species Beringiana beringiana (Middendorff, 1851): synonymy

Anodonta cellensis var. beringiana Middendorff, 1851

Middendorff, 1851. Reise Nord. Ost. Sibiriens 2(1): 284, pl. 28, figs. 4-7; pl. 29, figs. 1-4.
(intro'd as intra-sp. of Anodonta cellensis Pfeiffer, 1821)

Anodonta beringiana — Simpson (1900), Simpson (1914), Burch (1975), R.I. Johnson (1980), Turgeon et al. (1988), ... Froufe et al. (2017), Zieritz et al. (2018), Inoue et al. (2018), Tomilova et al. (2019), Galal-Khallaf et al. (2024)
Anodonta cygnea beringiana — Hannibal (1912)
Anodonta (s.s.) beringiana — Frierson (1927), Zhadin (1938), Zhadin (1965), Haas (1969)
Anodonta marginata beringiana — Modell (1945)
Anodonta (Sinanodonta) beringiana — Habe (1977)
Beringiana beringiana — Zatravkin (1983), Zatravkin & Bogatov (1987), Korniushin (1998), Bogatov & Starobogatov (2001), Starobogatov et al. (2004), ... Dai et al. (2025), Wu et al. (2025), FMCS (2025), Chen et al. (2025), Dai et al. (2026)
Beringiana baringiana [sic] — Bogatov & Zatravkin (1988)
Sinanodonta beringiana — Williams et al. (2017), FMCS (2021)
Buldowskia beringiana beringiana — Bogatov (2022)

Anodonta youcanensis Lea, 1867

‘Lea’ Lea, 1867. Obs., index I-XI: 63.
‘Lea’ Paetel, 1890. Cat. Conch.-Samml. 3: 186.
(error for Anodonta youkanensis Lea, 1867)

Anodonta beringiana — Simpson (1900), Simpson (1914)
Beringiana beringiana — Bieler (2021)

Anodonta youconensis Lea, 1867

Lea, 1867. Proc. Acad. Nat. Sci. 19: 81.
(incorrect original spelling of Anodonta youkanensis Lea, 1867)

Anodonta beringiana — Simpson (1900), Simpson (1914)
Anodonta cygnea beringiana — Hannibal (1912)
Anodonta (s.s.) beringiana — Frierson (1927)
Anodonta (s.s.) oregonensis — Haas (1969)
Beringiana youkanensis — Zatravkin & Bogatov (1987)
Beringiana youconensis — Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Beringiana beringiana — Bieler (2021)

Anodonta youkanensis Lea, 1867

Lea, 1867. Proc. Acad. Nat. Sci. 19: 81 [as ‘youconensis’].
Lea, 1868. Jour. Acad. Nat. Sci. 6 [O. 12]: 287 [47], pl. 40, fig. 99.
(correct name for Anodonta youconensis Lea, 1867)
Type(s): FIGURED SYNTYPE USNM_86624, Upper Yukon R., Alaska.

Anodonta beringiana — Simpson (1900), Simpson (1914), Burch (1975), Graf (2007), IUCN (2011)
Anodonta cygnea beringiana — Hannibal (1912)
Anodonta (s.s.) beringiana — Frierson (1927)
Anodonta (s.s.) oregonensis — Haas (1969)
Beringiana youkonensis [sic] — Zatravkin (1983), Bogatov & Starobogatov (2001), Starobogatov et al. (2004), Sayenko et al. (2009)
Beringiana youkanensis — Zatravkin & Bogatov (1987)
Beringiana yuokonensis [sic] — Bogatov & Zatravkin (1988)
Beringiana youconensis — Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Beringiana beringiana — Lopes-Lima et al. (2020), Bolotov et al. (2020), Bieler (2021)
Buldowskia beringiana youkonensis [sic] — Bogatov (2022)

Anodonta youkonensis ‘Lea’ of Authors

‘Lea’ of Authors.
Simpson, 1893. Nautilus 6(12): 135.
Zatravkin, 1983. [in] I.M. Likharev, Molluscs: 42.
Bogatov & Starobogatov, 2001. Zool. Zhur. 80: 30.
Starobogatov et al., 2004. Key Freshw. Invert. Russia 6 (molluscs, polychaetes, nemerteans): 41.
Kantor & Sysoev, 2005. Cat. Moll. Russia: 332.
Sayenko et al., 2009. Zool. Zhur. 88(11): 1306.
Bogatov, 2022. Lg. Bivalve Moll. Russia’s Fresh Waters: 73.
(error for Anodonta youkanensis Lea, 1867)

Beringiana youkonensis [sic] — Bogatov & Starobogatov (2001)
Beringiana beringiana — Bieler (2021), Bulakhova et al. (2023)

Anodonta beringiana var. taranetzi Zhadin, 1938

Zhadin, 1938. Faune USSR Mollusques 4(1): 133, fig. 49.
Zhadin, 1965. Moll. USSR: 255, fig. 273.
(intro'd as intra-sp. of Anodonta cellensis var. beringiana Middendorff, 1851)

Anodonta (s.s.) beringiana var. taranetzi — Zhadin (1938), Zhadin (1965)
Anodonta (s.s.) beringiana — Haas (1969)
Beringiana taranetzi — Zatravkin (1983), Zatravkin & Bogatov (1987), Bogatov & Zatravkin (1988), Vinarski & Kantor (2016)
Kunashiria taranetzi — Starobogatov et al. (2004), Sayenko et al. (2009)
Kunashiria taranetsi [sic] — Kantor & Sysoev (2005), Kantor et al. (2010)
Anodonta woodiana japonica — Graf (2007)
Sinanodonta woodiana — He & Zhuang (2013)
Beringiana beringiana — Lopes-Lima et al. (2020), Bolotov et al. (2020)
Kunashiria japonica taranetzi — Bogatov (2022)

Amuranodonta sihotealinica Zatravkin & Starobogatov, 1984

Zatravkin & Starobogatov, 1984. Zool. Zhur. 63(12): 1788, figs. 9-10.

Amuranodonta sp. nov. — Zatravkin (1983)
Arsenievinaia sihotealinica — Zatravkin & Bogatov (1987), Bogatov & Zatravkin (1988), Martynov & Chernyshev (1992), Bogatov & Sayenko (2002)
Kunashiria haconensis — Starobogatov et al. (2004), Kantor & Sysoev (2005), Sayenko et al. (2009), Kantor et al. (2010)
Sinanodonta woodiana — He & Zhuang (2013)
Beringiana haconensis — Vinarski & Kantor (2016)
Beringiana beringiana — Lopes-Lima et al. (2020), Bolotov et al. (2020)
Kunashiria japonica haconensis — Bogatov (2022)

Arsenievinaia coptzevi Zatravkin & Bogatov, 1987

Zatravkin & Bogatov, 1987. Biv. Far East: 104, sp. 51, figs. 20 б, 24 в-г.

Arsenievinaia coptzevi — Zatravkin & Bogatov (1987), Bogatov & Zatravkin (1988)
Kunashiria coptzevi — Starobogatov et al. (2004), Kantor & Sysoev (2005), Sayenko et al. (2009), Kantor et al. (2010)
Anodonta woodiana japonica — Graf (2007)
Sinanodonta woodiana — He & Zhuang (2013)
Beringiana coptzevi — Vinarski & Kantor (2016)
Beringiana beringiana — Lopes-Lima et al. (2020), Bolotov et al. (2020)
Kunashiria japonica haconensis — Bogatov (2022)

Arsenievinaia zimini Zatravkin & Bogatov, 1987

Zatravkin & Bogatov, 1987. Biv. Far East: 102, sp. 50, figs. 24 а-б.

Arsenievinaia zimini — Zatravkin & Bogatov (1987), Bogatov & Zatravkin (1988)
Arsenievinaia sihotealinica — Martynov & Chernyshev (1992)
Kunashiria zimini — Starobogatov et al. (2004), Kantor & Sysoev (2005), Sayenko et al. (2009), Kantor et al. (2010)
Anodonta woodiana japonica — Graf (2007)
Sinanodonta woodiana — He & Zhuang (2013)
Beringiana zimini — Vinarski & Kantor (2016)
Beringiana beringiana — Lopes-Lima et al. (2020), Bolotov et al. (2020)
Kunashiria japonica haconensis — Bogatov (2022)

Arsenievinaia alimovi Bogatov & Zatravkin, 1988

Bogatov & Zatravkin, 1988. Trudy Zool. Inst. 187: 165, fig. 5.

Arsenievinaia alimovi — Bogatov & Zatravkin (1988)
Kunashiria coptzevi — Starobogatov et al. (2004), Kantor & Sysoev (2005), Sayenko et al. (2009), Kantor et al. (2010)
Sinanodonta woodiana — He & Zhuang (2013)
Beringiana coptzevi — Vinarski & Kantor (2016)
Beringiana beringiana — Lopes-Lima et al. (2020), Bolotov et al. (2020)
Kunashiria japonica taranetzi — Bogatov (2022)

Arsenievinaia zarjaensis Bogatov & Zatravkin, 1988

Bogatov & Zatravkin, 1988. Trudy Zool. Inst. 187: 166, fig. 6.

Arsenievinaia zarjaensis — Bogatov & Zatravkin (1988)
Kunashiria zarjaensis — Starobogatov et al. (2004), Kantor & Sysoev (2005), Sayenko et al. (2009), Kantor et al. (2010)
Anodonta woodiana japonica — Graf (2007)
Sinanodonta woodiana — He & Zhuang (2013)
Beringiana zarjaensis — Vinarski & Kantor (2016)
Beringiana beringiana — Lopes-Lima et al. (2020), Bolotov et al. (2020)
Kunashiria japonica japonica — Bogatov (2022)

Arsenievinaia compressa Bogatov & Starobogatov, 1996

Bogatov & Starobogatov, 1996. Zool. Zhur. 75: 1332. fig’d.

Kunashiria compressa — Starobogatov et al. (2004), Kantor & Sysoev (2005), Sayenko et al. (2009), Kantor et al. (2010)
Anodonta woodiana japonica — Graf (2007)
Sinanodonta woodiana — He & Zhuang (2013)
Beringiana compressa — Vinarski & Kantor (2016)
Beringiana beringiana — Lopes-Lima et al. (2020), Bolotov et al. (2020)
Kunashiria japonica f. compressa — Bogatov (2022)

Arsenievinaia zatravkini Bogatov & Starobogatov, 1996

Bogatov & Starobogatov, 1996. Zool. Zhur. 75: 1331, fig’d.

Kunashiria taranetzi — Starobogatov et al. (2004)
Kunashiria taranetsi [sic] — Kantor & Sysoev (2005), Kantor et al. (2010)
Beringiana taranetzi — Vinarski & Kantor (2016)
Beringiana beringiana — Lopes-Lima et al. (2020), Bolotov et al. (2020)
Kunashiria japonica taranetzi — Bogatov (2022)

Beringiana compressa Sayenko & Bogatov, 1998

Sayenko & Bogatov, 1998. Zool. Zhur. 77(12): 1414, figs. 2-4.
(homonym of Arsenievinaia compressa Bogatov & Starobogatov, 1996)

Beringiana compressa — Bogatov & Starobogatov (2001), Starobogatov et al. (2004), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Anodonta beringiana — Graf (2007)
Beringiana beringiana — Lopes-Lima et al. (2020), Bolotov et al. (2020), Bulakhova et al. (2023)
Buldowskia beringiana f. compressa — Bogatov (2022)

Anemina (Buldowskia) adotymensys Labay & Shulga, 1999

Labay & Shulga, 1999. Ruthenica 9: 79, fig’d.

Amuranodonta sitaensis — Starobogatov et al. (2004), Kantor & Sysoev (2005), Kantor et al. (2010)
Anemina arcaeformis — He & Zhuang (2013)
Anemina (Buldowskia) sitaensis — Vinarski & Kantor (2016)
Beringiana beringiana — Lopes-Lima et al. (2020), Bolotov et al. (2020)

Anemina (Buldowskia) lacustris Labay & Shulga, 1999

Labay & Shulga, 1999. Ruthenica 9: 78, fig’d.

Amuranodonta parva — Starobogatov et al. (2004), Kantor & Sysoev (2005), Kantor et al. (2010)
Anemina arcaeformis — He & Zhuang (2013)
Anemina (Buldowskia) parva — Vinarski & Kantor (2016)
Beringiana beringiana — Lopes-Lima et al. (2020), Bolotov et al. (2020)

Kunashiria iturupica Bogatov et al., 1999

Bogatov et al., 1999. Ruthenica 9: 60, fig’d.

Kunashiria iturupica — Starobogatov et al. (2004), Kantor & Sysoev (2005), Sayenko et al. (2009), Kantor et al. (2010)
Anodonta woodiana japonica — Graf (2007)
Sinanodonta woodiana — He & Zhuang (2013)
Beringiana iturupica — Vinarski & Kantor (2016)
Beringiana beringiana — Lopes-Lima et al. (2020), Bolotov et al. (2020)
Kunashiria japonica taranetzi — Bogatov (2022)

Kunashiria sinanodontoides Bogatov et al., 1999

Bogatov et al., 1999. Ruthenica 9: 59, fig’d.

Kunashiria sinanodontoides — Starobogatov et al. (2004), Kantor & Sysoev (2005), Sayenko et al. (2009), Kantor et al. (2010)
Anodonta woodiana japonica — Graf (2007)
Sinanodonta woodiana — He & Zhuang (2013)
Beringiana sinanodontoides — Vinarski & Kantor (2016)
Beringiana beringiana — Lopes-Lima et al. (2020), Bolotov et al. (2020)
Kunashiria japonica taranetzi — Bogatov (2022)

Kunashiria japonica boreosakhalinensis Labay & Shulga, 1999

Labay & Shulga, 1999. Ruthenica 9: 77, fig’d.
(intro'd as intra-sp. of Anodonta japonica Clessin, 1874)

Kunashiria haconensis — Starobogatov et al. (2004), Kantor & Sysoev (2005), Kantor et al. (2010)
Beringiana haconensis — Vinarski & Kantor (2016)
Beringiana beringiana — Lopes-Lima et al. (2020), Bolotov et al. (2020)
Kunashiria japonica japonica — Bogatov (2022)

Beringiana chereshnevi Bogatov & Starobogatov, 2001

Bogatov & Starobogatov, 2001. Zool. Zhur. 80: 26, figs. 2а-б, 4ж.

Beringiana chereshnevi — Bogatov & Starobogatov (2001), Starobogatov et al. (2004), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Anodonta beringiana — Graf (2007)
Beringiana beringiana — Lopes-Lima et al. (2020), Bolotov et al. (2020)
Buldowskia beringiana youkonensis [sic] — Bogatov (2022)

Beringiana derzhavini Bogatov & Starobogatov, 2001

Bogatov & Starobogatov, 2001. Zool. Zhur. 80: 27, figs. 2в-г, 4а.

Beringiana derzhavini — Bogatov & Starobogatov (2001), Starobogatov et al. (2004), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Anodonta beringiana — Graf (2007)
Beringiana beringiana — Lopes-Lima et al. (2020), Bolotov et al. (2020), Bulakhova et al. (2023)
Buldowskia beringiana kamchatica — Bogatov (2022)

Beringiana georgiensis Bogatov & Starobogatov, 2001

Bogatov & Starobogatov, 2001. Zool. Zhur. 80: 28, figs. 2ж-з, 4з.

Beringiana georgiensis — Bogatov & Starobogatov (2001)
Anodonta beringiana — Graf (2007)
Buldowskia beringiana youkonensis [sic] — Bogatov (2022)

Beringiana kamchatica Bogatov & Starobogatov, 2001

Bogatov & Starobogatov, 2001. Zool. Zhur. 80: 27, figs. 2д-е, 4г.

Beringiana kamchatica — Bogatov & Starobogatov (2001), Starobogatov et al. (2004), Kantor & Sysoev (2005), Sayenko et al. (2009), Kantor et al. (2010), Vinarski & Kantor (2016)
Anodonta beringiana — Graf (2007)
Beringiana beringiana — Lopes-Lima et al. (2020), Bolotov et al. (2020), Bulakhova et al. (2023)
Buldowskia beringiana kamchatica — Bogatov (2022)

Sinanodonta japonica ‘Clessin’ of Authors

‘Clessin’ of Authors.
Sano et al., 2017. J. Wat. Res. Prot. 9: Table 2. [in part]
Sano et al., 2020. Plankton Benthos Res. 15(2): Table 1. [in part]
(misidentified reference to Anodonta japonica Clessin, 1874)

Beringiana beringiana — Sano et al. (2022), Bulakhova et al. (2023)

Anodon youkamensis Sowerby, 1870

‘Lea’ Sowerby, 1870. Conch. Icon. 17: index.
(error for Anodonta youkanensis Lea, 1867)

Beringiana beringiana — Bieler (2021)

Anodonta yukonensis Simpson, 1896

‘Lea’ Simpson, 1896. Proc. U.S. Nat. Mus. 18: 328.
(error for Anodonta youkanensis Lea, 1867)

Beringiana beringiana — Bieler (2021)

Beringiana baringiana Bogatov & Zatravkin, 1988

‘Middendorff’ Bogatov & Zatravkin, 1988. Trudy Zool. Inst. 187: 167.
(error for Anodonta cellensis var. beringiana Middendorff, 1851)

Beringiana yuokonensis Bogatov & Zatravkin, 1988

‘Lea’ Bogatov & Zatravkin, 1988. Trudy Zool. Inst. 187: 167.
(error for Anodonta youkanensis Lea, 1867)

Amuranodonta sichotealinica Korniushin, 1998

‘Zatravkin & Starobogativ’ Korniushin, 1998. Malac. Review Bivalvia 7: 79.
(error for Amuranodonta sihotealinica Zatravkin & Starobogatov, 1984)

Beringiana georginesis Bogatov & Starobogatov, 2001

Bogatov & Starobogatov, 2001. Zool. Zhur. 80: 28.
(error for Beringiana georgiensis Bogatov & Starobogatov, 2001)

Kunashiria taranetsi Kantor & Sysoev, 2005

‘Shadin’ Kantor & Sysoev, 2005. Cat. Moll. Russia: 339.
Kantor et al., 2010. Cat. Contin. Moll. Russia: 279.
(error for Anodonta beringiana var. taranetzi Zhadin, 1938)

Beringiana georginensis Graf, 2007

‘Bogatov & Starobogatov’ Graf, 2007. Proc. Acad. Nat. Sci. 156: 86.
(error for Beringiana georgiensis Bogatov & Starobogatov, 2001)