Johnson, N.A., C.H. Smith, J.M. Pfeiffer, C.R. Randklev, J.D. Williams & J.D. Austin. 2018. Integrative taxonomy resolves taxonomic uncertainty for freshwater mussels being considered for protection under the U.S. Endangered Species Act. Scientific Reports 8. 15892 (16 pp.). Available electronically.
(click the figure number to see the cladogram)
+-------------------------------- Amblema_plicata
|
--+ +----------------------------- Elliptio_crassidens
| |
+--+ +-------------------------- Megalonaias_nervosa
| |
+--+ +----------------------- Uniomerus_tetralasmus
| |
+--+ +-------------------- Theliderma_metanevra
| |
| | +-- Quadrula_quadrula
+--+ +--------------+
| | +-- Quadrula_verrucosa
| |
| | +----- Pustulosa_kieneriana
+--+ +--------+
| | | +-- Pustulosa_infucata
| | +--+
| | +-- Pustulosa_kleiniana
+--+
| +-- Pustulosa_pustulosa
| +--------+
| | +-- Pustulosa_succissa
+--+
| +-------- Cyclonaias_tuberculata
+--+
| +----- Pustulosa_necki
+--+
| +-- Pustulosa_nodulata
+--+
+-- Pustulosa_petrina