Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Anodontini | Cristariina

genus Pletholophus Simpson, 1900: cladomics

4 Recent valid species (4 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 4 (4 total)
analyzed in 42 cladograms in 24 publications
maximum number of terminals in an analysis = 4 (mean = 1.57, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 10
maximum number of included terminals = 4, (mean = 3.4 median = 4)
proportion of cladograms that support basic monophyly = 0.9

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Pletholophus cladogram distances
PCoA Ordination

NJ Tree of Pletholophus cladogram distances
NJ Tree

Cladograms

Chen et al., 2026 [Publ 4805], fig. 4

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 30 29 8 **
Cladomics cladogram Cristaria Beringiana Mimo Acudonta Sinanodonta Anemina Amuranodonta Buldowskia Pletholophus honglinhensis Pletholophus guangzhouensis Pletholophus reinianus Pletholophus tenuis

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cst iT tT bi riS oT roT roS iD
0.900 4 4 2 4 629 29 8 2.0
Cladomics cladogram Mimo Acudonta Sinanodonta Beringiana Pletholophus honglinhensis Pletholophus tenuis Pletholophus reinianus Pletholophus guangzhouensis

Wu et al., 2025 [Publ 4442], fig. 2

cst iT tT bi riS oT roT roS iD
0.889 4 12 10 4 31 29 7 8.0
Cladomics cladogram Cristaria Beringiana Acudonta Sinanodonta Pletholophus guangzhouensis Pletholophus tenuis Pletholophus reinianus Pletholophus honglinhensis Anemina Amuranodonta Buldowskia

Dai et al., 2024 [Publ 4385], fig. 3-BI

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 27 23 6 0.0
Cladomics cladogram Amuranodonta Anemina Buldowskia Cristaria Beringiana Sinanodonta Pletholophus honglinhensis Pletholophus guangzhouensis Pletholophus tenuis Pletholophus reinianus

Dai et al., 2024 [Publ 4385], fig. 3-ML

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 27 23 6 0.0
Cladomics cladogram Amuranodonta Anemina Buldowskia Cristaria Beringiana Sinanodonta Pletholophus honglinhensis Pletholophus guangzhouensis Pletholophus tenuis Pletholophus reinianus

Dai et al., 2026 [Publ 4757], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 30 26 6 0.0
Cladomics cladogram Amuranodonta Anemina Buldowskia Cristaria Beringiana Sinanodonta Pletholophus honglinhensis Pletholophus guangzhouensis Pletholophus tenuis Pletholophus reinianus

Xiang et al., 2025 [Publ 4433], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 25 24 7 1.0
Cladomics cladogram Pletholophus honglinhensis Pletholophus guangzhouensis Pletholophus tenuis Beringiana Mimo Sinanodonta Cristaria Anemina Amuranodonta Buldowskia

Bogan et al., 2023 [Publ 4295], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 3 2 2 1.0
Cladomics cladogram Cristaria Sinanodonta Pletholophus honglinhensis Pletholophus tenuis Pletholophus reinianus

Bogan et al., 2023 [Publ 4295], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 29 25 6 --
Cladomics cladogram Cristaria Amuranodonta Anemina Buldowskia Pletholophus honglinhensis Pletholophus tenuis Beringiana Sinanodonta

Lopes-Lima et al., 2020 [Publ 4080], fig. 9

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 4 2 2 --
Cladomics cladogram Cristaria Sinanodonta Pletholophus tenuis Pletholophus reinianus

Wu et al., 2025 [Publ 4442], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 16 14 7 --
Cladomics cladogram Cristaria Amuranodonta Anemina Buldowskia Beringiana Pletholophus guangzhouensis Acudonta Sinanodonta

Wang et al., 2025 [Publ 4488], fig. 3B

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 63 14 7 --
Cladomics cladogram Cristaria Amuranodonta Anemina Buldowskia Beringiana Pletholophus guangzhouensis Acudonta Sinanodonta

Wu et al., 2026 [Publ 4811], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 94 14 7 --
Cladomics cladogram Cristaria Amuranodonta Anemina Buldowskia Beringiana Pletholophus guangzhouensis Acudonta Sinanodonta

Wu et al., 2026 [Publ 4825], fig. 6B

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 64 13 7 --
Cladomics cladogram Cristaria Amuranodonta Anemina Buldowskia Beringiana Pletholophus guangzhouensis Acudonta Sinanodonta

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 121 23 6 --
Cladomics cladogram Cristaria Anemina Amuranodonta Buldowskia Pletholophus tenuis Beringiana Sinanodonta

Sano et al., 2020 [Publ 4138], fig. 2

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 68 10 5 --
Cladomics cladogram Pletholophus reinianus Beringiana Sinanodonta

Sano et al., 2020 [Publ 4138], fig. 3

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 68 10 5 --
Cladomics cladogram Sinanodonta Pletholophus reinianus Beringiana

Sano et al., 2020 [Publ 4138], fig. 4

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 68 10 5 --
Cladomics cladogram Sinanodonta Pletholophus reinianus Beringiana

Sano et al., 2020 [Publ 4138], fig. 1

cst iT tT bi riS oT roT roS iD
0.286 1 1 0 1 33 9 5 --
Cladomics cladogram Lanceolaria oxyrhyncha 1 Lanceolaria oxyrhyncha 2 Cristaria plicata 1 Cristaria plicata 2 Sinanodonta lauta Beringiana japonica Beringiana fukuharai 1 Beringiana beringiana Beringiana fukuharai 2 Beringiana fukuharai 3 Buldowskia iwakawai 1 Buldowskia iwakawai 2 Sinanodonta tumens 1 Sinanodonta tumens 2 Pletholophus reinianus Anemina Lanceolaria lanceolata Cristaria plicata 3 Lanceolaria gladiola Sinanodonta angula Lepidodesma Scriptolamprotula Acuticosta Cuneopsis Nodularia Schistodesmus Inversiunio

Chen et al., 2023 [Publ 4302], fig. 1

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 40 5 5 --
Cladomics cladogram Pletholophus tenuis Sinanodonta Beringiana

Chen et al., 2023 [Publ 4302], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 5 5 --
Cladomics cladogram Cristaria Anemina Amuranodonta Pletholophus tenuis Sinanodonta Beringiana

Dai et al., 2025 [Publ 4440], fig. 1

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 43 5 5 --
Cladomics cladogram Pletholophus tenuis Sinanodonta Beringiana

Dai et al., 2025 [Publ 4440], fig. 2

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 43 5 5 --
Cladomics cladogram Pletholophus tenuis Sinanodonta Beringiana

Chen et al., 2025 [Publ 4487], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 5 5 --
Cladomics cladogram Cristaria Anemina Amuranodonta Pletholophus tenuis Sinanodonta Beringiana

Chen et al., 2025 [Publ 4487], fig. 3

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 40 5 5 --
Cladomics cladogram Anemina Amuranodonta Pletholophus tenuis Sinanodonta Beringiana

Sano et al., 2017 [Publ 3799], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 20 7 4 --
Cladomics cladogram Cristaria Buldowskia Pletholophus reinianus Sinanodonta Beringiana

Sano et al., 2017 [Publ 3799], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 32 8 4 --
Cladomics cladogram Cristaria Buldowskia Beringiana Pletholophus reinianus Sinanodonta

Sano et al., 2022 [Publ 4311], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 11 11 4 --
Cladomics cladogram Beringiana Sinanodonta Pletholophus reinianus Cristaria Buldowskia

Sano et al., 2022 [Publ 4311], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 17 17 4 --
Cladomics cladogram Beringiana Sinanodonta Pletholophus reinianus Cristaria Buldowskia

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 80 4 3 --
Cladomics cladogram Cristaria Anemina Pletholophus tenuis Sinanodonta

Huang et al., 2019 [Publ 3843], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 200 8 3 --
Cladomics cladogram Anemina Cristaria Pletholophus tenuis Sinanodonta

Huang et al., 2019 [Publ 3843], fig. S3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 198 8 3 --
Cladomics cladogram Anemina Cristaria Pletholophus tenuis Sinanodonta

Huang et al., 2019 [Publ 3843], fig. S4

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 200 8 3 --
Cladomics cladogram Pletholophus tenuis Sinanodonta

Bolotov et al., 2017 [Publ 3761], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 138 5 2 --
Cladomics cladogram Cristaria Pletholophus tenuis Sinanodonta

Bolotov et al., 2017 [Publ 3761], fig. S2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 138 5 2 --
Cladomics cladogram Cristaria Pletholophus tenuis Sinanodonta

Bolotov et al., 2017 [Publ 3764], fig. S1

cst iT tT bi riS oT roT roS iD
0.250 1 1 0 1 124 5 2 --
Cladomics cladogram Alasmidonta Anodonta anatina Anodonta cygnea Brachyanodon Lasmigona Pseudanodonta Pyganodon Simpsonaias Strophitus Sinanodonta lauta Cristaria plicata Cristaria radiata Pletholophus tenuis Sinanodonta lucida Sinanodonta jourdyi Lanceolaria gladiola Lanceolaria grayii Lanceolaria danzhouensis

Bolotov et al., 2018 [Publ 3826], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 81 3 2 --
Cladomics cladogram Cristaria Pletholophus tenuis Sinanodonta

Kano et al., 2019 [Publ 4194], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 2 2 --
Cladomics cladogram Sinanodonta Pletholophus tenuis Beringiana

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 75 4 2 --
Cladomics cladogram Cristaria Pletholophus tenuis Sinanodonta

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 75 4 2 --
Cladomics cladogram Cristaria Pletholophus tenuis Sinanodonta

Bogan et al., 2023 [Publ 4295], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 6 5 2 --
Cladomics cladogram Cristaria Sinanodonta Pletholophus honglinhensis

Bogan et al., 2023 [Publ 4295], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 12 11 2 --
Cladomics cladogram Pletholophus honglinhensis Cristaria Sinanodonta