Lopes-Lima, M., L. Burlakova, A. Karatayev, A. Gomes-dos-Santos, A. Zieritz et al. 2019. Revisiting the North American freshwater mussel genus Quadrula sensu lato (Bivalvia: Unionidae): Phylogeny, taxonomy and species delineation. Zoologica Scripta 48(3): 313-336. Available electronically.
(click the figure number to see the cladogram)
+-- Margaritifera_falcata
+--------------------------------------+
| +-- Cumberlandia_monodonta
|
| +-- Anodonta_nuttalliana
--+ +-----------------------------------+
| | +-- Pyganodon_grandis
| |
| | +-- Amblema_elliottii
+--+ +--------------------------------+
| | +-- Amblema_plicata
| |
| | +-- Lampsilis_ornata
+--+ +-----------------------------+
| | +-- Potamilus_leptodon
| |
| | +-- Pleurobema_riddellii
+--+ +--------------------------+
| | +-- Elliptio_crassidens
| |
| | +----- Megalonaias_nervosa
+--+ |
| +--------------------+ +-- Uniomerus_carolinianus
| | | |
| | +--+-- Uniomerus_declivis
| | |
+--+ +-- Uniomerus_tetralasmus
|
| +----------------------- Quadrula_quadrula
| |
| | +-- Quadrula_verrucosa
+--+ +-----------------+
| | +-- Quadrula_nobilis
| |
+--+ +-- Theliderma_metanevra
| +--------------+
| | +-- Theliderma_johnsoni
| |
+--+ +----- Pustulosa_kieneriana
| +--------+
| | | +-- Pustulosa_infucata
| | +--+
+--+ +-- Pustulosa_kleiniana
|
| +----------- Cyclonaias_tuberculata
| |
+--+ +-- Pustulosa_pustulosa
| +-----+
| | +-- Pustulosa_succissa
+--+
| +----- Pustulosa_nodulata
+--+
| +-- Pustulosa_necki
+--+
+-- Pustulosa_petrina
+-- Cumberlandia_monodonta
+-----------------------------------+
| +-- Margaritifera_falcata
|
| +-- Anodonta_nuttalliana
--+ +--------------------------------+
| | +-- Pyganodon_grandis
| |
| | +-------------------------------- Megalonaias_nervosa
+--+ |
| | +-- Pleurobema_oviforme
| | +-----+
| | | +-- Pleurobema_riddellii
| | |
+--+ +--------------------+ +-- Lampsilis_ornata
| | | +--+
| | | | +-- Potamilus_leptodon
| | +--+
| | | +-- Amblema_elliottii
| | +--+
+--+ +-- Amblema_plicata
|
| +-------------------------- Uniomerus_tetralasmus
| |
| | +----------- Theliderma_cylindrica
| | +-----------+
| | | | +-------- Theliderma_sparsa
+--+ | +--+
| | | +----- Theliderma_intermedia
| | +--+
| | | +-- Theliderma_metanevra
| | +--+
+--+ +-- Theliderma_johnsoni
|
| +----- Quadrula_quadrula
| +--------------+
| | | +-- Quadrula_nobilis
| | +--+
| | +-- Quadrula_verrucosa
+--+
| +----------------- Pustulosa_kieneriana
| |
| | +-- Pustulosa_infucata
+--+ +-----------+
| | +-- Pustulosa_kleiniana
| |
+--+ +-- Pustulosa_pustulosa
| +--------+
| | +-- Pustulosa_succissa
+--+
| +-------- Cyclonaias_tuberculata
+--+
| +----- Pustulosa_necki
+--+
| +-- Pustulosa_petrina
+--+
+-- Pustulosa_nodulata
+-- Margaritifera_falcata
+-----------------------------------+
| +-- Cumberlandia_monodonta
|
| +-- Anodonta_nuttalliana
| +--------------------------------+
--+ | +-- Pyganodon_grandis
| |
| | +-- Amblema_elliottii
| | +-----+
| | | +-- Amblema_plicata
+--+ |
| +-----------------------+ +-- Pleurobema_riddellii
| | | +--+
| | | | +-- Pleurobema_oviforme
| | +--+
| | | +-- Lampsilis_ornata
+--+ +--+
| +-- Potamilus_leptodon
|
| +----------------------------- Megalonaias_nervosa
| |
+--+ +-------------------------- Uniomerus_tetralasmus
| |
| | +-- Theliderma_metanevra
+--+ +--------------------+
| | +-- Theliderma_johnsoni
| |
+--+ +-------------------- Quadrula_quadrula
| |
| | +-- Quadrula_verrucosa
+--+ +--------------+
| | +-- Quadrula_nobilis
| |
| | +----- Pustulosa_kieneriana
+--+ +--------+
| | | +-- Pustulosa_infucata
| | +--+
| | +-- Pustulosa_kleiniana
+--+
| +-- Pustulosa_pustulosa
| +--------+
| | +-- Pustulosa_succissa
+--+
| +-------- Cyclonaias_tuberculata
+--+
| +----- Pustulosa_necki
+--+
| +-- Pustulosa_petrina
+--+
+-- Pustulosa_nodulata