Page last updated
Wed 1 October 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Publication Lopes-Lima et al. (2021): taxonomic opinions
     publication summary | cladograms

Complete Citation

Lopes-Lima, M., M.E. Gürlek, Ü. Kebapçi, H. ŞerefliŞan, T. Yanik et al. 2021. Diversity, biogeography, evolutionary relationships, and conservation of Eastern Mediterranean freshwater mussels (Bivalvia: Unionidae). Molecular Phylogenetics and Evolution 163. 107261 (23 pp.). Available electronically.

Taxonomic Opinions

Amblema plicata

Anodonta anatina (Linnaeus, 1758)
+ Anodonta cyrea Drouët, 1881
+ Anodonta taurica Bourguignat, 1880
+ Anodonta apollonica Bourguignat, 1880
+ Anodonta gallandi Bourguignat, 1880
+ Anodonta byzantina Drouët, 1881
+ Anodonta ectina Bourguignat, 1881
+ Anodonta gaudioni Drouët, 1881
+ Anodonta georgiana Drouët, 1881
+ Anodonta lenkoranensis Drouët, 1881
+ Anodonta mahometana Bourguignat, 1881
+ Anodonta maritzana Bourguignat, 1881
+ Anodonta pseudodopsis Locard, 1883
+ Anodonta brandti Drouët, 1888
+ Anodonta cilicica Kobelt & Rolle, 1895
+ Anodonta cygnea var. armenica Ovtschinnikov, 1935

Anodonta cygnea (Linnaeus, 1758)
+ Anodonta gravida Drouët, 1879

Anodonta exulcerata

Anodonta seddoni sp. nov. = Anodonta seddonae

Anodonta vescoiana Bourguignat, 1856
+ Anodonta schlaeflii Mousson, 1874
+ Anodonta vescoiana var. mesopotamica Mousson, 1874
+ Anodonta (Euphrata) bahlikiana Pallary, 1939

Anodontoides denigratus

Coelatura aegyptiaca

Gonidea angulata (Lea, 1838)
+ Anodonta randalli Trask, 1854
+ Anodonta randalli Trask, 1854

Lamellidens corrianus

Lamprotula leaii

Lampsilis ovata

Leguminaia anatolica sp. nov.

Leguminaia cf. saulcyi (Bourguignat, 1852) = Leguminaia saulcyi
+ Unio michonii Bourguignat, 1852
+ Unio tripolitanus Bourguignat, 1852
+ Pseudodon chantrei Locard, 1883
+ Leguminaia (sec. Leguminaia) locardi Simpson, 1900
+ Leguminaia tripolitana var. kebirensis Pallary, 1939

Leguminaia cf. wheatleyi (Lea, 1862) = Leguminaia wheatleyi
+ Monocondyloea mardinensis Lea, 1865
+ Leguminaia bourguignati Locard, 1883
+ Leguminaia wheatleyi var. euphratica Kobelt, 1912
+ Leguminaia (mardinensis var.) naegelei Kobelt, 1913
+ Leguminaia graeteri Kobelt, 1913
+ Leguminaia nisibina Kobelt, 1913
+ Microdondylaea haasii Kobelt, 1914
+ Leguminaia mardinensis var. truncata Pallary, 1939

Microcondylaea bonellii

Middendorffinaia mongolica

Nodularia douglasiae

Potomida acarnanica

Potomida littoralis (Cuvier, 1798)
+ Mysca (Potomida) corrugata Swainson, 1840

Potomida semirugata (Lamarck, 1819)
+ Unio delesserti Bourguignat, 1852
+ Unio emesaensis Lea, 1865
+ Unio rothi Bourguignat, 1863
+ Unio simonis Tristram, 1865
+ Unio ferussacianus Lea, 1868
+ Unio galilaei Locard, 1883
+ Unio luynesi Locard, 1883
+ Unio rhomboidopsis Locard, 1883
+ Unio timius Locard, 1883
+ Unio tinctus Drouët, 1893
+ Unio rollei Kobelt & Rolle, 1895
+ Unio lycicus Kobelt & Rolle, 1895
+ Unio wagneri Kobelt & Rolle, 1895
+ Unio tracheae Kobelt & Rolle, 1895
+ Unio (Rhombunio ?) abrus Kobelt, 1913
12 more...

Pronodularia japanensis

Pseudanodonta complanata

Pseudodontopsis euphratica Bourguignat, 1852
+ Unio opperti Bourguignat, 1856
+ Unio churchillianus Bourguignat, 1857
+ Monocondyloea rhomboidea Lea, 1859
+ Pseudodon pachyolenus Bourguignat, 1880
+ Pseudodontopsis babylonica Kobelt, 1913
+ Pseudodontopsis piestius Kobelt, 1913

Pseudunio homsensis

Pyganodon cataracta

Sinanodonta woodiana

Unio bruguierianus Bourguignat, 1853
+ Unio orientalis Bourguignat, 1852
+ Unio prusii Bourguignat, 1856
+ Unio vescoi Bourguignat, 1856
+ Unio schwerzenbachi Bourguignat, 1856
+ Unio kotschyi Küster, 1862
+ Unio turcicus Küster, 1862
+ Unio kullethensis Lea, 1865
+ Unio mardinensis Lea, 1865
+ Unio orphaensis Lea, 1865
+ Unio vicarius Westerlund & Blanc, 1879
+ Unio rothi var. komarowi O. Boettger, 1880
+ Unio araxenus Drouët, 1881
+ Unio heldreichi O. Boettger, 1886
+ Unio thiesseae Drouët, 1892
+ Unio (sec. Lymnium) pseudonymus Simpson, 1900
15 more...

Unio bruguierianus Bourguignat, 1853 = Unio mardinensis

Unio bruguierianus Bourguignat, 1853 = Unio vicarius

Unio caffer

Unio cf. courtillieri = Unio crassus

Unio cf. desectus = Unio desectus

Unio crassus

Unio damascensis Lea, 1863
+ Unio orontesensis Lea, 1863
+ Unio syriacus Lea, 1863

Unio delicatus Lea, 1863
+ Unio anemprosthus Locard, 1883
+ Unio antiochianus Locard, 1883
+ Unio axiacus Locard, 1883
+ Unio chantrei Locard, 1883
+ Unio jauberti Locard, 1883
+ Unio subtigridis Locard, 1883
+ Unio alexandri Kobelt & Rolle, 1895
+ Unio cilicicus Kobelt & Rolle, 1895
+ Unio cilicicus var. jenemterensis Kobelt & Rolle, 1895
+ Unio cilicicus var. adanensis Kobelt & Rolle, 1895
+ Unio cilicicus var. subsaccatus Kobelt & Rolle, 1895
+ Unio (cilicicus var. ?) berytensis Kobelt & Rolle, 1895
+ Unio kobelti Kobelt & Rolle, 1895
+ Unio kuweikensis Kobelt, 1913
+ Unio (cilicicus var.) anceyi Kobelt, 1913
2 more...

Unio delphinus

Unio durieui

Unio elongatulus

Unio eucirrus Bourguignat, 1857

Unio foucauldianus

Unio gibbus

Unio hueti Bourguignat, 1855

Unio ionicus

Unio mancus

Unio pictorum (Linnaeus, 1758)
+ Unio gaudioni Drouët, 1881
+ Unio ascanius Kobelt, 1913
+ Unio gentilis Hesse, 1911

Unio ravoisieri

Unio sesirmensis Kobelt, 1913

Unio terminalis Bourguignat, 1852
+ Unio grelloisianus Bourguignat, 1856
+ Unio jordanicus Bourguignat, 1856
+ Unio lunulifer Bourguignat, 1856
+ Unio lorteti Locard, 1883
+ Unio pietri Locard, 1883
+ Unio ellipsoideus Locard, 1883
+ Unio genezarethanus Locard, 1883
+ Unio prosacrus Locard, 1883
+ Unio raymondi Locard, 1883
+ Unio tiberiadensis Locard, 1883
+ Unio tristrami Locard, 1883
+ Unio zabulonicus Locard, 1883
+ Unio herodes Kobelt & Rolle, 1895
+ Unio kisonis Kobelt & Rolle, 1895
+ Unio raymondopsis Kobelt, 1913
1 more...

Unio tigridis Bourguignat, 1852
+ Unio mussolianus Küster, 1861
+ Unio bourguignatianus Lea, 1863
+ Unio dignatus Lea, 1863
+ Unio mosulensis Lea, 1863
+ Unio rasus Lea, 1863
+ Margaron (Unio) tigris Lea, 1870
+ Unio calliopsis Kobelt, 1913
+ Unio tigridis var. haruni Kobelt, 1912
+ Unio hyperamblius Kobelt, 1913
+ Unio siouffi Kobelt, 1913
+ Unio dignatus assuricus Kobelt, 1913
+ Unio dignatus ninusi Kobelt, 1913
+ Unio dignatus semiramidis Kobelt, 1913
+ Unio (Nemrodia) tammouziana Pallary, 1939
+ Unio (Nemrodia) tammouziana var. major Pallary, 1939
3 more...

Unio tumidiformis

Unio tumidus Philipsson in Retzius, 1788

Utterbackia peggyae