Sayenko, E.M., M. Soroka, Y.B. Akiyama, T. Uechi, K. Ito & M. Kondo. 2021. Taxonomic status of genera Nodularia, Middendorffinaia and Inversiunio (Bivalvia: Unionidae) from South-East Asia: morphometric, genetic and GenBank data. Systematics and Biodiversity 19: 54-73. Available electronically.
(click the figure number to see the cladogram)
+-- Unio_pictorum
+--------------------+
| +-- Unio_nanus
|
| +-------- Inversiunio_jokohamensis
--+ |
| +-----------+ +----- Inversiunio_yanagawensis__2
| | | |
| | +--+----- Inversiunio_verrucosus
| | |
+--+ | +-- Inversiunio_reinianus
| +--+
| +-- Inversiunio_yanagawensis__1
|
| +----------------- Middendorffinaia_mongolica
+--+
| +-------------- Nodularia_breviconcha
+--+
| +----------- Nodularia_huana
+--+
| +-------- Nodularia_nuxpersicae
+--+
| +----- Nodularia_nipponensis
+--+
| +-- Nodularia_douglasiae
+--+
+-- Nodularia_dualobtusa