Lopes-Lima, M., I.N. Bolotov, V.T. Do, D.C. Aldridge, M.M. Fonseca et al. 2018. Expansion and systematics redefinition of the most threatened freshwater mussel family, the Margaritiferidae. Molecular Phylogenetics and Evolution 127: 98-118. Available electronically.
(click the figure number to see the cladogram)
+-- Neotrigonia_margaritacea
+--------------------------+
| +-- Neotrigonia_lamarckii
|
| +----- Triplodon_corrugatus
| +--------------------+
--+ | | +-- Velesunio_ambiguus
| | +--+
| | +-- Hyridella_australis
| |
| | +----------- Mutela_hargeri
| | |
+--+ +-----------+ +-- Aspatharia_pfeifferiana
| | | +-----+
| | | | +-- Chambardia_wahlbergi
| | +--+
| | | +----- Etheria_elliptica
| | +--+
| | | +-- Lamproscapha_ensiformis
| | +--+
+--+ +-- Tamsiella_schroeteriana
|
| +----- Potomida_littoralis
| +--------------+
| | | +-- Lampsilis_cardium
| | +--+
| | +-- Unio_pictorum
| |
+--+ +----- Gibbosula_rochechouartii
| +-----------+
| | | +-- Gibbosula_crassa
| | +--+
| | +-- Gibbosula_laosensis
| |
+--+ +-------- Cumberlandia_monodonta
| +-----+
| | | +----- Pseudunio_marocanus
| | +--+
| | | +-- Pseudunio_auricularius
| | +--+
+--+ +-- Pseudunio_homsensis
|
| +-- Margaritifera_dahurica
| +--------+
| | +-- Margaritifera_margaritifera
+--+
| +-------- Margaritifera_falcata
| |
+--+ +-- Margaritifera_middendorffi
| +--+
| | +-- Margaritifera_laevis
+--+
| +-- Margaritifera_hembeli
+--+
+-- Margaritifera_marrianae
+-- Neotrigonia_margaritacea
+-----------------------------+
| +-- Neotrigonia_lamarckii
|
| +----- Triplodon_corrugatus
| +-----------------------+
--+ | | +-- Velesunio_ambiguus
| | +--+
| | +-- Hyridella_australis
| |
| | +----------- Mutela_hargeri
| | |
+--+ +--------------+ +-- Chambardia_wahlbergi
| | | +-----+
| | | | +-- Aspatharia_pfeifferiana
| | +--+
| | | +----- Etheria_elliptica
| | +--+
| | | +-- Tamsiella_schroeteriana
| | +--+
+--+ +-- Lamproscapha_ensiformis
|
| +----- Potomida_littoralis
| +-----------------+
| | | +-- Lampsilis_cardium
| | +--+
| | +-- Unio_pictorum
| |
+--+ +----- Gibbosula_rochechouartii
| +--------------+
| | | +-- Gibbosula_laosensis
| | +--+
| | +-- Gibbosula_crassa
| |
+--+ +-------- Cumberlandia_monodonta
| +--------+
| | | +----- Pseudunio_marocanus
| | +--+
| | | +-- Pseudunio_auricularius
| | +--+
+--+ +-- Pseudunio_homsensis
|
| +-- Margaritifera_margaritifera
| +-----------+
| | +-- Margaritifera_dahurica
+--+
| +----------- Margaritifera_falcata
+--+
| +-------- Margaritifera_laevis
+--+
| +----- Margaritifera_middendorffi
+--+
| +-- Margaritifera_hembeli
+--+
+-- Margaritifera_marrianae