Page last updated
Wed 1 October 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae | Unionini | Unio

species Unio mancus Lamarck, 1819: specimens
     135 available nominal species synonyms (200 total synonyms)

Recent Taxonomic Opinions

Unio mancus — Graf (2007), Graf & Cummings (2007), Graf & Cummings (2007), Araujo et al. (2009), Dhora (2009), ... Wu et al. (2024), Wu et al. (2024), Dai et al. (2024), Hou et al. (2024), Dai et al. (2025)
Unio delphinus — Araujo et al. (2009)
Unio mancus mancus — Prie et al. (2012), Prie (2012), Prié & Puillandre (2014), Lopes-Lima et al. (2017), Marrone et al. (2019)
Unio mancus turtonii — Prie (2012), Prie et al. (2012), Prié & Puillandre (2014), Lopes-Lima et al. (2017), Marrone et al. (2019), Lyubas et al. (2022)
Unio elongatulus turtoni — Fauna Europaea (2012)
Unio elongatulus requienii — Fauna Europaea (2012)
Unio mancus requienii — Prie et al. (2012), Prie (2012), Prié & Puillandre (2014), Lopes-Lima et al. (2017), Marrone et al. (2019), Lyubas et al. (2022)
Unio elongatulus gargottae — Fauna Europaea (2012)
Unio mancus moquinianus — Prie (2012), Prié & Puillandre (2014)
Unio elongatulus moquinianus — Fauna Europaea (2012)
Unio mancus aleronii — Prie (2012)
Unio elongatulus rousii — Fauna Europaea (2012)
Unio elongatulus lawleyianus — Fauna Europaea (2012)
Unio mancus brindosianus — Prie (2012)
Unio elongatulus bourgeticus — Fauna Europaea (2012)
Unio mancus bourgeticus — Prie (2012), Prié & Puillandre (2014)
Unio elongatulus mancus — Fauna Europaea (2012)
Unio pictorum — He & Zhuang (2013), Prié & Puillandre (2014), Araujo et al. (2018)
Unio mancus var. aleronii — Prié & Puillandre (2014)
Unio mancus gargottae — Lopes-Lima et al. (2017)
Unio cf. elongatulus — Lopes-Lima et al. (2017)
— Lopes-Lima et al. (2024)

Range

Spain to Italy, in basins draining to the Mediterranean.

Map of Georeferenced Specimens

MNHN_22466. La Saône, à Mâcon (Saône-et-Loire)

France

MNHN_24472. Aramon (Gard)

MNHN_22265. Bois-Vieux (Jura)

MNHN_22267. Espalion (Aveyron)

MNHN_22224. Fréjus (Var)

MNHN_22266. l’Osse (Gers)

MNHN_22514. l’Ouvez (Ardèche)

MNHN_22578. La Brizotte, près d’Auxonne (Cote-d’Or)

MNHN_22089. La Grande-Garonne, à Fréjus (Var)

MNHN_22434. La Grande-Garonne, à Fréjus (Var)

MNHN_22158. La Loire à Nantes (Loire-Infereur)

MNHN_22513. La Loire à Nantes (Loire-Infereur)

MNHN_22426. La Marne à Lagny (Seine-et-Marne)

MNHN_22516. La Meurthe-et-Moselle

MNHN_22383. La Saône, à Auxonne (Côte-d’Or)

MNHN_22517. La Saône, à Collonges (Rhône)

MNHN_22515. la Saône, à Saint-Germain au-Mont-d’Or

MNHN_22580. la Saône, à Saint-Germain-au-Mont-d’Or

MNHN_22207. La Siagne, à Cannes (Alpes-Maritimes)

MNHN_22579. la Tille, près d’Auxonne (Cote-d’Or)

MNHN_22582. Le Menthon (Ain)

MNHN_22469. le Rhône à Irigny (Rhône)

MNHN_22468. Le Rhône au confluent

MNHN_22581. Le Rhône, à Avignon

MNHN_22597. Le Rhône, à Avignon

MNHN_22584. le Rhône, à Beaucaire (Gard)

MNHN_22510. Pentoise (Jura)

MNHN_22467. Saint-Laurent-d’Ain près Mâcon

MNHN_22583. Varennes (Jura)

MNHN_22090. Vidauban (Var)

Georeferenced Genbank Vouchers

France

MNCN_N134. (16S: KX400414, COI: KX400028, go to Genbank) Bourget Lake (Rhone) River, France.

MNCN_N135. (16S: KX400415, COI: KX400029, go to Genbank) ditto.

MNCN_N136. (COI: KX400030, go to Genbank) ditto.

MNCN_N137. (16S: KX400416, go to Genbank) ditto.

MNCN_N138. (16S: KX400419, go to Genbank) ditto.

MNCN_N139. (16S: KX400421, go to Genbank) ditto.

MNCN_N140. (16S: KX400426, go to Genbank) ditto.

MNCN_N141. (COI: KX400040, go to Genbank) ditto.

MNCN_N143. (COI: KX400041, go to Genbank) ditto.

MNCN_N147. (COI: KX400044, go to Genbank) ditto.

MNCN_N148. (COI: KX400045, go to Genbank) ditto.

MNCN_N149. (COI: KX400046, go to Genbank) ditto.

MNCN_N150. (16S: KX400431, COI: KX400047, go to Genbank) ditto.

MNCN_N133. (16S: KX400413, COI: KX400027, go to Genbank) Bourget Lake (Rhone), France.

MNHN_IM-2009-22386. (16S: KC703602, 28S: KC703851, COI: KC703950, go to Genbank) France: Ain River.

MNHN_IM-2009-22387. (16S: KC703603, 28S: KC703852, COI: KC703951, go to Genbank) ditto.

MNHN_IM-2009-22388. (16S: KC703604, 28S: KC703853, COI: KC703952, go to Genbank) ditto.

MNHN_IM-2009-22389. (16S: KC703605, 28S: KC703854, COI: KC703953, go to Genbank) ditto.

MNHN_IM-2009-22395. (16S: KC703610, 28S: KC703859, COI: KC703958, go to Genbank) ditto.

MNHN_IM-2009-17765. (16S: KC703460, 28S: KC703705, go to Genbank) France: Aisne River.

MNHN_IM-2009-17803. (16S: KC703483, 28S: KC703728, COI: JX046638, go to Genbank) ditto.

MNHN_IM-2009-12582. (16S: KC703365, 28S: KC703631, COI: JX046565, go to Genbank) France: Ardèche River.

MNHN_IM-2009-12571. (16S: KC703355, 28S: KC703625, COI: JX046556, go to Genbank) France: Argens River.

MNHN_IM-2009-12579. (16S: KC703363, COI: JX046563, go to Genbank) ditto.

MNHN_IM-2009-21274. (16S: KC703574, 28S: KC703823, COI: JX046678, go to Genbank) France: Authion River.

MNHN_IM-2009-21275. (16S: KC703575, 28S: KC703824, COI: JX046679, go to Genbank) ditto.

MNHN_IM-2009-21276. (16S: KC703576, 28S: KC703825, COI: JX046680, go to Genbank) ditto.

MNHN_IM-2009-17815. (16S: KC703493, 28S: KC703740, COI: JX046647, go to Genbank) France: Basse River.

MNHN_IM-2009-17816. (16S: KC703494, 28S: KC703741, COI: JX046648, go to Genbank) ditto.

MNHN_IM-2009-17817. (16S: KC703495, 28S: KC703742, COI: JX046649, go to Genbank) ditto.

MNHN_IM-2009-17818. (16S: KC703496, 28S: KC703743, COI: JX046650, go to Genbank) ditto.

MNHN_IM-2009-17819. (28S: KC703744, COI: JX046651, go to Genbank) ditto.

MNHN_IM-2009-17820. (16S: KC703497, 28S: KC703745, COI: JX046652, go to Genbank) ditto.

MNHN_IM-2009-17821. (16S: KC703498, 28S: KC703746, COI: JX046653, go to Genbank) ditto.

MNHN_IM-2009-12710. (16S: KC703426, 28S: KC703671, COI: JX046592, go to Genbank) France: Belle River.

MNHN_IM-2009-12711. (16S: KC703427, 28S: KC703672, COI: JX046593, go to Genbank) ditto.

MNHN_IM-2009-12712. (16S: KC703428, 28S: KC703673, go to Genbank) ditto.

MNHN_IM-2009-12713. (16S: KC703429, 28S: KC703674, COI: JX046594, go to Genbank) ditto.

MNHN_IM-2009-12714. (16S: KC703430, 28S: KC703675, COI: JX046595, go to Genbank) ditto.

MNHN_IM-2009-12591. (16S: KC703368, COI: JX046568, go to Genbank) France: Boivre River.

MNHN_IM-2009-12581. (16S: KC703364, 28S: KC703630, COI: JX046564, go to Genbank) France: Cèze River.

MNHN_IM-2009-21293. (16S: KC703587, 28S: KC703836, COI: JX046691, go to Genbank) France: Cher River.

MNHN_IM-2009-21294. (16S: KC703588, 28S: KC703837, COI: JX046692, go to Genbank) ditto.

MNHN_IM-2009-21295. (16S: KC703589, 28S: KC703838, COI: JX046693, go to Genbank) ditto.

MNHN_IM-2009-21296. (16S: KC703590, 28S: KC703839, COI: JX046694, go to Genbank) ditto.

MNHN_IM-2009-21298. (16S: KC703592, 28S: KC703841, COI: JX046696, go to Genbank) ditto.

MNHN_IM-2009-21299. (16S: KC703593, 28S: KC703842, COI: JX046697, go to Genbank) ditto.

MNHN_IM-2009-21300. (16S: KC703594, 28S: KC703843, COI: JX046698, go to Genbank) ditto.

MNHN_IM-2009-21267. (16S: KC703569, 28S: KC703818, COI: JX046675, go to Genbank) France: Drée River.

MNHN_IM-2009-21268. (16S: KC703570, 28S: KC703819, COI: JX046676, go to Genbank) ditto.

MNHN_IM-2009-21269. (16S: KC703571, 28S: KC703820, COI: JX046677, go to Genbank) ditto.

MNHN_IM-2009-21270. (16S: KC703572, 28S: KC703821, go to Genbank) ditto.

MNHN_IM-2009-21271. (16S: KC703573, 28S: KC703822, go to Genbank) ditto.

MNHN_IM-2009-17785. (16S: KC703466, 28S: KC703711, go to Genbank) France: Ellé River.

MNHN_IM-2009-17786. (16S: KC703467, 28S: KC703712, go to Genbank) ditto.

MNHN_IM-2009-17787. (16S: KC703468, 28S: KC703713, COI: JX046632, go to Genbank) ditto.

MNHN_IM-2009-17788. (16S: KC703469, 28S: KC703714, COI: JX046633, go to Genbank) ditto.

MNHN_IM-2009-21277. (16S: KC703577, 28S: KC703826, COI: JX046681, go to Genbank) France: Etrepes River.

MNHN_IM-2009-21278. (16S: KC703578, 28S: KC703827, COI: JX046682, go to Genbank) ditto.

MNHN_IM-2009-21279. (16S: KC703579, 28S: KC703828, COI: JX046683, go to Genbank) ditto.

MNHN_IM-2009-21280. (16S: KC703580, 28S: KC703829, COI: JX046684, go to Genbank) ditto.

MNHN_IM-2009-21281. (16S: KC703581, 28S: KC703830, COI: JX046685, go to Genbank) ditto.

MNHN_IM-2009-21282. (16S: KC703582, 28S: KC703831, COI: JX046686, go to Genbank) ditto.

MNHN_IM-2009-21283. (16S: KC703583, 28S: KC703832, COI: JX046687, go to Genbank) ditto.

MNHN_IM-2009-17784. (16S: KC703465, 28S: KC703710, COI: JX046631, go to Genbank) France: Golo River.

MNHN_IM-2009-12569. (16S: KC703353, 28S: KC703623, COI: JX046554, go to Genbank) France: Hérault River.

MNHN_IM-2009-12572. (16S: KC703356, COI: JX046557, go to Genbank) ditto.

MNHN_IM-2009-22391. (16S: KC703607, 28S: KC703856, COI: KC703955, go to Genbank) France: Lake Aiguebelette.

MNHN_IM-2009-12701. (16S: KC703417, 28S: KC703662, COI: JX046586, go to Genbank) France: Lake Bourget.

MNHN_IM-2009-12702. (16S: KC703418, 28S: KC703663, go to Genbank) ditto.

MNHN_IM-2009-12704. (16S: KC703420, 28S: KC703665, COI: JX046588, go to Genbank) ditto.

MNHN_IM-2009-12706. (16S: KC703422, 28S: KC703667, COI: JX046590, go to Genbank) ditto.

MNHN_IM-2009-12707. (16S: KC703423, 28S: KC703668, go to Genbank) ditto.

MNHN_IM-2009-12708. (16S: KC703424, 28S: KC703669, COI: JX046591, go to Genbank) ditto.

MNHN_IM-2009-12709. (16S: KC703425, 28S: KC703670, go to Genbank) ditto.

MNHN_IM-2009-12570. (16S: KC703354, 28S: KC703624, COI: JX046555, go to Genbank) France: Lamalou River.

MNHN_IM-2009-12598. (16S: KC703370, 28S: KC703635, COI: JX046569, go to Genbank) France: Loutz River.

MNHN_IM-2009-12599. (16S: KC703371, COI: JX046570, go to Genbank) ditto.

MNHN_IM-2009-12715. (16S: KC703431, 28S: KC703676, COI: JX046596, go to Genbank) France: Né (Charente) River.

MNHN_IM-2009-12716. (16S: KC703432, 28S: KC703677, COI: JX046597, go to Genbank) ditto.

MNHN_IM-2009-12717. (16S: KC703433, 28S: KC703678, COI: JX046598, go to Genbank) ditto.

MNHN_IM-2009-12735. (16S: KC703441, 28S: KC703686, COI: JX046609, go to Genbank) France: Orbu River.

MNHN_IM-2009-12736. (16S: KC703442, 28S: KC703687, COI: JX046610, go to Genbank) ditto.

MNHN_IM-2009-12747. (16S: KC703443, 28S: KC703688, COI: JX046615, go to Genbank) ditto.

MNHN_IM-2009-12748. (16S: KC703444, 28S: KC703689, COI: JX046616, go to Genbank) ditto.

MNHN_IM-2009-12752. (16S: KC703445, 28S: KC703690, go to Genbank) ditto.

MNHN_IM-2009-21218. (16S: KC703528, 28S: KC703776, COI: JX046662, go to Genbank) France: Ourcq River.

MNHN_IM-2009-21220. (28S: KC703778, go to Genbank) ditto.

MNHN_IM-2009-21222. (16S: KC703531, 28S: KC703780, COI: JX046663, go to Genbank) ditto.

MNHN_IM-2009-21224. (16S: KC703532, 28S: KC703781, COI: JX046664, go to Genbank) ditto.

MNHN_IM-2009-21225. (16S: KC703533, 28S: KC703782, go to Genbank) ditto.

MNHN_IM-2009-21228. (16S: KC703536, 28S: KC703785, COI: JX046665, go to Genbank) ditto.

MNHN_IM-2009-21229. (16S: KC703537, 28S: KC703786, COI: JX046666, go to Genbank) ditto.

MNHN_IM-2009-21231. (16S: KC703539, 28S: KC703788, COI: JX046667, go to Genbank) ditto.

MNHN_IM-2009-21235. (16S: KC703541, 28S: KC703790, COI: JX046670, go to Genbank) ditto.

MNHN_IM-2009-17810. (16S: KC703490, 28S: KC703735, COI: JX046643, go to Genbank) France: Petit Morin River.

MNHN_IM-2009-17812. (28S: KC703737, COI: JX046644, go to Genbank) ditto.

MNHN_IM-2009-17813. (16S: KC703492, 28S: KC703738, COI: JX046645, go to Genbank) ditto.

MNHN_IM-2009-17814. (28S: KC703739, COI: JX046646, go to Genbank) ditto.

MNHN_IM-2009-12605. (16S: KC703375, 28S: KC703638, COI: JX046573, go to Genbank) France: Rhône River.

MNHN_IM-2009-21208. (28S: KC703766, go to Genbank) ditto.

MNHN_IM-2009-22392. (16S: KC703608, 28S: KC703857, COI: KC703956, go to Genbank) France: Saône River.

MNHN_IM-2009-21242. (16S: KC703547, 28S: KC703797, COI: JX046672, go to Genbank) France: Seine River.

MNHN_IM-2009-21245. (16S: KC703549, 28S: KC703799, COI: JX046673, go to Genbank) ditto.

MNHN_IM-2009-12602. (16S: KC703373, COI: JX046571, go to Genbank) France: Seugne River.

MNHN_IM-2009-12604. (16S: KC703374, 28S: KC703637, COI: JX046572, go to Genbank) ditto.

MNHN_IM-2009-12699. (16S: KC703416, 28S: KC703661, COI: JX046585, go to Genbank) France: Suran River.

MNHN_IM-2009-17806. (16S: KC703486, 28S: KC703731, COI: JX046639, go to Genbank) ditto.

MNHN_IM-2009-17807. (16S: KC703487, 28S: KC703732, COI: JX046640, go to Genbank) ditto.

MNHN_IM-2009-17808. (16S: KC703488, 28S: KC703733, COI: JX046641, go to Genbank) ditto.

MNHN_IM-2009-17809. (16S: KC703489, 28S: KC703734, COI: JX046642, go to Genbank) ditto.

MNHN_IM-2009-12723. (16S: KC703437, 28S: KC703682, go to Genbank) France: Taravu River.

MNHN_IM-2009-12724. (16S: KC703438, 28S: KC703683, go to Genbank) ditto.

MNHN_IM-2009-12725. (16S: KC703439, 28S: KC703684, COI: JX046599, go to Genbank) ditto.

MNHN_IM-2009-12726. (16S: KC703440, 28S: KC703685, COI: JX046600, go to Genbank) ditto.

MNHN_IM-2009-12758. (16S: KC703446, 28S: KC703691, COI: JX046620, go to Genbank) France: Taviganu River.

MNHN_IM-2009-12759. (16S: KC703447, 28S: KC703692, COI: JX046621, go to Genbank) ditto.

MNHN_IM-2009-12760. (16S: KC703448, 28S: KC703693, COI: JX046622, go to Genbank) ditto.

MNHN_IM-2009-12761. (16S: KC703449, 28S: KC703694, COI: JX046623, go to Genbank) ditto.

MNHN_IM-2009-12666. (16S: KC703393, 28S: KC703647, COI: JX046575, go to Genbank) France: Vonne River.

MNHN_IM-2009-12667. (16S: KC703394, COI: JX046576, go to Genbank) ditto.

MNHN_IM-2009-12668. (16S: KC703395, COI: JX046577, go to Genbank) ditto.

Spain

MNCN_Fw1487-1. (16S: EF635011, COI: AY522849, go to Genbank) Spain: Ebro River, Sastago.

MNCN_Fw1487-2. (16S: EF635012, COI: AY522850, go to Genbank) ditto.