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  +-------------------------------------- Neotrigonia_margaritacea   
  |                                                                  
  |                             +-------- Velesunio_ambiguus         
  |  +--------------------------+                                    
  |  |                          |  +----- Hyridella_australis        
--+  |                          +--+                                 
  |  |                             |  +-- Triplodon_corrugatus       
  |  |                             +--+                              
  |  |                                +-- Castalia_ambigua           
  |  |                                                               
  +--+                             +----- Etheria_elliptica          
     |  +--------------------------+                                 
     |  |                          |  +-- Lamproscapha_ensiformis    
     |  |                          +--+                              
     |  |                             +-- Chambardia_wahlbergi       
     |  |                                                            
     |  |                             +-- Margaritifera_margaritifera
     |  |  +--------------------------+                              
     +--+  |                          +-- Cumberlandia_monodonta     
        |  |                                                         
        |  |                          +-- Trapezidens_dolichorhynchus
        |  |              +-----------+                              
        |  |              |           +-- Lamellidens_savadiensis    
        |  |              |                                          
        |  |  +-----------+           +-- Leoparreysia_olivacea      
        +--+  |           |  +--------+                              
           |  |           |  |        +-- Leoparreysia_tavoyensis    
           |  |           +--+                                       
           |  |              |  +-------- Coelatura_aegyptiaca       
           |  |              +--+                                    
           |  |                 |  +----- Indochinella_pugio         
           |  |                 +--+                                 
           |  |                    |  +-- Radiatula_chaudhurii       
           |  |                    +--+                              
           +--+                       +-- Scabies_mandarinus         
              |                                                      
              |                    +----- Unio_pictorum              
              |  +-----------------+                                 
              |  |                 |  +-- Pyganodon_grandis          
              |  |                 +--+                              
              |  |                    +-- Lasmigona_costata          
              |  |                                                   
              |  |           +----------- Quadrula_quadrula          
              |  |  +--------+                                       
              +--+  |        |  +-------- Amblema_plicata            
                 |  |        +--+                                    
                 |  |           |  +----- Potamilus_alatus           
                 |  |           +--+                                 
                 |  |              |  +-- Lampsilis_cardium          
                 |  |              +--+                              
                 |  |                 +-- Truncilla_truncata         
                 +--+                                                
                    |           +-------- Ensidens_telus             
                    |  +--------+                                    
                    |  |        |  +----- Lens_micropterus           
                    |  |        +--+                                 
                    |  |           |  +-- Physunio                   
                    |  |           +--+                              
                    |  |              +-- Physunio_modelli           
                    +--+                                             
                       |        +-------- Gonidea_angulata           
                       |  +-----+                                    
                       |  |     |  +----- Potomida_littoralis        
                       |  |     +--+                                 
                       |  |        |  +-- Pronodularia_japanensis    
                       +--+        +--+                              
                          |           +-- Lamprotula_caveata         
                          |                                          
                          |  +----------- Pilsbryoconcha_carinifera  
                          |  |                                       
                          +--+        +-- Pseudodon_vondembuschianus 
                             |  +-----+                              
                             |  |     +-- Pseudodon_cambodjensis     
                             +--+                                    
                                |  +----- Sundadontina_tumida        
                                +--+                                 
                                   |  +-- Nyeinchanconcha_nyeinchani 
                                   +--+                              
                                      +-- Bineurus_exilis