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Data last updated
Mon 1 Dec 2025
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Publication Pfeiffer et al. (2021): taxonomic opinionspublication summary | cladograms
Complete CitationPfeiffer, J.M., D.L. Graf, K.S. Cummings & L.M. Page. 2021. Taxonomic revision of a radiation of Southeast Asian freshwater mussels (Unionidae: Gonideinae: Contradentini+Rectidentini). Invertebrate Systematics 35: 394-470. Available electronically.
Taxonomic OpinionsBalwantia soleniformis (Benson, 1836) Ctenodesma borneensis (Issel, 1874)
+ Unio plicatulus Lea, 1859
+ Unio pensitatus Fischer & Crosse, 1894 Ctenodesma scheibeneri Haas, 1927 Elongaria orientalis (Lea, 1840)
+ Unio productus Mousson, 1849
+ Unio productior Lea, 1852
+ Unio bithynicus Kobelt, 1893 Elongaria trompi (Drouët & Chaper, 1892) Ensidens dugasti (Morlet, 1892), stat. rev.
+ Ensidens sagittarius Muanta et al., 2019 Ensidens ingallsianus (Lea, 1852)
+ Unio pazii Lea, 1862 Ensidens jaculus (Rochebrune, 1882), stat. rev.
+ Ensidens sagittarius Muanta et al., 2019 Ensidens sagittarius (Lea, 1856) Ensidens spiculus sp. nov.
+ Ensidens sagittarius Bolotov et al., 2017 Ensidens telus sp. nov.
+ Ensidens ingallsianus Muanta et al., 2019
+ Ensidens sagittarius Pfeiffer & Graf, 2015 Hyriopsis bialata Simpson, 1900
+ Unio delphinus Lea, 1840
+ Unio delphinus Gruner, 1841
+ Unio megapterus Morelet, 1864
+ Unio delphinopterus Dautzenberg & Fischer, 1905
+ Hyriopsis gracilis Haas, 1910 Hyriopsis delaportei (Crosse & Fischer, 1876) Hyriopsis desowitzi Brandt, 1974 Hyriopsis khoratensis sp. nov. Hyriopsis kratiensis sp. nov. Hyriopsis myersiana (Lea, 1856)
+ Unio housei Lea, 1856
+ Unio sutrangensis Morlet, 1889
+ Unio stungtrangensis Fischer & Dautzenberg, 1904
+ Unio patrangensis Morlet, 1904
+ Hyriopsis delaportei Muanta et al., 2019 Hyriopsis phuphaniensis sp. nov. Hyriopsis sakhonensis sp. nov. Hyriopsis undescribed = Hyriopsis panhai Hyriopsis velthuizeni (Schepman, 1896) Lamprotula cornuumlunae Lens comptus (Deshayes in Deshayes & Jullien 1876), comb. nov.
+ Harmandia munensis Brandt, 1974
+ Contradens semmelinki fultoni Lopes-Lima et al., 2017 Lens contradens (Lea, 1838), comb. nov.
+ Contortus contortus Araujo et al., 2017
+ Unio javanus Lea, 1840
+ Unio exilis Dunker, 1846
+ Unio mutatus Mousson, 1849
+ Unio tumidulus Lea, 1856
+ Unio cambodiensis Lea, 1856
+ Unio ascia Hanley, 1856
+ Unio rusticus Lea, 1856
+ Unio sumatrensis Lea, 1859
+ Unio mederianus Küster, 1861
+ Unio paivanus Morelet, 1865
+ Unio asperulus Lea, 1866
+ Margaron (Unio) dimotus Lea, 1870
+ Unio semidecoratus Morlet, 1889
+ Unio dautzenbergi Morlet, 1889
9 more... Lens eximius (Lea, 1856)
+ Unio semiquadrata Sowerby, 1866 Lens inornatus (Lea, 1856), comb. nov. Lens lugens (Drouët & Chaper, 1892), comb. nov.
+ Unio saccellus Drouët & Chaper, 1892
+ Unio fulvaster Drouët & Chaper, 1892 Lens maenamensis sp. nov.
+ Contradens crossei Jeratthitikul et al., 2019 Lens micropterus (Morelet, 1866), comb. nov.
+ Unio cambojensis Sowerby, 1866
+ Margaron (Unio) versus Lea, 1870
+ Unio inornatus Reeve, 1865
+ Unio crossei Deshayes & Jullien, 1876
+ Unio semialatus Deshayes & Jullien, 1876
+ Diplodon ludovicianum Rochebrune, 1882
+ Unio inaequalis Rochebrune, 1882
+ Unio fischerianus Morlet, 1883
+ Unio siamensis Morlet, 1889
+ Unio soboles Fischer, 1891
+ Contradens contradens Pfeiffer & Graf, 2015 Lens misellus (Morelet, 1865)
+ Unio siamensis Lea, 1866 Lens novoselovi (Konopleva et al., 2019) Lens pallegoixi (Sowerby, 1867) Lens rolfbrandti (Jeratthitikul & Panha, 2019) Lens semmelinki (von Martens, 1891)
+ Unio semmelincki Haas, 1914 Leoparreysia subcircularis (Brandt, 1974) Monodontina cambodjensis (Petit, 1865) = Pseudodon cambodjensis Parvasolenaia neotriangularis Parvasolenaia rivularis Parvasolenaia triangularis Physunio massini (Morelet, 1864) Physunio modelli Brandt, 1974 Physunio pellucidus sp. nov. Physunio superbus (Lea, 1843)
+ Unio gravidus Lea, 1856
+ Unio velaris Hanley, 1856
+ Unio abnormis Morelet, 1862
+ Unio superbus Sowerby, 1867 Pressidens exanthematicus (Küster, 1861)
+ Pseudodon insularis Drouët, 1894
+ Pressidens moellendorffi Haas, 1910 Prisodon obliquus Schumacher, 1817
+ Unio macropterus Dunker, 1846 Prohyriopsis stolata (von Martens, 1900) Radiatula gratiosus (Philippi, 1843) comb. nov. = Indonaia gratiosa Rectidens sumatrensis (Dunker, 1852)
+ Unio perakensis Morgan, 1885
+ Unio lingulatus Drouët & Chaper, 1892
+ Unio prolongatus Drouët, 1894
+ Microcondylaea gracilis Strubell, 1897
+ Unio palembangensis Strubell, 1897
+ Unio pahangensis E.A. Smith, 1899
+ Unio palembangensis var. pahumbaensis Martens, 1900
+ Unio pressirostris Martens, 1900
+ Unio perakensis Preston, 1912
+ Rectidens prestoni Simpson, 1914 Solenaia carinata = Sinosolenaia carinata Solenaia emarginata (Lea, 1860)
+ Solenaia khwaenoiensis Deein et al., 2004 Solenaia oleivora = Sinosolenaia iridinea
+ Mycetopus iridineus Heude, 1874 Solenaia oleivora = Sinosolenaia oleivora Trapezoideus foliaceus (Gould, 1843) Trapezoideus lenya Bolotov, Konopleva, Vikhrev, Gofarov, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Tomilova, Tanmuangpak, Tumpeesuwan & Kondakov, 2020 Trapezoideus subviridis sp. nov. Yaukthwa avaensis Bolotov, Konopleva, Vikhrev, Gofarov, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Tomilova, Tanmuangpak, Tumpeesuwan & Kondakov, 2020 Yaukthwa baniensis (Bolotov, Konopleva, Vikhrev, Gofarov, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Tomilova, Tanmuangpak, Tumpeesuwan & Kondakov, 2020), comb. nov.
+ Balwantia myanmarensis Bolotov et al., 2020 Yaukthwa dalliana (Frierson, 1913) Yaukthwa elongatula Bolotov, Konopleva, Vikhrev, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Gofarov, Tomilova & Kondakov, 2019, comb. res. Yaukthwa inlenensis Konopleva, Pfeiffer, Vikhrev, Kondakov, Gofarov, Aksenova, Lunn, Chan & Bolotov, 2019 Yaukthwa nesemanni (Konopleva, Vikhrev & Bolotov in Bolotov et al., 2017) Yaukthwa paiensis Konopleva, Pfeiffer, Vikhrev, Kondakov, Gofarov, Aksenova, Lunn, Chan & Bolotov, 2019 Yaukthwa panhai (Konopleva, Bolotov & Kondakov in Bolotov et al., 2017) Yaukthwa peguensis (Anthony, 1865) Yaukthwa rectangularis (Tapparone-Canefri, 1889), comb. nov. Yaukthwa zayleymanensis (Preston, 1912)
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