Shirai, A., T. Kondo & T. Kajita. 2010. Molecular markers reveal genetic contamination of endangered freshwater pearl mussels in pearl culture farms in Japan . Venus 68: 151-163. https://doi.org/10.18941/venus.68.3-4_151.
(click the figure number to see the cladogram)
+----- Pronodularia_japanensis
--+
| +-- Sinohyriopsis_cumingii
+--+
+-- Sinohyriopsis_schlegelii