Zieritz, A., E. Froufe, I. Bolotov, D,V, Gonçakves, D.C. Aldridge et al. 2021. Mitogenomic phylogeny and fossil-calibrated mutation rates for all F- and M-type mtDNA genes of the largest freshwater mussel family, the Unionidae (Bivalvia). Zoological Journal of the Linnean Society 193(3): 1088-1107. Available electronically.
(click the figure number to see the cladogram)
+----- Pseudunio_marocanus
+--------------------------+
| | +-- Margaritifera_falcata
| +--+
| +-- Margaritifera_dahurica
|
| +----------- Quadrula_quadrula
| |
| +-----------------+ +-------- Toxolasma_parvum
| | | |
| | +--+ +-- Potamilus_alatus
| | | +--+
| | | | +-- Potamilus_leptodon
--+ | +--+
| | | +-- Venustaconcha_ellipsiformis
| | +--+
| | +-- Lampsilis_ornata
| |
| | +----- Chamberlainia_somsakpanhai
| | +--------------+
| | | | +-- Sinohyriopsis_cumingii
| | | +--+
| | | +-- Sinohyriopsis_schlegelii
| | |
| | | +-- Lens_contradens
+--+ +-----+ +--+
| | | | +-- Physunio_superbus
| | | +-----------+
| | | | | +-- Hyriopsis_bialata
| | | | +--+
| | | | +-- Rectidens_sumatrensis
| | | |
| | +--+ +-------- Potomida_littoralis
| | | +-----+
| | | | | +----- Pronodularia_japanensis
| | | | +--+
| | | | | +-- Lamprotula_caveata
| | | | +--+
| | +--+ +-- Lamprotula_leaii
| | |
| | | +-- Pilsbryoconcha_exilis
| | | +--------+
+--+ | | +-- Pseudodon_vondembuschianus
| +--+
| | +-------- Ptychorhynchus_pfisteri
| +--+
| | +----- Microcondylaea_bonellii
| +--+
| | +-- Sinosolenaia_oleivora
| +--+
| +-- Sinosolenaia_carinata
|
| +----------------------- Lepidodesma_languilati
| |
| | +----- Aculamprotula_tortuosa
| | +--------+
| | | | +-- Aculamprotula_tientsinensis
| | | +--+
| | | +-- Aculamprotula_coreana
| | +-----+
+--+ | | +----- Nodularia_douglasiae
| | | +-----+
| | | | | +-- Schistodesmus_lampreyanus
| | | | +--+
| | +--+ +-- Tchangsinaia_piscicula
| | |
| | | +-------- Unio_tumidus
| | +--+
| | | +----- Unio_nanus
+--+ +--+
| | +-- Unio_pictorum
| +--+
| +-- Unio_delphinus
|
| +-- Lanceolaria_grayii
| +--------------+
| | +-- Lanceolaria_lanceolata
| |
| | +-------- Cristaria_plicata
+--+ +-----+
| | | +----- Anemina_harlandi
| | +--+
| | | +-- Sinanodonta_lucida
+--+ +--+
| +-- Sinanodonta_woodiana
|
| +----------- Anodonta_anatina
+--+
| +-------- Platynaias_compressa
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_imbecillis
+--+
+-- Utterbackia_peninsularis
+----------------------- Pseudunio_marocanus
|
| +----- Quadrula_quadrula
| +--------------+
| | | +-- Venustaconcha_ellipsiformis
--+ | +--+
| | +-- Potamilus_alatus
| |
| | +----- Unio_tumidus
| | +-----+
| | | | +-- Unio_delphinus
+--+ | +--+
| +-----+ +-- Unio_nanus
| | |
| | | +-------- Sinanodonta_woodiana
| | +--+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Utterbackia_peninsularis
| | +--+
+--+ +-- Pyganodon_grandis
|
| +-- Chamberlainia_somsakpanhai
| +-----------+
| | +-- Sinohyriopsis_cumingii
| |
| | +-- Lens_contradens
| | +--+
+--+ | +-- Physunio_superbus
| +-----+
| | | +-- Hyriopsis_bialata
| | +--+
| | +-- Rectidens_sumatrensis
+--+
| +-- Pilsbryoconcha_exilis
| +-----+
| | +-- Pseudodon_vondembuschianus
| |
+--+ +-- Pronodularia_japanensis
| +--+
| | +-- Potomida_littoralis
+--+
| +-- Microcondylaea_bonellii
+--+
+-- Sinosolenaia_carinata