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Wed 1 October 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

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Publication Inoue et al. (2020): cladograms
     publication summary | cladograms

Complete Citation

Inoue, K., K.S. Cummings, J.S. Tiemann, T.D. Miller, N.A. Johnson et al. 2020. A new species of freshwater mussel in the genus Popenaias Frierson, 1927, from the Gulf coast rivers of central Mexico (Bivalvia: Unionida: Unionidae) with comments on the genus. Zootaxa 4816(4): 457-490. Available electronically.

Cladogram Figures

(click the figure number to see the cladogram)

Cladogram Figure 2
  +----------------------- Quadrula_quadrula          
  |                                                   
  |                    +-- Reginaia_ebenus            
  |  +-----------------+                              
  |  |                 +-- Reginaia_rotulata          
--+  |                                                
  |  |              +----- Amblema_plicata            
  |  |  +-----------+                                 
  |  |  |           |  +-- Amblema_elliottii          
  |  |  |           +--+                              
  +--+  |              +-- Amblema_neislerii          
     |  |                                             
     |  |           +----- Popenaias_popeii           
     |  |        +--+                                 
     |  |        |  |  +-- Popenaias_berezai          
     |  |        |  +--+                              
     |  |        |     +-- Popenaias_semirasa         
     +--+        |                                    
        |  +-----+     +-- Psoronaias_crocodilorum__1 
        |  |     |     |                              
        |  |     |     +-- Psoronaias_crocodilorum__2 
        |  |     |     |                              
        |  |     |     +-- Psoronaias_crocodilorum__3 
        |  |     |     |                              
        |  |     +-----+-- Psoronaias_guatemalensis__1
        |  |           |                              
        |  |           +-- Psoronaias_guatemalensis__2
        +--+           |                              
           |           +-- Psoronaias_crocodilorum__4 
           |           |                              
           |           +-- Psoronaias_crocodilorum__5 
           |                                          
           |           +-- Glebula_rotundata          
           |  +--------+                              
           |  |        +-- Cyrtonaias_berlandierii    
           |  |                                       
           +--+  +-------- Potamilus_alatus           
              |  |                                    
              |  |     +-- Ptychobranchus_fasciolaris 
              +--+  +--+                              
                 |  |  +-- Truncilla_truncata         
                 |  |                                 
                 |  |  +-- Ortmanniana_ligamentina    
                 +--+  |                              
                    |  +-- Epioblasma_triquetra       
                    |  |                              
                    |  +-- Lampsilis_cardium          
                    |  |                              
                    +--+-- Ligumia_recta              
                       |                              
                       +-- Cambarunio_iris            
                       |                              
                       +-- Pachynaias_spheniopsis     
                       |                              
                       +-- Mesonaias_explicata        
Cladogram Figure 3
  +----------------------------------- Quadrula_quadrula          
  |                                                               
  |                                +-- Reginaia_ebenus            
  |  +-----------------------------+                              
--+  |                             +-- Reginaia_rotulata          
  |  |                                                            
  |  |                          +----- Amblema_neislerii          
  |  |  +-----------------------+                                 
  |  |  |                       |  +-- Amblema_plicata            
  +--+  |                       +--+                              
     |  |                          +-- Amblema_elliottii          
     |  |                                                         
     |  |                       +----- Popenaias_popeii           
     |  |              +--------+                                 
     |  |              |        |  +-- Popenaias_berezai          
     +--+              |        +--+                              
        |  +-----------+           +-- Popenaias_semirasa         
        |  |           |                                          
        |  |           |  +----------- Psoronaias_crocodilorum__4 
        |  |           |  |                                       
        |  |           +--+        +-- Psoronaias_crocodilorum__1 
        |  |              |  +-----+                              
        |  |              |  |     +-- Psoronaias_guatemalensis__1
        |  |              +--+                                    
        +--+                 |  +----- Psoronaias_crocodilorum__3 
           |                 +--+                                 
           |                    |  +-- Psoronaias_guatemalensis__2
           |                    +--+                              
           |                       +-- Psoronaias_crocodilorum__2 
           |                                                      
           |  +----------------------- Cyrtonaias_berlandierii    
           |  |                                                   
           +--+  +-------------------- Ptychobranchus_fasciolaris 
              |  |                                                
              +--+  +----------------- Ligumia_recta              
                 |  |                                             
                 +--+  +-------------- Potamilus_alatus           
                    |  |                                          
                    +--+  +----------- Pachynaias_spheniopsis     
                       |  |                                       
                       |  |        +-- Epioblasma_triquetra       
                       +--+  +-----+                              
                          |  |     +-- Truncilla_truncata         
                          |  |                                    
                          +--+     +-- Mesonaias_explicata        
                             |  +--+                              
                             |  |  +-- Cambarunio_iris            
                             +--+                                 
                                |  +-- Ortmanniana_ligamentina    
                                +--+                              
                                   +-- Lampsilis_cardium          
 
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