus Lasmigona Alasmidonta

King et al., 1999 [Publ 2699], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 5 4 --
Cladomics cladogram Pterosyna Lasmigona Prolasmidonta Strophitus undulatus Platynaias

Huang et al., 2002 [Publ 2657], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 33 4 4 --
Cladomics cladogram Utterbackiana Utterbackia Strophitus radiatus Alasmidonta Lasmigona

Huang et al., 2002 [Publ 2657], fig. 2

cst iT tT bi riS oT roT roS iD
0.833 1 1 0 1 33 4 4 --
Cladomics cladogram Strophitus radiatus Schistodesmus Lepidodesma Acuticosta Lanceolaria Sinanodonta Cristaria Nodularia Tchangsinaia Utterbackia Utterbackiana Alasmidonta Lasmigona

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 80 4 4 --
Cladomics cladogram Pyganodon Simpsonaias Strophitus undulatus Alasmidonta Lasmigona

Bolotov et al., 2017 [Publ 3761], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 138 4 4 --
Cladomics cladogram Lasmigona Alasmidonta Simpsonaias Pyganodon Strophitus undulatus

Bolotov et al., 2017 [Publ 3761], fig. S2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 138 4 4 --
Cladomics cladogram Pyganodon Simpsonaias Strophitus undulatus Alasmidonta Lasmigona

Bolotov et al., 2017 [Publ 3764], fig. S1

cst iT tT bi riS oT roT roS iD
0.833 1 1 0 1 124 4 4 --
Cladomics cladogram Alasmidonta Anodonta anatina Anodonta cygnea Brachyanodon Lasmigona Pseudanodonta Pyganodon Simpsonaias Strophitus undulatus Sinanodonta lauta Cristaria plicata Cristaria radiata Pletholophus Sinanodonta lucida Sinanodonta jourdyi Lanceolaria gladiola Lanceolaria grayii Lanceolaria danzhouensis

Bolotov et al., 2018 [Publ 3826], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 81 4 4 --
Cladomics cladogram Pyganodon Simpsonaias Strophitus undulatus Alasmidonta Lasmigona

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 121 4 4 --
Cladomics cladogram Pyganodon Simpsonaias Strophitus undulatus Alasmidonta Lasmigona

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 75 4 4 --
Cladomics cladogram Pyganodon Simpsonaias Strophitus undulatus Alasmidonta Lasmigona

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 75 4 4 --
Cladomics cladogram Alasmidonta Lasmigona Pyganodon Strophitus undulatus Simpsonaias

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 29 4 3 --
Cladomics cladogram Alasmidonta Lasmigona Strophitus undulatus Pyganodon

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 29 4 3 --
Cladomics cladogram Alasmidonta Lasmigona Strophitus undulatus Pyganodon

Graf & Cummings, 2006 [Publ 1968], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 46 3 3 --
Cladomics cladogram Lasmigona Alasmidonta Pyganodon Strophitus undulatus

Hoeh et al., 2009 [Publ 2716], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 46 3 3 --
Cladomics cladogram Lasmigona Alasmidonta Strophitus undulatus Pyganodon

Hoeh et al., 2009 [Publ 2716], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 45 3 3 --
Cladomics cladogram Lasmigona Alasmidonta Strophitus undulatus Pyganodon

Hoeh et al., 2009 [Publ 2716], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 45 3 3 --
Cladomics cladogram Lasmigona Alasmidonta Strophitus undulatus Pyganodon

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 24 3 3 --
Cladomics cladogram Lasmigona Strophitus undulatus Alasmidonta Pyganodon

Wu et al., 2018 [Publ 3836], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 4 3 --
Cladomics cladogram Strophitus undulatus Platynaias Alasmidonta Pyganodon

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 2 2 --
Cladomics cladogram Strophitus undulatus Platynaias Pyganodon

Wu et al., 2021 [Publ 4366], fig. 2_I

cst iT tT bi riS oT roT roS iD
0.750 1 1 0 1 90 2 2 --
Cladomics cladogram Pyganodon Strophitus undulatus

Wu et al., 2021 [Publ 4366], fig. 6

cst iT tT bi riS oT roT roS iD
0.750 1 1 0 1 90 2 2 --
Cladomics cladogram Pyganodon Strophitus undulatus

Wu et al., 2023 [Publ 4296], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 22 2 2 --
Cladomics cladogram Lasmigona Strophitus undulatus Pyganodon

Liu et al., 2023 [Publ 4310], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 23 2 2 --
Cladomics cladogram Lasmigona Strophitus undulatus Pyganodon

Liu et al., 2023 [Publ 4310], fig. 5

cst iT tT bi riS oT roT roS iD
0.750 1 1 0 1 23 2 2 --
Cladomics cladogram Pyganodon Strophitus undulatus

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 33 1 1 --
Cladomics cladogram Strophitus undulatus Pyganodon

Hoeh et al., 2001 [Publ 2652], fig. 14f2

cst iT tT bi riS oT roT roS iD
0.667 1 1 0 1 30 1 1 --
Cladomics cladogram Anodonta Strophitus undulatus

Hoeh et al., 2001 [Publ 2652], fig. 14f4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 1 1 --
Cladomics cladogram Strophitus undulatus Pyganodon

Hoeh et al., 2002 [Publ 2653], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 1 1 --
Cladomics cladogram Pyganodon Strophitus undulatus

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

cst iT tT bi riS oT roT roS iD
0.667 1 1 0 1 30 1 1 --
Cladomics cladogram Anodonta Strophitus undulatus

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 1 1 --
Cladomics cladogram Pyganodon Strophitus undulatus