Lane, T.W., E.M. Hallerman & J.W. Jones. 2016. Phylogenetic and taxonomic assessment of the endangered Cumberland bean, Villosa trabalis and purple bean, Villosa perpurpurea (Bivalvia: Unionidae). Conservation Genetics 17(5): 1109-1124. Available electronically.
(click the figure number to see the cladogram)
+----- Paetulunio_fabalis
|
+----- Epioblasma_capsaeformis
--+
| +-- Venustaconcha_trabalis__1
| |
| +-- Venustaconcha_trabalis__2
+--+
+-- Venustaconcha_ellipsiformis
|
+-- Venustaconcha_pleasii