Jeratthitikul, E., C. Sucharit & P. Prasankok. 2019. Molecular phylogeny of the Indochinese freshwater mussel genus Scabies Haas, 1911 (Bivalvia: Unionidae). Tropical Natural History 19(1): 21-36. Available electronically.
(click the figure number to see the cladogram)
+-- Lamellidens_mainwaringi
+-----------------------------------------------+
| +-- Trapezidens_exolescens
--+
| +----------------------------------------------- Leoparreysia_canefrii
| |
+--+ +-------------------------------------------- Parreysia_keralaensis
| |
+--+ +----------------------------------------- Coelatura_aegyptiaca
| |
| | +----- Indonaia_rugosa
+--+ +--------------------------------+
| | | +-- Indonaia_caerulea
| | +--+
| | +-- Indonaia_andersoniana
+--+
| +----------------------------------- Indochinella_pugio
| |
| | +----- Radiatula_mouhoti
+--+ +--------------------------+
| | | +-- Radiatula_myitkyinae
| | +--+
+--+ +-- Radiatula_chaudhurii
|
| +----------------------------- Scabiellus_songkramensis
+--+
| +-------------------------- Harmandia_somboriensis
+--+
| +----------------------- Unionetta_fabagina
+--+
| +-------------------- Scabies_crispata
+--+
| +----------------- Scabies_mandarinus
+--+
| +-------------- Scabies_pilata
+--+
| +----------- Scabies_nucleus
+--+
| +-------- Scabies_humilis
+--+
| +----- Scabies_scobinatus
+--+
| +-- Scabies_phaselus
+--+
+-- Scabies_anceps