Zanatta, D.T. & A.T. Harris. 2013. Phylogeography and genetic variability of the freshwater mussels (Bivalvia: Unionidae) Ellipse, Venustaconcha ellipsiformis (Conrad 1836), and Bleeding Tooth, V. pleasii (Marsh 1891) . American Malacological Bulletin 31(2): 267-279. Available electronically.
(click the figure number to see the cladogram)
+----------- Pyganodon_grandis
--+
| +-------- Lampsilis_cardium
+--+
| +----- Epioblasma_rangiana
+--+
| +-- Venustaconcha_pleasii
+--+
+-- Venustaconcha_ellipsiformis