The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!
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Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae | Anodontini | Cristariina genus Pletholophus Simpson, 1900: cladomics
4 Recent valid species (4 total species) Cladomics pages are experimental — even more so than the rest of this site!
The purpose of the cladomics pages for each genus and family-group level taxon is to
provide statistics and references indicating the extent to which published phylogenetic
work supports the classification used here. The Cladomics Summary provides an
overview of the number of studies that have included terminal species representing this
taxon as well as an indication of how frequently it has been recovered as monophyletic.
Following the summary, the available published clades representing the taxon are listed.
For more information, visit the Cladomics FARQ (frequently
asked rhetorical questions) page. Abbreviations for the various statistics are explained on
the FARQ! Cladomics Summarynumber of species = 4 (4 total)
analyzed in 34 cladograms in 17 publications
maximum number of terminals in an analysis =
4 (mean = 1.44, median = 1) Basic Monophylynumber of cladograms that test basic monophyly
(i.e., more than 1 included species) = 7
maximum number of included terminals = 4, (mean =
3.14 median = 3)
proportion of cladograms that support basic monophyly
= 0.86
Pairwise Cladogram ClusteringThe graphs below depict in two dimensions all pair-wise ingroup
distances within the cladome. See the Cladmoics
FARQ for more information.
Wu et al., 2025 [Publ 4442], fig. 2 ** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 12 | 10 | 4 | 31 | 29 | 7 | 0 | +----------------- Cristaria
|
| +----- Beringiana
--+ +--------+
| | | +-- Acudonta
| | +--+
| | +-- Sinanodonta
+--+
| +----- Pletholophus_guangzhouensis **
| +-----+
| | | +-- Pletholophus_tenuis **
| | +--+
+--+ +-- Pletholophus_reinianus **
|
| +-------- Pletholophus_honglinhensis **
+--+
| +----- Anemina
+--+
| +-- Amuranodonta
+--+
+-- Buldowskia
Dai et al., 2024 [Publ 4385], fig. 3-BI iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 27 | 23 | 6 | 8 | +----- Amuranodonta
+-----------+
| | +-- Anemina
| +--+
--+ +-- Buldowskia
|
| +-------------- Cristaria
| |
+--+ +-- Beringiana
| +--------+
| | +-- Sinanodonta
+--+
| +-------- Pletholophus_honglinhensis **
+--+
| +----- Pletholophus_guangzhouensis **
+--+
| +-- Pletholophus_tenuis **
+--+
+-- Pletholophus_reinianus **
Dai et al., 2024 [Publ 4385], fig. 3-ML iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 27 | 23 | 6 | 8 | +----- Amuranodonta
+-----------+
| | +-- Anemina
| +--+
--+ +-- Buldowskia
|
| +-------------- Cristaria
| |
+--+ +-- Beringiana
| +--------+
| | +-- Sinanodonta
+--+
| +-------- Pletholophus_honglinhensis **
+--+
| +----- Pletholophus_guangzhouensis **
+--+
| +-- Pletholophus_tenuis **
+--+
+-- Pletholophus_reinianus **
Bogan et al., 2023 [Publ 4295], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 3 | 2 | 2 | 9 | +----------- Cristaria
--+
| +-------- Sinanodonta
+--+
| +----- Pletholophus_honglinhensis **
+--+
| +-- Pletholophus_tenuis **
+--+
+-- Pletholophus_reinianus **
Xiang et al., 2025 [Publ 4433], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 25 | 24 | 7 | 9 | +----- Pletholophus_honglinhensis **
+--------+
| | +-- Pletholophus_guangzhouensis **
| +--+
| +-- Pletholophus_tenuis **
--+
| +----- Beringiana
| +-----+
| | | +-- Mimo
| | +--+
+--+ +-- Sinanodonta
|
| +-------- Cristaria
+--+
| +----- Anemina
+--+
| +-- Amuranodonta
+--+
+-- Buldowskia
Lopes-Lima et al., 2020 [Publ 4080], fig. 9 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 4 | 2 | 2 | -- | +-------- Cristaria
--+
| +----- Sinanodonta
+--+
| +-- Pletholophus_tenuis **
+--+
+-- Pletholophus_reinianus **
Bogan et al., 2023 [Publ 4295], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 29 | 25 | 6 | -- | +----------- Cristaria
|
| +----- Amuranodonta
--+ +--+
| | | +-- Anemina
| | +--+
| | +-- Buldowskia
+--+
| +-- Pletholophus_honglinhensis **
| +--+
| | +-- Pletholophus_tenuis **
+--+
| +-- Beringiana
+--+
+-- Sinanodonta
Dai et al., 2025 [Publ 4440], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 44 | 5 | 5 | -- | +----- Pletholophus_reinianus **
--+
| +-- Sinanodonta
+--+
+-- Beringiana
Bolotov et al., 2017 [Publ 3764], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 124 | 5 | 2 | -- | -- Pletholophus_tenuis **
Sano et al., 2022 [Publ 4311], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 17 | 17 | 4 | -- | +-- Beringiana
+-----+
| +-- Sinanodonta
--+
| +----- Pletholophus_reinianus **
+--+
| +-- Cristaria
+--+
+-- Buldowskia
Huang et al., 2019 [Publ 3843], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 198 | 8 | 3 | -- | +-------- Anemina
--+
| +----- Cristaria
+--+
| +-- Pletholophus_tenuis **
+--+
+-- Sinanodonta
Huang et al., 2019 [Publ 3843], fig. S4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 198 | 8 | 3 | -- | +-- Pletholophus_tenuis **
--+
+-- Sinanodonta
Bogan et al., 2023 [Publ 4295], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 6 | 5 | 2 | -- | +----- Cristaria
--+
| +-- Sinanodonta
+--+
+-- Pletholophus_honglinhensis **
Bolotov et al., 2017 [Publ 3761], fig. S2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 138 | 5 | 2 | -- | +----- Cristaria
--+
| +-- Pletholophus_tenuis **
+--+
+-- Sinanodonta
Sano et al., 2020 [Publ 4138], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 67 | 10 | 5 | -- | +----- Pletholophus_reinianus **
--+
| +-- Beringiana
+--+
+-- Sinanodonta
Sano et al., 2017 [Publ 3799], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 32 | 8 | 4 | -- | +-- Cristaria
+-----+
| +-- Buldowskia
--+
| +----- Beringiana
+--+
| +-- Pletholophus_reinianus **
+--+
+-- Sinanodonta
Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 75 | 4 | 2 | -- | +----- Cristaria
--+
| +-- Pletholophus_tenuis **
+--+
+-- Sinanodonta
Dai et al., 2025 [Publ 4440], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 44 | 5 | 5 | -- | +----- Pletholophus_reinianus **
--+
| +-- Sinanodonta
+--+
+-- Beringiana
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 79 | 4 | 3 | -- | +-------- Cristaria
--+
| +----- Anemina
+--+
| +-- Pletholophus_tenuis **
+--+
+-- Sinanodonta
Kano et al., 2019 [Publ 4194], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 2 | 2 | -- | +----- Sinanodonta
--+
| +-- Pletholophus_tenuis **
+--+
+-- Beringiana
Bogan et al., 2023 [Publ 4295], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 12 | 11 | 2 | -- | +----- Pletholophus_honglinhensis **
--+
| +-- Cristaria
+--+
+-- Sinanodonta
Bolotov et al., 2017 [Publ 3761], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 138 | 5 | 2 | -- | +----- Cristaria
--+
| +-- Pletholophus_tenuis **
+--+
+-- Sinanodonta
Sano et al., 2022 [Publ 4311], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 11 | 4 | -- | +-- Beringiana
+-----+
| +-- Sinanodonta
--+
| +----- Pletholophus_reinianus **
+--+
| +-- Cristaria
+--+
+-- Buldowskia
Sano et al., 2020 [Publ 4138], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 9 | 5 | -- | -- Pletholophus_reinianus **
Sano et al., 2017 [Publ 3799], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 20 | 7 | 4 | -- | +-- Cristaria
+-----+
| +-- Buldowskia
--+
| +----- Pletholophus_reinianus **
+--+
| +-- Sinanodonta
+--+
+-- Beringiana
Chen et al., 2023 [Publ 4302], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 39 | 5 | 5 | -- | +----------- Cristaria
|
--+ +-- Anemina
| +-----+
| | +-- Amuranodonta
+--+
| +----- Pletholophus_tenuis **
+--+
| +-- Sinanodonta
+--+
+-- Beringiana
Sano et al., 2020 [Publ 4138], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 67 | 10 | 5 | -- | +----- Sinanodonta
--+
| +-- Pletholophus_reinianus **
+--+
+-- Beringiana
Wu et al., 2025 [Publ 4442], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 16 | 14 | 7 | -- | +----------- Cristaria
|
| +----- Amuranodonta
--+ +--+
| | | +-- Anemina
| | +--+
| | +-- Buldowskia
+--+
| +-- Beringiana
| +--+
| | +-- Pletholophus_guangzhouensis **
+--+
| +-- Acudonta
+--+
+-- Sinanodonta
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 120 | 23 | 6 | -- | +-------- Cristaria
|
| +----- Anemina
+--+
--+ | +-- Amuranodonta
| +--+
| +-- Buldowskia
|
| +----- Pletholophus_tenuis **
+--+
| +-- Beringiana
+--+
+-- Sinanodonta
Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 75 | 4 | 2 | -- | +----- Cristaria
--+
| +-- Pletholophus_tenuis **
+--+
+-- Sinanodonta
Sano et al., 2020 [Publ 4138], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 67 | 10 | 5 | -- | +----- Sinanodonta
--+
| +-- Pletholophus_reinianus **
+--+
+-- Beringiana
Bolotov et al., 2018 [Publ 3826], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 82 | 3 | 2 | -- | +----- Cristaria
--+
| +-- Pletholophus_tenuis **
+--+
+-- Sinanodonta
Chen et al., 2023 [Publ 4302], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 39 | 5 | 5 | -- | +----- Pletholophus_tenuis **
--+
| +-- Sinanodonta
+--+
+-- Beringiana
Huang et al., 2019 [Publ 3843], fig. S3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 196 | 8 | 3 | -- | +-------- Anemina
--+
| +----- Cristaria
+--+
| +-- Pletholophus_tenuis **
+--+
+-- Sinanodonta
|
