Sano, I., A. Shirai, T. Kondo & J.-I. Miyazaki. 2017. Phylogenetic relationships of Japanese Unionoida (Mollusca: Bivalvia) based on mitochondrial 16S rDNA sequences. Journal of Water Resources and Protection 9: 493-509. Available electronically.
(click the figure number to see the cladogram)
+-- Neotrigonia_margaritacea
+-----------------------------+
| +-- Neotrigonia_lamarckii
|
| +-------- Margaritifera_laevis__2
| +--------------------+
| | | +----- Margaritifera_laevis__1
--+ | +--+
| | | +-- Margaritifera_middendorffi
| | +--+
| | +-- Margaritifera_laevis__3
| |
| | +-------- Obovalis_omiensis
+--+ +-----------------+
| | | +----- Inversidens_brandtii
| | +--+
| | | +-- Pronodularia_japanensis
| | +--+
| | +-- Sinohyriopsis_schlegelii
| |
| | +-------------------- Lanceolaria_grayii
+--+ +--+
| | | +----------------- Nodularia_nipponensis
| | +--+
| | | +-------------- Inversiunio_jokohamensis
| | +--+
| | | +----------- Inversiunio_yanagawensis__1
| | +--+
| | | +-------- Inversiunio_yanagawensis__2
| | +--+
+--+ | +----- Inversiunio_yanagawensis__3
| +--+
| | +-- Inversiunio_yanagawensis__4
| +--+
| +-- Inversiunio_reinianus
|
| +-- Cristaria_plicata
| +-----------+
| | +-- Buldowskia_iwakawai
+--------+
| +----------- Pletholophus_reinianus
| |
+--+ +-- Sinanodonta_tumens
| +-----+
| | +-- Sinanodonta_lauta
+--+
| +----- Beringiana_fukuharai
+--+
| +-- Beringiana_japonica
+--+
+-- Beringiana_beringiana
+-- Neotrigonia_lamarckii
+--------------------------+
| +-- Neotrigonia_margaritacea
|
| +-- Margaritifera_laevis
| +-----------------------+
| | +-- Margaritifera_middendorffi
| |
--+ | +-- Obovalis_omiensis
| | +--------+
| | | +-- Ptychorhynchus_pfisteri
| | |
| | +-----------+ +-- Inversidens_brandtii
| | | | +-----+
| | | | | +-- Sinosolenaia_oleivora
+--+ | | |
| | +--+ +-- Pronodularia_japanensis
| | | +--+
| | | | +-- Lamprotula_leaii
| | +--+
| | | +-- Sinohyriopsis_schlegelii
| | +--+
| | +-- Sinohyriopsis_cumingii
| |
| | +-- Cristaria_plicata
+--+ +-----------+
| | +-- Buldowskia_iwakawai
| |
| +-----+ +----- Beringiana_fukuharai
| | | +-----+
| | | | | +-- Beringiana_japonica
| | | | +--+
| | +--+ +-- Beringiana_beringiana
| | |
| | | +-------- Pletholophus_reinianus
| | +--+
| | | +----- Sinanodonta_woodiana
| | +--+
| | | +-- Sinanodonta_lauta
+--+ +--+
| +-- Sinanodonta_tumens
|
| +----------- Tchangsinaia_piscicula
| +-----+
| | | +-------- Nodularia_nipponensis__1
| | +--+
| | | +----- Nodularia_breviconcha
| | +--+
| | | +-- Nodularia_nipponensis__2
| | +--+
+--+ +-- Nodularia_douglasiae
|
| +-- Schistodesmus_xinyuensis
| +-----------+
| | +-- Acuticosta_ovata
| |
| | +-------- Lepidodesma_languilati
+--+ +--+
| | | +----- Lanceolaria_grayii__1
| | +--+
| | | +-- Lanceolaria_lanceolata
+--+ +--+
| +-- Lanceolaria_grayii__2
|
| +-------- Inversiunio_jokohamensis
| |
+--+ +----- Inversiunio_yanagawensis__3
| |
+--+ +-- Inversiunio_yanagawensis__1
| |
+--+-- Inversiunio_yanagawensis__2
|
+-- Inversiunio_reinianus