Gangloff, M.M., B.A. Hamstead, E.F. Abernethy & P.D. Hartfield. 2013. Genetic distinctiveness of Ligumia recta, the black sandshell, in the Mobile River Basin and implications for conservation. Conservation Genetics 14: 913-916. Available electronically.
(click the figure number to see the cladogram)
+-- Ligumia_recta
+--+
| +-- Sagittunio_nasutus
--+
| +-- Venustaconcha_ellipsiformis
+--+
+-- Venustaconcha_pleasii