Publication Whelan et al. (2023): cladograms

publication summary

Complete Citation

Whelan, N.V., N.A. Johnson, A.S. Williams, M.A. Perkins, C.E. Beaver & J.W. Mays. 2023. Hidden in the hills: phylogeny of the freshwater mussel genus Alasmidonta (Bivalvia: Unionidae) and description of a new species. Zoological Journal of the Linnean Society 198(2): 650-676. Available electronically.

Cladogram Figures

(click the figure number to see the cladogram)

Cladogram Figure 2
                                +-------- Anemina_arcaeformis             
  +-----------------------------+                                         
  |                             |  +----- Sinanodonta_lucida              
  |                             +--+                                      
  |                                |  +-- Sinanodonta_woodiana            
  |                                +--+                                   
--+                                   +-- Sinanodonta_tumens              
  |                                                                       
  |  +----------------------------------- Anodonta_cygnea                 
  |  |                                                                    
  |  |                                +-- Anodonta_nuttalliana            
  |  |  +-----------------------------+                                   
  +--+  |                             +-- Pseudanodonta_complanata        
     |  |                                                                 
     |  |  +----------------------------- Anodonta_anatina                
     |  |  |                                                              
     |  |  |                       +----- Utterbackia_peninsularis__2     
     +--+  |                    +--+                                      
        |  |                    |  |  +-- Utterbackia_peninsularis__1     
        |  |                    |  +--+                                   
        |  |  +-----------------+     +-- Utterbackia_imbecillis          
        |  |  |                 |                                         
        |  |  |                 |  +----- Pyganodon_grandis__2            
        +--+  |                 +--+                                      
           |  |                    |  +-- Pyganodon_grandis__1            
           |  |                    +--+                                   
           |  |                       +-- Pyganodon_lacustris             
           |  |                                                           
           |  |                       +-- Utterbackiana_couperiana        
           |  |                    +--+                                   
           |  |                    |  +-- Utterbackiana_heardi            
           |  |              +-----+                                      
           |  |              |     |  +-- Utterbackiana_suborbiculata     
           +--+              |     +--+                                   
              |              |        +-- Utterbackiana_hartfieldorum     
              |  +-----------+                                            
              |  |           |     +----- Lasmigona_costata               
              |  |           |  +--+                                      
              |  |           |  |  |  +-- Pseudodontoideus_subvexus       
              |  |           |  |  +--+                                   
              |  |           +--+     +-- Pseudodontoideus_connasaugaensis
              |  |              |                                         
              |  |              |  +----- Pterosyna_complanata            
              |  |              +--+                                      
              |  |                 |  +-- Arcidens_confragosus            
              |  |                 +--+                                   
              |  |                    +-- Arcidens_wheeleri               
              +--+                                                        
                 |     +----------------- Sulcularia_viridis              
                 |     |                                                  
                 |  +--+              +-- Anodontoides_denigratus         
                 |  |  |  +-----------+                                   
                 |  |  |  |           +-- Anodontoides_ferussacianus      
                 |  |  +--+                                               
                 |  |     |  +----------- Strophitus_undulatus            
                 |  |     +--+                                            
                 |  |        |  +-------- Strophitus_pascagoulaensis__2   
                 |  |        +--+                                         
                 |  |           |  +----- Strophitus_radiatus             
                 +--+           +--+                                      
                    |              |  +-- Strophitus_pascagoulaensis__1   
                    |              +--+                                   
                    |                 +-- Strophitus_williamsi            
                    |                                                     
                    |              +----- Prolasmidonta_heterodon         
                    |  +-----------+                                      
                    |  |           |  +-- Platynaias_subviridis           
                    |  |           +--+                                   
                    |  |              +-- Platynaias_compressa            
                    +--+                                                  
                       |              +-- Alasminota_holstonia            
                       |  +-----------+                                   
                       |  |           +-- Alasminota_etowaensis           
                       |  |                                               
                       +--+           +-- Alasmidonta_raveneliana         
                          |  +--------+                                   
                          |  |        +-- Alasmidonta_marginata           
                          |  |                                            
                          +--+        +-- Alasmidonta_varicosa            
                             |  +-----+                                   
                             |  |     +-- Alasmidonta_uwharriensis        
                             +--+                                         
                                |  +----- Alasmidonta_undulata            
                                +--+                                      
                                   |  +-- Alasmidonta_triangulata         
                                   +--+                                   
                                      +-- Alasmidonta_arcula
Cladogram Figure 3
  +-------------------------------- Anodonta_anatina                
  |                                                                 
  |                             +-- Utterbackiana_heardi            
  |                          +--+                                   
  |                          |  +-- Utterbackiana_couperiana        
  |  +-----------------------+                                      
--+  |                       |  +-- Utterbackiana_hartfieldorum     
  |  |                       +--+                                   
  |  |                          +-- Utterbackiana_suborbiculata     
  |  |                                                              
  |  |                          +-- Utterbackia_peninsularis        
  |  |  +-----------------------+                                   
  +--+  |                       +-- Pyganodon_grandis               
     |  |                                                           
     |  |     +-------------------- Alasminota_etowaensis           
     |  |     |                                                     
     |  |     |              +----- Sulcularia_viridis              
     |  |  +--+  +-----------+                                      
     |  |  |  |  |           |  +-- Platynaias_subviridis           
     |  |  |  |  |           +--+                                   
     +--+  |  |  |              +-- Prolasmidonta_heterodon         
        |  |  +--+                                                  
        |  |     |              +-- Anodontoides_ferussacianus      
        |  |     |  +-----------+                                   
        |  |     |  |           +-- Anodontoides_denigratus         
        |  |     +--+                                               
        |  |        |  +----------- Strophitus_undulatus            
        |  |        +--+                                            
        |  |           |  +-------- Strophitus_pascagoulaensis__2   
        |  |           +--+                                         
        +--+              |  +----- Strophitus_pascagoulaensis__1   
           |              +--+                                      
           |                 |  +-- Strophitus_radiatus             
           |                 +--+                                   
           |                    +-- Strophitus_williamsi            
           |                                                        
           |                    +-- Arcidens_confragosus            
           |                 +--+                                   
           |                 |  +-- Arcidens_wheeleri               
           |        +--------+                                      
           |        |        |  +-- Pseudodontoideus_connasaugaensis
           |        |        +--+                                   
           |        |           +-- Pseudodontoideus_subvexus       
           +--------+                                               
                    |           +-- Alasmidonta_marginata           
                    |  +--------+                                   
                    |  |        +-- Alasmidonta_raveneliana         
                    |  |                                            
                    +--+        +-- Alasmidonta_varicosa            
                       |  +-----+                                   
                       |  |     +-- Alasmidonta_uwharriensis        
                       +--+                                         
                          |  +----- Alasmidonta_undulata            
                          +--+                                      
                             |  +-- Alasmidonta_triangulata         
                             +--+                                   
                                +-- Alasmidonta_arcula
Cladogram Figure 4
                          +-- Anodonta_anatina                   
  +-----------------------+                                      
  |                       +-- Anodonta_cygnea                    
  |                                                              
  |  +----------------------- Sulcularia_viridis                 
  |  |                                                           
  |  |                 +----- Utterbackiana_couperiana           
--+  |                 |                                         
  |  |     +-----------+  +-- Utterbackiana_heardi               
  |  |     |           |  |                                      
  |  |     |           +--+-- Utterbackiana_hartfieldorum        
  |  |     |              |                                      
  |  |  +--+              +-- Utterbackiana_suborbiculata        
  |  |  |  |                                                     
  +--+  |  |  +-------------- Alasminota_etowaensis              
     |  |  |  |                                                  
     |  |  +--+           +-- Arcidens_confragosus               
     |  |     |  +--------+                                      
     |  |     |  |        +-- Arcidens_wheeleri                  
     |  |     +--+                                               
     |  |        |  +-------- Pseudodontoideus_connasaugaensis__3
     |  |        +--+                                            
     |  |           |  +----- Pseudodontoideus_connasaugaensis__2
     |  |           +--+                                         
     +--+              |  +-- Pseudodontoideus_subvexus          
        |              +--+                                      
        |                 +-- Pseudodontoideus_connasaugaensis__1
        |                                                        
        |              +----- Alasmidonta_raveneliana            
        |              |                                         
        |              |  +-- Alasmidonta_uwharriensis           
        |  +-----------+--+                                      
        |  |           |  +-- Alasmidonta_varicosa               
        |  |           |                                         
        |  |           |  +-- Alasmidonta_arcula                 
        |  |           +--+                                      
        +--+              +-- Alasmidonta_triangulata            
           |                                                     
           |  +-------------- Platynaias_subviridis              
           |  |                                                  
           |  |     +-------- Strophitus_pascagoulaensis         
           |  |  +--+                                            
           +--+  |  |  +----- Strophitus_undulatus               
              |  |  +--+                                         
              |  |     |  +-- Strophitus_radiatus                
              |  |     +--+                                      
              +--+        +-- Strophitus_williamsi               
                 |                                               
                 |        +-- Pyganodon_grandis                  
                 |  +-----+                                      
                 |  |     +-- Pyganodon_cataracta                
                 +--+                                            
                    |  +----- Utterbackia_peninsularis           
                    +--+                                         
                       |  +-- Anodontoides_ferussacianus         
                       +--+                                      
                          +-- Anodontoides_denigratus