Bivalvia | Unionida | Hyrioidea | Hyriidae | Hyriinae | Rhipidodontini | Diplodon

species Diplodon chilensis (Gray, 1828): specimens

28 available nominal species synonyms (42 total synonyms)

Recent Taxonomic Opinions

Diplodon chilensisGraf & Cummings (2007), Fabbri et al. (2008), Hoeh et al. (2009), Bogan (2010), IUCN (2011), ... Graf & Cummings (2021), Torres et al. (2024), Nagel (2025), Campbell & Lydeard (2026), Prié et al. (2026)
Diplodon patagonicusGraf & Cummings (2007), Bogan (2010)
Diplodon chilensis chilensisParada & Peredo (2008)
Diplodon chilensis patagonicusParada & Peredo (2008)
Diplodon (s.s.) chilensisRumi et al. (2008), Pereira et al. (2014), Torres et al. (2018)
Rhipidodonta charruanaNagel (2025)

Range

Southern Chile and Argentina, on both sides of the Andes. (South America: SA6)

Map of Georeferenced Specimens

specimens map of Diplodon chilensis

MUSSELp Specimen Records

Type Specimens

image of lot SMF_3856
SMF_3856
Patagonien am Fuss der Kordilleren
lectotype
Unio andina Ihering, 1893

image of lot SMF_4322
SMF_4322
Laguna Villavira, S. Chile
possible syntype
Unio foncki Philippi, 1869

image of lot BMNH_MP_365
BMNH_MP_365
Id. of Chiloe, Chile
holotype
Unio fragilis Reeve, 1865

image of lot MNHN_IM-2000-1703
MNHN_IM-2000-1703
Rio Carampagua, S.O. de Concepcion, Chili
syntypes
Unio gassiei Küster, 1856

image of lot SMF_3850
SMF_3850
Prov. Santiago, Chile
idiotype
Unio jacobaeus Philippi, 1869

image of lot USNM_186117A
USNM_186117A
Nahuel Huape, Argentina
paratype
Diplodon huapensis Bartsch, 1906

image of lot SMF_3857
SMF_3857
Patagonien am Fuss der Kordillere
paralectotype
Unio andina Ihering, 1893

image of lot BMNH_1965145
BMNH_1965145
I. of Chiloe
paralectotypes
Unio auratus Sowerby, 1866

image of lot SMF_3852
SMF_3852
Rio Guayellua, S. Chile
possible syntype
Unio foncki Philippi, 1869

image of lot BMNH_1854-12-4-851
BMNH_1854-12-4-851
Rio Negro, Patagonia
paralectotypes
Unio patagonica d’Orbigny, 1835

image of lot SMF_3853
SMF_3853
Rio Negro, Patagonien
paralectotype
Unio patagonica d’Orbigny, 1835

Other Specimens by Collection | show by country

11 Specimens from AMNH

image of lot AMNH_92260
AMNH_92260
Lago Correntoso, Parque Nacional Nahuel Huapi, Pcia. Rio Negro, Argentina

image of lot AMNH_35039
AMNH_35039
Lake Fellafquen, Chebut, Argentina

image of lot AMNH_30901
AMNH_30901
Rio Negros, Patagonia

image of lot AMNH_132931
AMNH_132931
Dalcahue, Isla de Chiloe, Chile

image of lot AMNH_91549
AMNH_91549
Rio Maipu, near Isla Maipu, South Santiago, Chile

image of lot AMNH_146552
AMNH_146552
Rio Maullin, Llanguihue, Los Lagos, Chile

image of lot AMNH_30925
AMNH_30925
South America

image of lot AMNH_30926
AMNH_30926
South America

image of lot AMNH_30916
AMNH_30916
South America

1 Specimens from AMS

image of lot AMS_126594
AMS_126594
Valdivia, Chile

7 Specimens from ANSP

image of lot ANSP_41887
ANSP_41887
Quillota, Valparaiso, Chili

image of lot ANSP_125831
ANSP_125831
Rio Camamnagua, Chili

image of lot ANSP_41795
ANSP_41795
So. America

9 Specimens from BMNH

image of lot BMNH_MP_034
BMNH_MP_034
Largo, Argentina

image of lot BMNH_1964138
BMNH_1964138
Patagonia

image of lot BMNH_MP_024
BMNH_MP_024
[Lima] [Chile]

image of lot BMNH_54-12-4-848
BMNH_54-12-4-848
La Laguna, Chili

image of lot BMNH_MP_379
BMNH_MP_379
[no locality available - South America]

11 Specimens from CM

image of lot CM_61.9743
CM_61.9743
Lake Nahuel Huapi, Argentina

image of lot CM_61-136
CM_61-136
Lake Nahuel Huapi, Neuquen, Argentina

image of lot CM_61.5851
CM_61.5851
Rio Limay, Patagonia, Argentina

image of lot CM_61.9246
CM_61.9246
Rio Limay, Patagonia, Argentina

image of lot CM_61.9247
CM_61.9247
Rio Limay, Patagonia, Argentina

image of lot CM_61.9245
CM_61.9245
Colchol River, Nuevo Imperial, Chile

image of lot CM_61.2027
CM_61.2027
Island of Chiloe, Chile

image of lot CM_MP_23
CM_MP_23
Rio Choapa, between La Serena and Calera, Prov. Coquimbo, Chile

image of lot CM_61.9744
CM_61.9744
Valdivia, Chile

11 Specimens from DMNH

image of lot DMNH_173115
DMNH_173115
Arroyo Pilcan, Rio Limay drainage, Rio Negro, Argentina

image of lot DMNH_174544
DMNH_174544
Arroyo Pilcan, Rio Limey drainage, Rio Negro, Argentina

image of lot DMNH_173113
DMNH_173113
Arroyo Pilear, near Rio Negro, Argentina

image of lot DMNH_173071
DMNH_173071
Lago Fatalafquen, Chubut, Argentina

image of lot DMNH_173114
DMNH_173114
Lago Fetalanfquia, Chubut, Argentina

image of lot DMNH_174561
DMNH_174561
Lago Villarino, Neuquen, Argentina

image of lot DMNH_173116
DMNH_173116
Lago Villarino, Neuquen, Argentina

image of lot DMNH_33151
DMNH_33151
Caldevia, Chili

image of lot DMNH_56331
DMNH_56331
Valencia, Chile

38 Specimens from FMNH

image of lot FMNH_21836
FMNH_21836
Argentina

image of lot FMNH_179984
FMNH_179984
Dalcahue, Ancud, Chiloe, Chile; 42°23’S, 73°40’W

image of lot FMNH_115739
FMNH_115739
Dalcahue, Id. Chiloe, Chile

image of lot FMNH_198226
FMNH_198226
Lago Azul, Llanquihue, Chiloe, Chile; 41°57’S, 71°51’W

image of lot FMNH_179988
FMNH_179988
Lago Blanco, Llanquihue, Chiloe, Chile; 41°57’S, 71°53’W

image of lot FMNH_198229
FMNH_198229
Lago Chico, NE of Lago Paloma, Territorio de Aysen, Chile; 45°54’S, 72°09’W

image of lot FMNH_198222
FMNH_198222
Lago de las Rocas, Llanouihue, Chiloe, Chile; 42°02’S, 71°50’W

image of lot FMNH_198210
FMNH_198210
Lago Esmeralda, Territorio de Aisen, Chile; 47°19’S, 72°34’W

image of lot FMNH_198216
FMNH_198216
Lago la Paloma, Territorio de Aysen, Chile; 45°53’S, 72°06’W

image of lot FMNH_179989
FMNH_179989
Lago Rinihue, Valdiva, Valdiva, Chile; 39°49’S, 72°28’W

image of lot FMNH_198223
FMNH_198223
Lago Taguatagua, Llanouihue, Chiloe, Chile; 41°42’S, 72°08’W

image of lot FMNH_179986
FMNH_179986
Lago Totoral, Llanquihue, Chiloe, Chile; 41°54’S, 71°54’W

image of lot FMNH_198228
FMNH_198228
Lago Totoral, Llanquihue, Chiloe, Chile; 41°54’S, 71°54’W

image of lot FMNH_198224
FMNH_198224
Lago Villarrica, Pucon, Villarica, Cautin, Chile; 39°16’S, 71°58’W

image of lot FMNH_198219
FMNH_198219
Lago Villarrica, Pucon, Villarica, Cautin, Chile; 39°16’S, 71°58’W

image of lot FMNH_21835
FMNH_21835
Lake Llanquihue, Chile

image of lot FMNH_198218
FMNH_198218
Los Ganchos, Estero; Traiguen, Bio-Bio, Chile; 38°19’S, 73°08’W

image of lot FMNH_198211
FMNH_198211
Los Ganchos, Traiguen, Bio-Bio, Chile; 38°19’S, 73°08’W

image of lot FMNH_179985
FMNH_179985
Mina Carmen, Illapel, Coquimbo, Chile; 32°06’S, 71°17’W

image of lot FMNH_121885
FMNH_121885
on river bank, E of Puerto Montt, Petroue, Chile

image of lot FMNH_94156
FMNH_94156
Puerto Montt, Chile

image of lot FMNH_179987
FMNH_179987
Puerto Pichidangui, Petorca, Aconcagua, Chile; 32°09’S, 71°33’W

image of lot FMNH_198227
FMNH_198227
Quebrada Centinela, at Lago Llanquihue, Llanquihue, Chiloe, Chile; 41°05’S, 73°13’W

image of lot FMNH_198213
FMNH_198213
Rio Choapa, Salamanca, Illapel, Coquimbo, Chile; 31°47’S, 70°58’W

image of lot FMNH_198220
FMNH_198220
Rio Cogoti (Las Tinajas), Ovalle, Coquimbo, Chile; 31°04’S, 71°02’W

image of lot FMNH_198215
FMNH_198215
Rio Higuera, La Higuera, Petorca, Aconcagua, Chile; 32°25’S, 71°25’W

image of lot FMNH_198225
FMNH_198225
Rio la Poza, Llanquihue, Valdiva, Chile; 40°17’S, 73°02’W

image of lot FMNH_198209
FMNH_198209
Rio Limari, Ovalle, Coquimbo, Chile; 30°36’S, 71°12’W

image of lot FMNH_21834
FMNH_21834
Rio Maullico, South Chile

image of lot FMNH_198214
FMNH_198214
Rio Puelo, Laguna de la Division, Llanquihue, Chiloe, Chile; 41°39’S, 72°19’W

image of lot FMNH_198212
FMNH_198212
Rio Puelo, Laguna Mallin Grande, Llanquihue, Chiloe, Chile; 41°39’S, 72°19’W

image of lot FMNH_179983
FMNH_179983
Rio Quilimari, Illapel, Coquimbo, Chile; 32°08’S, 71°32’W

image of lot FMNH_64970
FMNH_64970
Rio Valdivia, Chile

image of lot FMNH_120389
FMNH_120389
Valdivia, Chile

image of lot FMNH_89819
FMNH_89819
Valdivia, Chile

image of lot FMNH_198217
FMNH_198217
Lago Blanca, Tacna, Peru-Chile; 17°38’S, 69°38’W

1 Specimens from IRSNB

image of lot IRSNB_MP_0933
IRSNB_MP_0933
Bogota [No - Chile]

23 Specimens from MCZ

image of lot MCZ_MP.38
MCZ_MP.38
Lago Correntoso, Parque Nacional de Nahuel Huapi, Prov. Rio Negro, Argentina

image of lot MCZ_MP.50
MCZ_MP.50
Lago Correntoso, Parque Nacional de Nahuel Huapi, Prov. Rio Negro, Argentina

image of lot MCZ_MP.51
MCZ_MP.51
Lago Correntoso, Parque Nacional de Nahuel Huapi, Prov. Rio Negro, Argentina

image of lot MCZ_118706
MCZ_118706
Lago Nahuel-Huapi, Neuquen, Argentina

image of lot MCZ_MP.66
MCZ_MP.66
Cautin, Villarrica, Chile

image of lot MCZ_17030
MCZ_17030
Chile, South America

image of lot MCZ_MP.27
MCZ_MP.27
Chile, South America

image of lot MCZ_MP.14
MCZ_MP.14
Lago Esmeralda, Peulla, Chile

image of lot MCZ_MP.13
MCZ_MP.13
Lake Llanquihue, Chile

image of lot MCZ_MP.63
MCZ_MP.63
Peralillo, Departmento de Iltapel, Rio Choapa, Prov. Coquimbo, Chile

image of lot MCZ_17037
MCZ_17037
Rio Mayo, Mexico [Chile or Peru]

image of lot MCZ_197983
MCZ_197983
Rio Valdivia, S. Chile

image of lot MCZ_MP.37
MCZ_MP.37
Sao Paulo, Brazil. [must be Chile!]

image of lot MCZ_MP.34
MCZ_MP.34
Valdivia, Chile

image of lot MCZ_MP.35
MCZ_MP.35
South America

image of lot MCZ_17027
MCZ_17027
South America

image of lot MCZ_MP.19
MCZ_MP.19
South America

MCZ_MP.017. Chiloe Island, Chile

8 Specimens from MNHN

image of lot MNHN_MP_0985
MNHN_MP_0985
Rio Negro, Patagonia

image of lot MNHN_MP_0978
MNHN_MP_0978
Bolivie [No - Chile]

image of lot MNHN_MP_0866
MNHN_MP_0866
Island Chiloe

image of lot MNHN_MP_0991
MNHN_MP_0991
Province Llanguihue, Sud Chili

image of lot MNHN_MP_0967
MNHN_MP_0967
Quillota, Chili

image of lot MNHN_MP_1406
MNHN_MP_1406
Rio Carampangue, Chili

image of lot MNHN_MP_0878
MNHN_MP_0878
Riviere Maypo, Chili

image of lot MNHN_MP_0885
MNHN_MP_0885
Valdivia, Chile

2 Specimens from PUCRS-MCP

PUCRS-MCP_8868. Argentina: Neuquen: (Atlantic Ocean - Negro): Lago Lacar: Quila, Quina, próximo a San Martin de Los Andes, junto ao Lago Lacar

PUCRS-MCP_2854. Chile: Los Lagos: (Pacific Ocean): Chiloé, Região X de Los Lagos: -42.67667 -73.99333

47 Specimens from SMF

image of lot SMF_11219
SMF_11219
Araucaria

image of lot SMF_11233
SMF_11233
Arroyo Quelimalal, Prov. Neugen, Argentina

image of lot SMF_11238
SMF_11238
Lago Lacar, Argentinien

image of lot SMF_11236
SMF_11236
Patagonien

image of lot SMF_11237
SMF_11237
Rio Negro, b. Carmen de Patagones, Argent.

image of lot SMF_11232
SMF_11232
Rio Negro, b. Carmen de Patagones, Argent.

image of lot SMF_11240
SMF_11240
Rio Negro, b. Potrero Serrado, Patagonien

image of lot SMF_644
SMF_644
Rio Neuquén, Patagonien

image of lot SMF_11239
SMF_11239
See Nahuel Huape, Argent. Kordillere

image of lot SMF_11235
SMF_11235
Buenos Aires [No - Chile]

image of lot SMF_11217
SMF_11217
Concepcion, Chile

image of lot SMF_11215
SMF_11215
Conger River, Chile

image of lot SMF_168325
SMF_168325
Fluss Maullin, Sud Chile

image of lot SMF_11222
SMF_11222
Gegend [Area] von Valparaiso, Chile

image of lot SMF_11213
SMF_11213
Guayellua, Chile

image of lot SMF_11205
SMF_11205
Insel Chiloe, Chile

image of lot SMF_11204
SMF_11204
Insel Chiloe, Chile

image of lot SMF_11230
SMF_11230
Insel Chiloe, Chile

image of lot SMF_11225
SMF_11225
Insel Chiloe, Chile

image of lot SMF_11206
SMF_11206
Insel Chiloe, Chile

image of lot SMF_11229
SMF_11229
Insel Chiloe, Chile

image of lot SMF_11231
SMF_11231
Laguna Ganyetta, Chile

image of lot SMF_11207
SMF_11207
Prov. Llanquihue, Chile

image of lot SMF_168330
SMF_168330
Prov. Llanquihui, S. Chile

image of lot SMF_168327
SMF_168327
Prov. Llanquihui, S. Chile

image of lot SMF_168328
SMF_168328
Province Llanquihue, Chile

image of lot SMF_11228
SMF_11228
Quillota bei Valparaiso, Chile

image of lot SMF_11218
SMF_11218
Quillota bei Valparaiso, Chile

image of lot SMF_11226
SMF_11226
Quintero, nord van Valparasiso, Chile

image of lot SMF_11216
SMF_11216
Reloncavi bei Puerto Montt, S Chile

image of lot SMF_11212
SMF_11212
Rio Carampangue, Chile

image of lot SMF_11274
SMF_11274
Rio Kuljun, Chile

image of lot SMF_11210
SMF_11210
Rio Maule, M Chile

image of lot SMF_11220
SMF_11220
Rio Maule, S. Chile

image of lot SMF_11224
SMF_11224
Rio Maullin, S, Chile

image of lot SMF_11227
SMF_11227
Rio Maullin, S, Chile

image of lot SMF_11211
SMF_11211
Rio Maullin, S, Chile

image of lot SMF_11202
SMF_11202
S. Chile

image of lot SMF_11221
SMF_11221
San Juan, Prov. Valdivia, Chile

image of lot SMF_11208
SMF_11208
See Llanquihue, S, Chile

image of lot SMF_11223
SMF_11223
See Llanquihue, S, Chile

image of lot SMF_11209
SMF_11209
See Llanquihue, S, Chile

3 Specimens from UF

image of lot UF_270580
UF_270580
Peru ? [Chile]

image of lot UF_270582
UF_270582
Peru ? [Chile]

image of lot UF_270581
UF_270581
[no locality available - South America]

23 Specimens from UMMZ

image of lot UMMZ_22943
UMMZ_22943
Chile ? South America ?

image of lot UMMZ_111910
UMMZ_111910
Cholchol River, Imperial, Chile

image of lot UMMZ_111239
UMMZ_111239
Chotchol River, Imperial, Chile

image of lot UMMZ_111890
UMMZ_111890
Llanquihu Province, southern Chile

image of lot UMMZ_111238
UMMZ_111238
Moullin River, Southern Chile

image of lot UMMZ_111273
UMMZ_111273
Penaflor, Chile

image of lot UMMZ_111242
UMMZ_111242
Prov. Llanquihue, Chile

image of lot UMMZ_111882
UMMZ_111882
Rio Mayo, Mexico [Chile or Peru]

image of lot UMMZ_22928
UMMZ_22928
Valdivia, Chile

image of lot UMMZ_111891
UMMZ_111891
Valdivia, Chile

image of lot UMMZ_111240
UMMZ_111240
Valdivia, Chile

image of lot UMMZ_22921
UMMZ_22921
South America

image of lot UMMZ_22922
UMMZ_22922
South America

36 Specimens from USNM

image of lot USNM_359921
USNM_359921
L. Futalaufqua, Chubut, Argentina. Apparently drains into G de Corcovado and hence into the Pacific. Is in NW Chubut

image of lot USNM_359922
USNM_359922
L. Villarino, Argentina

image of lot USNM_628770
USNM_628770
Rio Negro, Patagonia

image of lot USNM_86105
USNM_86105
Rio Negro, S. America

image of lot USNM_762610
USNM_762610
ca Calafquen Lake, Valdiva Province, Chile

image of lot USNM_762611
USNM_762611
ca Temuco, Chile, Sta. 7

image of lot USNM_251239
USNM_251239
Chalchal River, Imperial, Chile

image of lot USNM_414170
USNM_414170
drainage canal, El Vergel, Angol, Chile

image of lot USNM_85622A
USNM_85622A
Id. of Chiloe, Chile

image of lot USNM_85619
USNM_85619
Isle of Chiloe, Chile

image of lot USNM_569009
USNM_569009
Isle of Chiloe, Chile

image of lot USNM_85626
USNM_85626
Isle of Chiloe, Chile

image of lot USNM_85623
USNM_85623
Peru [Chile]

image of lot USNM_85618
USNM_85618
Peru [Chile]

image of lot USNM_339726
USNM_339726
Prov. of Llanquihue, Chile

image of lot USNM_414171
USNM_414171
Ranaico R., Renaico, Chile

image of lot USNM_85620
USNM_85620
Rio Campagne, Chile

image of lot USNM_758637
USNM_758637
Rio Longotoma, Chile

image of lot USNM_523318
USNM_523318
Rio Mallico, at ford to Los Alpes, Angol, Chile

image of lot USNM_762619
USNM_762619
Temuco, Chile

image of lot USNM_622201
USNM_622201
Valdivia, Chile

image of lot USNM_504816
USNM_504816
Valdivia, Chile

image of lot USNM_758639
USNM_758639
Valdivia, Chile

image of lot USNM_131257
USNM_131257
Valparaiso, Chile

image of lot USNM_85622
USNM_85622
Valparaiso, Chile

image of lot USNM_85625
USNM_85625
Valparaiso, Chile

image of lot USNM_126300_01
USNM_126300_01
Behu, Malacca [no- South America]

image of lot USNM_40836
USNM_40836
Sth. America

image of lot USNM_58214
USNM_58214
W. S. America

37 Specimens from ZMB

image of lot ZMB_MP_0300
ZMB_MP_0300
Nahuel Huape, Argentina

image of lot ZMB_24898
ZMB_24898
Patagones

image of lot ZMB_34738
ZMB_34738
Rio Negro, Sud Brazil or Sud Argentinien

image of lot ZMB_MP_0282
ZMB_MP_0282
....Costa .. Contao, Chile

image of lot ZMB_MP_0311
ZMB_MP_0311
aus dem Rio Cruces nordlich von Valdivia

image of lot ZMB_MP_0305
ZMB_MP_0305
aus dem Rio Cruces nordlich von Valdivia

image of lot ZMB_9205
ZMB_9205
Chiloe

image of lot ZMB_MP_0284
ZMB_MP_0284
Estero del Potrerillo de las Yeguas en el eamino realei Chiloe

image of lot ZMB_48666
ZMB_48666
Frutillar, Chile

image of lot ZMB_MP_0229
ZMB_MP_0229
I. Chile

image of lot ZMB_MP_0299
ZMB_MP_0299
I. Chiloe

image of lot ZMB_MP_0322
ZMB_MP_0322
I. Chiloe

image of lot ZMB_MP_0338
ZMB_MP_0338
Ins. Chiloe

image of lot ZMB_MP_0317
ZMB_MP_0317
Ins. Chiloe

image of lot ZMB_91038
ZMB_91038
Lago Riso Paton, Pto Puynhuapi, Prov. Aysen, Chile

image of lot ZMB_MP_0314
ZMB_MP_0314
Mittel Chile, Villarica - See

image of lot ZMB_MP_0296
ZMB_MP_0296
Nancagua, Chile

image of lot ZMB_MP_0335
ZMB_MP_0335
Osorno, Valdivia

image of lot ZMB_52583
ZMB_52583
Puerto Montt

image of lot ZMB_52895
ZMB_52895
Puerto Montt, Chile

image of lot ZMB_MP_0279
ZMB_MP_0279
Rio Mayumo [?] Los Lagos [?], Chile

image of lot ZMB_MP_0316
ZMB_MP_0316
Valdivia

image of lot ZMB_MP_0309
ZMB_MP_0309
Valdivia

image of lot ZMB_MP_0321
ZMB_MP_0321
Valdivia, Chile

image of lot ZMB_MP_0319
ZMB_MP_0319
Valdivia, Chiloe

image of lot ZMB_MP_0298
ZMB_MP_0298
Valparaiso

image of lot ZMB_MP_0330
ZMB_MP_0330
Valpariaso u Santiago

image of lot ZMB_MP_0333
ZMB_MP_0333
[no locality available - South America]

image of lot ZMB_MP_0337
ZMB_MP_0337
[no locality available - South America]

image of lot ZMB_MP_0329
ZMB_MP_0329
[no locality available - South America]

image of lot ZMB_MP_0999
ZMB_MP_0999
[no locality available - South America]

12 Specimens from other collections

ZMA_210691. Lago Nahuel, Bariloche, Prov. de Rio Negro, Argentina

ZMA_14395. Lago Llanquihue, Frutillar, Chile

ZMA_419642. Lago Todos Los Santos, Petrohué, Chile

RMNH_326000. Lago Todos Los Santos, Petrohué, Chile

RMNH_325976. Laguna de los Jaras, Lonquimae [Lonquimay], prov. Caukin, Chili

RMNH_326530. Renaico River, Bio-Bio, Chile

ZMA_210628. Rio Aconcagua, Prov.de Valparaiso, Chile

RMNH_326532. Rio Valdivia, Chili

RMNH_326529. Rio Valdivia, Chili

ZMA_213666. Valdivia, Extre Madura [?], Spain [Chile]

ZMA_210010. X Region, S.Chile, Frutillar, L. Llanquihue

GenBank Vouchers

There are no georeferenced GenBank vouchers for this species in the MUSSELpdb.