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Data last updated
Mon 1 Dec 2025
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Publication Lopes-Lima et al. (2017): cladogramspublication summary | cladograms
Complete CitationLopes-Lima, M., E. Froufe, V.T. Do, M. Ghamizi, K.E. Mock et al. 2017. Phylogeny of the most species-rich freshwater bivalve family (Bivalvia: Unionida: Unionidae): Defining modern subfamilies and tribes. Molecular Phylogenetics and Evolution 106: 174-191. Available electronically.
Cladogram Figures(click the figure number to see the cladogram) Cladogram Figure 1-2
+----------------------------------------- Neotrigonia_margaritacea
|
| +-- Hyridella_depressa
| +-----------------------------------+
| | +-- Velesunio_ambiguus
--+ |
| | +-- Cumberlandia_monodonta
| | +--------------------------------+
| | | +-- Margaritifera_margaritifera
| | |
| | | +----------- Mutela_rostrata
+--+ | |
| | +--------------------+ +-- Aspatharia_pfeifferiana
| | | | +-----+
| | | | | +-- Chambardia_wahlbergi
| | | +--+
| | | | +----- Etheria_elliptica
| | | +--+
| | | | +-- Tamsiella_schroeteriana
+--+ | +--+
| | +-- Mycetopoda_siliquosa
| |
| | +-------- Trapezidens_dolichorhynchus
| | +--------------------+
| | | | +----- Leoparreysia_tavoyensis
| | | +--+
| | | | +-- Coelatura_aegyptiaca
| | | +--+
| | | +-- Indochinella_pugio
| | |
| | | +----- Unio_gibbus
+--+ | |
| | +-----------+----- Unio_tumidus
| | | |
| | | | +-- Unio_crassus
| | | +--+
| | +--+ +-- Unio_pictorum
| | | |
| | | | +-------------- Aculamprotula_tortuosa
| | | | |
| | | +--+ +-- Nodularia_nuxpersicae
| | | | +--------+
| | | | | +-- Nodularia_douglasiae
| | | +--+
| | | | +-------- Schistodesmus_xinyuensis
| | | +--+
| | | | +----- Tchangsinaia_piscicula
| | | +--+
| | +-----+ | +-- Cuneopsis_szechenyii
| | | | +--+
| | | | +-- Cuneopsis_rufescens
+--+ | |
| | | +-------- Lanceolaria_gladiola
| | | +--------+
| | | | | +----- Lanceolaria_grayii__2
| | | | +--+
| | | | | +-- Lanceolaria_lanceolata
| | | | +--+
| | | | +-- Lanceolaria_grayii__1
| | | |
| | +--+ +----------- Cristaria_plicata
| | | +--+
| | | | | +-------- Anemina_arcaeformis
| | | | +--+
| | | | | +----- Pletholophus_tenuis
| | | | +--+
| | | | | +-- Sinanodonta_jourdyi
| | | | +--+
| | +--+ +-- Sinanodonta_lucida
| | |
| | | +----- Pseudanodonta_complanata
| | | |
| | | +-----+ +-- Anodonta_anatina
| | | | | |
| | | | +--+-- Anodonta_cygnea
| | | | |
| | +--+ +-- Anodonta_nuttalliana
| | |
| | | +-- Pyganodon_grandis
+--+ | +-----+
| | | +-- Simpsonaias_ambigua
| +--+
| | +----- Strophitus_undulatus
| +--+
| | +-- Alasmidonta_marginata
| +--+
| +-- Lasmigona_costata
|
| +-- Lens_contradens
| +--+
| | +-- Lens_comptus
| +--+
| | | +-- Physunio_modelli
| | +--+
| | +-- unionoid_species
| |
| +--------------+ +----- Hyriopsis_desowitzi
| | | |
| | | +----- Rectidens_sumatrensis
| | | |
| | | | +-- Hyriopsis_khoratensis
| | +--+--+
| | | +-- Hyriopsis_myersiana
| | |
| | | +-- Ensidens_ingallsianus
| | | |
| | +--+-- Ensidens_telus
| | |
| | +-- Ensidens_jaculus
| |
| | +-------- Amblema_plicata
+--+ +--+
| | | +----- Cambarunio_iris
| | +--+
| | | +-- Lampsilis_cardium
| | +--+
| +--------+ +-- Ortmanniana_ligamentina
| | |
| | | +-- Quadrula_quadrula
| | | +-----+
| | | | +-- Quadrula_verrucosa
| | +--+
| | | +----- Elliptio_complanata
| | +--+
| | | +-- Pleurobema_sintoxia
| | +--+
| | +-- Eurynia_dilatata
+--+
| +-- Chamberlainia_somsakpanhai
| +-----+
| | +-- Sinohyriopsis_cumingii
| |
| +--------+ +-- Lamprotula_caveata
| | | +--+
| | | | +-- Lamprotula_leaii
| | +--+
| | | +-- Potomida_littoralis
| | +--+
| | +-- Pronodularia_japanensis
+--+
| +-- Sinosolenaia_carinata
| +-----+
| | +-- Sinosolenaia_oleivora
| +-----+
| | | +----- Gonidea_angulata
| | +--+
| | | +-- Microcondylaea_bonellii
+--+ +--+
| +-- Leguminaia_wheatleyi
|
| +----------- Parvasolenaia_rivularis
| |
+--+ +-- Pilsbryoconcha_exilis
| +-----+
| | +-- Pilsbryoconcha_acuta
+--+
| +----- Pseudodon_cambodjensis
+--+
| +-- Bineurus_exilis
+--+
+-- Nyeinchanconcha_nyeinchani
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