Bivalvia | Unionida | Unionoidea | Unionidae | Parreysiinae | Indochinellini

genus Radiatula Simpson, 1900: cladomics

7 Recent valid species (7 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 7 (7 total)
analyzed in 35 cladograms in 23 publications
maximum number of terminals in an analysis = 7 (mean = 3.57, median = 3)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 27
maximum number of included terminals = 7, (mean = 4.33 median = 3)
proportion of cladograms that support basic monophyly = 0.85

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Radiatula cladogram distances
PCoA Ordination

NJ Tree of Radiatula cladogram distances
NJ Tree

Cladograms

Bolotov et al., 2024 [Publ 4369], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
7 7 5 7 55 22 7 0 
                    +-- Pseudoscabies          
  +-----------------+                          
  |                 +-- Indonaia               
  |                                            
--+  +----------------- Indochinella           
  |  |                                         
  |  |        +-------- Scabiellus             
  |  |  +-----+                                
  +--+  |     |  +----- Harmandia              
     |  |     +--+                             
     |  |        |  +-- Unionetta              
     |  |        +--+                          
     +--+           +-- Scabies                
        |                                      
        |  +----------- Radiatula_crispisulcata **
        |  |                                   
        |  |     +----- Radiatula_chaudhurii **   
        +--+  +--+                             
           |  |  |  +-- Radiatula_myitkyinae **   
           |  |  +--+                          
           +--+     +-- Radiatula_kachinensis **  
              |                                
              |  +----- Radiatula_chindwinensis **
              +--+                             
                 |  +-- Radiatula_myitthanensis **
                 +--+                          
                    +-- Radiatula_mouhoti **

Bolotov et al., 2022 [Publ 4252], fig. 2

iTtTbiriSoTroTroSiD
7 7 5 7 54 20 6 0 
  +-------------------- Indonaia               
  |                                            
  |  +----------------- Indochinella           
--+  |                                         
  |  |        +-------- Scabiellus             
  |  |  +-----+                                
  +--+  |     |  +----- Harmandia              
     |  |     +--+                             
     |  |        |  +-- Unionetta              
     |  |        +--+                          
     +--+           +-- Scabies                
        |                                      
        |  +----------- Radiatula_crispisulcata **
        |  |                                   
        |  |     +----- Radiatula_chaudhurii **   
        +--+  +--+                             
           |  |  |  +-- Radiatula_myitkyinae **   
           |  |  +--+                          
           +--+     +-- Radiatula_kachinensis **  
              |                                
              |  +----- Radiatula_chindwinensis **
              +--+                             
                 |  +-- Radiatula_myitthanensis **
                 +--+                          
                    +-- Radiatula_mouhoti **

Bolotov et al., 2024 [Publ 4369], fig. S1

iTtTbiriSoTroTroSiD
7 7 5 7 56 22 7 6 
                       +-- Pseudoscabies          
  +--------------------+                          
  |                    +-- Indonaia               
  |                                               
--+  +-------------------- Indochinella           
  |  |                                            
  |  |           +-------- Scabiellus             
  |  |  +--------+                                
  +--+  |        |  +----- Harmandia              
     |  |        +--+                             
     |  |           |  +-- Unionetta              
     |  |           +--+                          
     +--+              +-- Scabies                
        |                                         
        |              +-- Radiatula_myitkyinae **   
        |  +-----------+                          
        |  |           +-- Radiatula_kachinensis **  
        +--+                                      
           |  +----------- Radiatula_chaudhurii **   
           +--+                                   
              |  +-------- Radiatula_crispisulcata **
              +--+                                
                 |  +----- Radiatula_mouhoti **      
                 +--+                             
                    |  +-- Radiatula_myitthanensis **
                    +--+                          
                       +-- Radiatula_chindwinensis **

Bolotov et al., 2022 [Publ 4252], fig. S1

iTtTbiriSoTroTroSiD
7 7 5 7 54 20 6 4 
  +-------------------- Indonaia               
  |                                            
  |  +----------------- Indochinella           
--+  |                                         
  |  |        +-------- Scabiellus             
  |  |  +-----+                                
  +--+  |     |  +----- Harmandia              
     |  |     +--+                             
     |  |        |  +-- Unionetta              
     |  |        +--+                          
     +--+           +-- Scabies                
        |                                      
        |        +----- Radiatula_chaudhurii **   
        |  +-----+                             
        |  |     |  +-- Radiatula_kachinensis **  
        |  |     +--+                          
        +--+        +-- Radiatula_myitkyinae **   
           |                                   
           |  +-------- Radiatula_crispisulcata **
           +--+                                
              |  +----- Radiatula_mouhoti **      
              +--+                             
                 |  +-- Radiatula_myitthanensis **
                 +--+                          
                    +-- Radiatula_chindwinensis **

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

iTtTbiriSoTroTroSiD
7 7 5 7 624 21 6 6 
        +----- Radiatula_kachinensis **  
  +-----+                             
  |     |  +-- Radiatula_chaudhurii **   
  |     +--+                          
--+        +-- Radiatula_myitkyinae **   
  |                                   
  |  +-------- Radiatula_crispisulcata **
  +--+                                
     |  +----- Radiatula_mouhoti **      
     +--+                             
        |  +-- Radiatula_chindwinensis **
        +--+                          
           +-- Radiatula_myitthanensis **

Bolotov et al., 2022 [Publ 4239], fig. 2

iTtTbiriSoTroTroSiD
6 6 4 6 52 20 6 3 
  +-------------------- Indonaia               
  |                                            
--+  +----------------- Indochinella           
  |  |                                         
  |  |        +-------- Scabiellus             
  +--+  +-----+                                
     |  |     |  +----- Harmandia              
     |  |     +--+                             
     |  |        |  +-- Unionetta              
     +--+        +--+                          
        |           +-- Scabies                
        |                                      
        |  +----------- Radiatula_crispisulcata **
        |  |                                   
        +--+        +-- Radiatula_chaudhurii **   
           |  +-----+                          
           |  |     +-- Radiatula_myitkyinae **   
           +--+                                
              |  +----- Radiatula_mouhoti **      
              +--+                             
                 |  +-- Radiatula_chindwinensis **
                 +--+                          
                    +-- Radiatula_myitthanensis **

Bolotov et al., 2022 [Publ 4239], fig. S1

iTtTbiriSoTroTroSiD
6 6 3 6 52 20 6 2 
  +----------------- Indonaia               
  |                                         
--+  +-------------- Indochinella           
  |  |                                      
  |  |     +-------- Scabiellus             
  +--+  +--+                                
     |  |  |  +----- Harmandia              
     |  |  +--+                             
     |  |     |  +-- Unionetta              
     +--+     +--+                          
        |        +-- Scabies                
        |                                   
        |  +-------- Radiatula_crispisulcata **
        |  |                                
        +--+     +-- Radiatula_chaudhurii **   
           |  +--+                          
           |  |  +-- Radiatula_myitkyinae **   
           +--+                             
              |  +-- Radiatula_myitthanensis **
              |  |                          
              +--+-- Radiatula_mouhoti **      
                 |                          
                 +-- Radiatula_chindwinensis **

Bolotov et al., 2022 [Publ 4250], fig. 8

iTtTbiriSoTroTroSiD
6 6 4 6 43 3 2 3 
  +----------------- Indonaia               
  |                                         
--+  +-------------- Indochinella           
  |  |                                      
  +--+  +----------- Radiatula_crispisulcata **
     |  |                                   
     +--+        +-- Radiatula_chaudhurii **   
        |  +-----+                          
        |  |     +-- Radiatula_myitkyinae **   
        +--+                                
           |  +----- Radiatula_mouhoti **      
           +--+                             
              |  +-- Radiatula_chindwinensis **
              +--+                          
                 +-- Radiatula_myitthanensis **

Bolotov et al., 2022 [Publ 4250], fig. S2

iTtTbiriSoTroTroSiD
6 6 4 6 9 6 2 1 
  +----------------- Indonaia               
  |                                         
--+  +-------------- Indochinella           
  |  |                                      
  +--+  +----------- Radiatula_crispisulcata **
     |  |                                   
     +--+        +-- Radiatula_chaudhurii **   
        |  +-----+                          
        |  |     +-- Radiatula_myitkyinae **   
        +--+                                
           |  +----- Radiatula_chindwinensis **
           +--+                             
              |  +-- Radiatula_mouhoti **      
              +--+                          
                 +-- Radiatula_myitthanensis **

Bolotov et al., 2022 [Publ 4239], fig. S2

iTtTbiriSoTroTroSiD
6 6 4 6 52 20 6 3 
  +-------------------- Indonaia               
  |                                            
--+  +----------------- Indochinella           
  |  |                                         
  |  |        +-------- Scabiellus             
  +--+  +-----+                                
     |  |     |  +----- Harmandia              
     |  |     +--+                             
     |  |        |  +-- Unionetta              
     +--+        +--+                          
        |           +-- Scabies                
        |                                      
        |  +----------- Radiatula_crispisulcata **
        |  |                                   
        +--+        +-- Radiatula_chaudhurii **   
           |  +-----+                          
           |  |     +-- Radiatula_myitkyinae **   
           +--+                                
              |  +----- Radiatula_mouhoti **      
              +--+                             
                 |  +-- Radiatula_myitthanensis **
                 +--+                          
                    +-- Radiatula_chindwinensis **

Bolotov et al., 2019 [Publ 3974], fig. 1

iTtTbiriSoTroTroSiD
5 5 3 5 13 1 1 2 
  +----------- Indochinella           
  |                                   
--+        +-- Radiatula_myitkyinae **   
  |  +-----+                          
  |  |     +-- Radiatula_chaudhurii **   
  +--+                                
     |  +----- Radiatula_chindwinensis **
     +--+                             
        |  +-- Radiatula_mouhoti **      
        +--+                          
           +-- Radiatula_myitthanensis **

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
4 4 2 4 113 17 6 3 
  +----------------- Indonaia               
  |                                         
--+  +-------------- Indochinella           
  |  |                                      
  |  |     +-------- Radiatula_crispisulcata **
  +--+  +--+                                
     |  |  |  +----- Radiatula_mouhoti **      
     |  |  +--+                             
     |  |     |  +-- Radiatula_chaudhurii **   
     +--+     +--+                          
        |        +-- Radiatula_myitkyinae **   
        |                                   
        |  +-------- Scabiellus             
        +--+                                
           |  +----- Scabies                
           +--+                             
              |  +-- Unionetta              
              +--+                          
                 +-- Harmandia

Bolotov et al., 2020 [Publ 4100], fig. S1

iTtTbiriSoTroTroSiD
4 15 12 4 107 16 6 14 
  +-------------------- Indonaia               
  |                                            
--+  +----------------- Indochinella           
  |  |                                         
  |  |           +----- Radiatula_mouhoti **      
  +--+  +--------+                             
     |  |        |  +-- Radiatula_myitkyinae **   
     |  |        +--+                          
     +--+           +-- Radiatula_chaudhurii **   
        |                                      
        |  +----------- Radiatula_crispisulcata **
        +--+                                   
           |  +-------- Scabiellus             
           +--+                                
              |  +----- Harmandia              
              +--+                             
                 |  +-- Unionetta              
                 +--+                          
                    +-- Scabies

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
3 3 1 3 121 9 3 4 
  +----------- Indochinella        
--+                                
  |  +-------- Scabies             
  +--+                             
     |  +----- Radiatula_mouhoti **   
     +--+                          
        |  +-- Radiatula_myitkyinae **
        +--+                       
           +-- Radiatula_chaudhurii **

Konopleva et al., 2019 [Publ 3900], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 48 9 3 4 
  +-------------- Indonaia            
--+                                   
  |  +----------- Indochinella        
  +--+                                
     |  +-------- Scabies             
     +--+                             
        |  +----- Radiatula_mouhoti **   
        +--+                          
           |  +-- Radiatula_myitkyinae **
           +--+                       
              +-- Radiatula_chaudhurii **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
3 3 1 3 197 9 3 4 
  +-------------- Indonaia            
--+                                   
  |  +----------- Indochinella        
  +--+                                
     |  +-------- Scabies             
     +--+                             
        |  +----- Radiatula_mouhoti **   
        +--+                          
           |  +-- Radiatula_myitkyinae **
           +--+                       
              +-- Radiatula_chaudhurii **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
3 3 1 3 195 9 3 4 
  +-------------- Indonaia            
--+                                   
  |  +----------- Indochinella        
  +--+                                
     |  +-------- Scabies             
     +--+                             
        |  +----- Radiatula_mouhoti **   
        +--+                          
           |  +-- Radiatula_myitkyinae **
           +--+                       
              +-- Radiatula_chaudhurii **

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 79 8 3 6 
  +-------------- Indonaia            
--+                                   
  |  +----------- Indochinella        
  +--+                                
     |  +-------- Scabies             
     +--+                             
        |  +----- Radiatula_myitkyinae **
        +--+                          
           |  +-- Radiatula_mouhoti **   
           +--+                       
              +-- Radiatula_chaudhurii **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
3 4 2 3 88 2 2 7 
  +----------- Indonaia            
--+                                
  |  +-------- Radiatula_myitkyinae **
  +--+                             
     |  +----- Scabies             
     +--+                          
        |  +-- Radiatula_mouhoti **   
        +--+                       
           +-- Radiatula_chaudhurii **

Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 1

iTtTbiriSoTroTroSiD
3 6 1 3 33 5 3 8 
  +-------- Indonaia            
  |                             
--+  +----- Indochinella        
  |  |                          
  +--+  +-- Radiatula_mouhoti **   
     |  |                       
     |  +-- Radiatula_myitkyinae **
     +--+                       
        +-- Radiatula_chaudhurii **
        |                       
        +-- Scabies

Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 2

iTtTbiriSoTroTroSiD
3 8 1 3 33 5 3 10 
  +-- Radiatula_mouhoti **   
  |                       
  +-- Radiatula_myitkyinae **
  |                       
  +-- Radiatula_chaudhurii **
--+                       
  +-- Indochinella        
  |                       
  +-- Indonaia            
  |                       
  +-- Scabies

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 197 9 3 6 
  +-------------- Indonaia            
--+                                   
  |  +----------- Indochinella        
  +--+                                
     |  +-------- Scabies             
     +--+                             
        |  +----- Radiatula_myitkyinae **
        +--+                          
           |  +-- Radiatula_chaudhurii **
           +--+                       
              +-- Radiatula_mouhoti **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
3 3 1 3 88 2 2 6 
  +----------- Indonaia            
--+                                
  |  +-------- Scabies             
  +--+                             
     |  +----- Radiatula_myitkyinae **
     +--+                          
        |  +-- Radiatula_mouhoti **   
        +--+                       
           +-- Radiatula_chaudhurii **

Jeratthitikul et al., 2019 [Publ 3922], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 20 15 6 4 
  +----------------- Indonaia            
  |                                      
--+  +-------------- Indochinella        
  |  |                                   
  |  |        +----- Radiatula_mouhoti **   
  +--+  +-----+                          
     |  |     |  +-- Radiatula_myitkyinae **
     |  |     +--+                       
     +--+        +-- Radiatula_chaudhurii **
        |                                
        |  +-------- Scabiellus          
        +--+                             
           |  +----- Harmandia           
           +--+                          
              |  +-- Unionetta           
              +--+                       
                 +-- Scabies

Pfeiffer et al., 2018 [Publ 3827], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 26 15 6 -- 
  +-------------------- Indonaia             
  |                                          
--+  +----------------- Indochinella         
  |  |                                       
  +--+              +-- Radiatula_chaudhurii ** 
     |  +-----------+                        
     |  |           +-- Radiatula_myitkyinae ** 
     +--+                                    
        |  +----------- Scabiellus           
        +--+                                 
           |  +-------- Unionetta_fabagina__2
           +--+                              
              |  +----- Harmandia            
              +--+                           
                 |  +-- Unionetta_fabagina__1
                 +--+                        
                    +-- Scabies

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
2 2 0 2 136 9 3 -- 
  +-------------- Indonaia            
  |                                   
  |           +-- Indochinella        
--+  +--------+                       
  |  |        +-- Scabies_crispata    
  |  |                                
  +--+        +-- Radiatula_myitkyinae **
     |  +-----+                       
     |  |     +-- Radiatula_chaudhurii **
     +--+                             
        |  +----- Scabies_mandarinus  
        +--+                          
           |  +-- Scabies_humilis     
           +--+                       
              +-- Scabies_phaselus

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
2 2 0 2 136 9 3 -- 
  +-------------- Indonaia            
  |                                   
  |           +-- Indochinella        
--+  +--------+                       
  |  |        +-- Scabies_crispata    
  |  |                                
  +--+        +-- Radiatula_myitkyinae **
     |  +-----+                       
     |  |     +-- Radiatula_chaudhurii **
     +--+                             
        |  +----- Scabies_mandarinus  
        +--+                          
           |  +-- Scabies_humilis     
           +--+                       
              +-- Scabies_phaselus

Konopleva et al., 2023 [Publ 4262], fig. 8

iTtTbiriSoTroTroSiD
1 1 0 1 81 13 6 -- 
  +----------------- Indonaia         
--+                                   
  |  +-------------- Indochinella     
  +--+                                
     |  +----------- Radiatula_mouhoti **
     +--+                             
        |  +-------- Scabiellus       
        +--+                          
           |  +----- Scabies          
           +--+                       
              |  +-- Unionetta        
              +--+                    
                 +-- Harmandia

Choi et al., 2020 [Publ 4169], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 9 0 0 -- 
-- Radiatula_mouhoti **

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
1 1 0 1 35 1 1 -- 
  +-- Radiatula_chaudhurii **
--+                       
  +-- Indochinella

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 39 2 2 -- 
  +----- Indochinella        
--+                          
  |  +-- Radiatula_chaudhurii **
  +--+                       
     +-- Scabies

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
1 1 0 1 121 0 0 -- 
-- Radiatula_mouhoti **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
1 1 0 1 55 2 2 -- 
  +----- Indonaia         
--+                       
  |  +-- Indochinella     
  +--+                    
     +-- Radiatula_mouhoti **

Konopleva et al., 2017 [Publ 3807], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 30 2 2 -- 
  +----- Indochinella        
--+                          
  |  +-- Radiatula_chaudhurii **
  +--+                       
     +-- Scabies

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
1 1 0 1 35 1 1 -- 
  +-- Radiatula_chaudhurii **
--+                       
  +-- Indochinella