species Diplodon charruanus (d’Orbigny, 1835): synonymy

Unio charruana d’Orbigny, 1835

d’Orbigny, 1835. Mag. Zool. 5(61-62): 35.
d’Orbigny, 1846. Voyage dans l’Amérique Méridionale 5(3): 606, pl. 71, figs. 8-14.
LECTOTYPE BMNH_1854-9-4-15 (by R.I. Johnson, 1971: 80), Rio del Rosario, Banda orientali, Uruguay.

Diplodon (s.s.) charruanus — Simpson (1900), Simpson (1914), Ortmann (1921), Morretes (1949), Haas (1969)
Unio charruana — Corsi (1900)
Diplodon charruanus — Haas (1930), Bonetto (1954), Bonetto (1959), Parodiz (1968), Mansur (1970), Graf & Cummings (2021)
Diplodon (Rhipidodonta) charruanus — Bonetto (1964), Bonetto (1965), Figueiras (1965), Bonetto & Mansur (1970), Mansur & Pereira (2006), Pereira et al. (2014), Torres et al. (2018)
Diplodon charruana — Schade (1965)
Rhipidodonta charruana — Simone (2006), Graf & Cummings (2007), Bogan (2010), Mansur et al. (2020), Bieler (2021), Torres et al. (2024), Nagel (2025)
Diplodon (Rhipidiodonta) [sic] charruanus — Rumi et al. (2008)

Unio faba d’Orbigny, 1835

d’Orbigny, 1835. Mag. Zool. 5(61-62): 35.
d’Orbigny, 1846. Voyage dans l’Amérique Méridionale 5(3): 606.
SYNTYPES BMNH_54-12-4-839, Banda orientali, Uruguay.

Diplodon (s.s.) charruanus — Simpson (1900), Simpson (1914), Ortmann (1921), Haas (1969)
Unio charruana — Corsi (1900)
Diplodon charruanus — Haas (1930), Bonetto (1954), Parodiz (1968), Mansur (1970)
Diplodon (Rhipidodonta) charruanus — Bonetto (1964), Bonetto (1965), Figueiras (1965), Bonetto & Mansur (1970)
Rhipidodonta charruana — Simone (2006)

Unio charruana d’Orbigny, 1851

(reference to Unio charruana d’Orbigny, 1835)
d’Orbigny, 1851. Rev. Mag. Zool. 3: ??.

Diplodon charruanus — Bonetto (1959)

Unio lepidus Lea, 1860

(renamed Margaron (Unio) lepidior Lea, 1870; homonym of Unio lepidus Gould, 1856)
Lea, 1860. Proc. Acad. Nat. Sci. 12: 91.
Lea, 1863. Jour. Acad. Nat. Sci. 5 [O. 10]: 389, pl. 50, fig. 306.
LECTOTYPE USNM_85624 (by Johnson & Baker, 1973: 160), Uruguay R., S. America.

Diplodon (s.s.) lepidior — Simpson (1900), Simpson (1914)
Diplodon rhuacoicus — Haas (1930)
Diplodon (Rhipidodonta) charruanus — Bonetto (1964), Bonetto (1965), Figueiras (1965)
Diplodon charruanus — Parodiz (1968), Mansur (1970)
Diplodon (s.s.) rhuacoicus — Haas (1969)
Rhipidodonta charruana — Simone (2006), Bieler (2021)

Unio parcus Lea, 1866

Lea, 1866. Proc. Acad. Nat. Sci. 18: 34.
Lea, 1868. Jour. Acad. Nat. Sci. 6 [O. 12]: 262, pl. 33, fig. 77.
HOLOTYPE USNM_85198, S. America.

Diplodon (s.s.) parcus — Simpson (1900), Simpson (1914)
Diplodon (s.s.) charruanus — Ortmann (1921), Haas (1969)
Diplodon charruanus — Haas (1930), Parodiz (1968), Mansur (1970)
Diplodon (s.s.) rhuacoicus — Bonetto (1964), Bonetto (1965), Figueiras (1965)
Rhipidodonta charruana — Simone (2006)

Margaron (Unio) lepidior Lea, 1870

(new name for Unio lepidus Lea, 1860)
Lea, 1870. Synopsis Fam. Naiades: 53.
OBJECTIVE SYNONYM USNM_85624, Uruguay R., S. America.

Diplodon (s.s.) lepidior — Simpson (1900), Simpson (1914)
Diplodon rhuacoicus — Haas (1930)
Diplodon (Rhipidodonta) charruanus — Bonetto (1964), Bonetto (1965), Figueiras (1965)
Diplodon charruanus — Parodiz (1968), Mansur (1970)
Diplodon (s.s.) rhuacoicus — Haas (1969)
Diplodon (s.s.) delodontus wimanii [sic] — Bonetto & Mansur (1970)
Rhipidodonta charruana — Simone (2006), Bieler (2021)

Unio aethiops var. piracicabana Ihering, 1893

(intro'd as intra-sp. of Unio aethiops Lea, 1860)
Ihering, 1893. Arch. Naturg. 59(1): 102. [in part]
LECTOTYPE SMF_4031 (by Haas, 1930: 193), Piracicaba, Geb. d Rio Tiete, St. Sao Paulo, Bras.

Diplodon (s.s.) æthiops var. piricicabana [sic] — Simpson (1900)
Diplodon (s.s.) aethiops var. piracicabanus — Simpson (1914)
Diplodon delodontus expansus — Haas (1930)
Diplodon (s.s.) delodontus expansus — Bonetto (1964), Bonetto (1965), Haas (1969)
Diplodon expansus — Parodiz (1968), Mansur (1970), Quintana (1982), Simone (2006)
Diplodon (s.s.) aethiops — Mansur & Pereira (2006)
Rhipidodonta charruana — Nagel (2025)

Unio aethiopiformis Simpson, 1900

‘von Ihering’ Simpson, 1900. Proc. U.S. Nat. Mus. 22: 894. [nomen nudum]
‘von Ihering’ Simpson, 1914. Descr. Cat. Naiades: 1312. [nomen nudum]
NOMEN NUDUM MCZ_192369, Rio Camaquam, Rio Grande do Sul, Brazil.

Diplodon aethiopiformis — Simpson (1914)
Diplodon charruanus — Haas (1930)
Diplodon (Rhipidodonta) charruanus — Bonetto (1964), Bonetto & Mansur (1970)
Diplodon parallelipipedon [sic] aethiops — Parodiz (1968), Mansur (1970)
Rhipidodonta charruana — Simone (2006)

Diplodon charruanus lujanensis Ihering, 1907

(intro'd as intra-sp. of Unio charruana d’Orbigny, 1835)
Ihering, 1907. Ann. Mus. Nac. Buenos Aires 14: 468.
NOMEN NUDUM SMF_3918, Lujan bei Buenous Aires Arg. ob Pampas Stufe.

Rhipidodonta charruana — Simone (2006)

Diplodon (s.s.) trivialis Simpson, 1914

Simpson, 1914. Descr. Cat. Naiades: 1250.
SYNTYPE USNM_162409, Taboticabal [= Jaboticabal], Sao Paulo, Brazil.

Diplodon (s.s.) trivialis — Simpson (1914), Morretes (1949)
Diplodon trivialis — Marshall (1917), Bonetto (1959)
Diplodon charruanus — Haas (1930), Bonetto (1954)
Diplodon uruguayensis X expansus — Parodiz (1968)
Diplodon (s.s.) charruanus — Haas (1969)
Diplodon uruguayensis x expansus — Mansur (1970)
Rhipidodonta charruana — Simone (2006)

Diplodon (s.s.) perplexus Simpson, 1914

Simpson, 1914. Descr. Cat. Naiades: 1248.
SYNTYPE USNM_150389, Maldonado, Uruguay.

Diplodon (s.s.) perplexus — Simpson (1914)
Diplodon perplexus — Marshall (1917)
Diplodon charruanus — Haas (1930), Bonetto (1954), Parodiz (1968), Mansur (1970)
Diplodon (Rhipidodonta) charruanus — Bonetto (1964), Bonetto (1965), Figueiras (1965), Bonetto & Mansur (1970)
Diplodon (s.s.) charruanus — Haas (1969)
Rhipidodonta charruana — Simone (2006)

Diplodon hidalgoi Haas, 1916

Haas, 1916. Trab. Mus. Nacion. Cienc. Nat., Madrid 25: 18, pl. 1, fig. 1.
HOLOTYPE MNCN_15.07/2, Rio Miguelete, Uruguay.

Diplodon (s.s.) charruanus — Ortmann (1921), Haas (1969)
Diplodon charruanus — Haas (1930), Bonetto (1954)
Diplodon (Rhipidodonta) charruanus — Bonetto (1964), Bonetto (1965), Figueiras (1965), Bonetto & Mansur (1970)
Diplodon piceus — Parodiz (1968), Mansur (1970)
Rhipidodonta charruana — Simone (2006), Nagel (2025)

Diplodon fortis Marshall, 1917

Marshall, 1917. Proc. U.S. Nat. Mus. 53(2209): 382, pl. 52, figs. 1-4.
HOLOTYPE (ORIG. DESIG.) USNM_270899, Rio Negro, Tacuarembo, Uruguay.

Diplodon fortis — Marshall (1917)
Diplodon (s.s.) charruanus — Ortmann (1921), Haas (1969)
Diplodon charruanus — Haas (1930), Bonetto (1954)
Diplodon (Rhipidodonta) charruanus — Bonetto (1964), Bonetto (1965), Figueiras (1965), Bonetto & Mansur (1970)
Diplodon piceus — Parodiz (1968), Mansur (1970)
Rhipidodonta charruana — Simone (2006)

Diplodon trivialoides Marshall, 1922

Marshall, 1922. Proc. U.S. Nat. Mus. 61(2437): 5, pl. 1, figs. 12, 14; pl. 2, figs. 3, 5; pl. 3, fig. 12.
HOLOTYPE (ORIG. DESIG.) USNM_335770, Arroyo Bellaco, Dept. Paysandu, Uruguay.

Diplodon trivialoides — Marshall (1922)
Diplodon charruanus — Haas (1930), Bonetto (1954), Parodiz (1968), Mansur (1970)
Diplodon (Rhipidodonta) charruanus — Bonetto (1964), Bonetto (1965), Figueiras (1965), Bonetto & Mansur (1970)
Diplodon (s.s.) charruanus — Haas (1969)
Rhipidodonta charruana — Simone (2006)

Unio lepidioir ‘Lea’ of Authors

(error for Margaron (Unio) lepidior Lea, 1870)
‘Lea’ of Authors.
Bonetto, 1964. Physis 24(68): 327.
Bonetto, 1965. Anais Congr. Lat.-Amer. Zool. 2: 50.
Bonetto & Mansur, 1970. Acta Zool. Lilloana 27: 249.
Simone, 2006. Land. Freshw. Moll. Brazil.: 267.

Unio sebastanae Simpson, 1900

‘von Ihering’ Simpson, 1900. Proc. U.S. Nat. Mus. 22: 894. [nomen nudum]
‘von Ihering’ Simpson, 1914. Descr. Cat. Naiades: 1312. [nomen nudum]

Diplodon sebastanae — Simpson (1914)

Rhipidodonta pacus Simone, 2006

(error for Unio parcus Lea, 1866)
‘Lea’ Simone, 2006. Land. Freshw. Moll. Brazil.: 267.