MUSSELpdb
Taxonomy
Species & Genus Richness of
freshwater mussels
by Taxonomy
Species & Genus Richness of
freshwater bivalves
by Family
Geography
Species Richness & Endemism of
freshwater mussels
by Geography
Data last updated
Mon 2 March 2026
Bivalvia | ... | Unionidae | Unioninae | Anodontini | Cristariina | Anemina harlandi
nominal species Anodon arcaeformis Heude, 1877
Anodon arcaeformis Heude, 1877 = Anemina harlandi
Heude, 1877. Conch. Fluv. Nanking (3): pl. 19, fig. 40.
Type(s): SYNTYPE MNHN_MP_3418, Kiang sou, Chine.
Taxonomic Opinions
= Anemina arcaeformis (Heude, 1877) — Dai et al., 2026: Table 1.
+ Anemina harlandi (Baird & H. Adams, 1867) — Chen et al., 2026: 54.
= Anemina arcaeformis (Heude, 1877) — Campbell & Lydeard, 2026: 293.
= Anemina arcaeformis (Heude, 1877) — Xiang et al., 2025: Tables 1-2.
= Anemina arcaeformis (Heude, 1877) — Dai et al., 2025: Tables 1-2.
= Anemina arcaeformis — Dai et al., 2025: Table 1.
= Anemina arcaeformis — Wu et al., 2025: 11, fig. 5, Tables S1, S5, S9.
= Anodonta arcaeformis — Perkins et al., 2025: Table S1.
= Anodonta arcaeformis — Perkins et al., 2025: Table S1.
= Anemina arcaeformis (Heude, 1877) — Chen et al., 2025: Table S3.
= Anemina arcaeformis — Hou et al., 2025: Table S3.
= Anemina arcaeformis — Wang et al., 2025: Table S2.
= Anemina arcaeformis (Heude, 1877) — Chen et al., 2025: Table 1.
= Anemina arcaeformis — Chen et al., 2025: Table 1.
= Anemina arcaeformis — Wu et al., 2024: 6.
= Anemina arcaeformis (Heude, 1877) — Dai et al., 2024: Table 1.
= Anemina arcaeformis — Wu et al., 2024: Table S3.
= Anemina arcaeformis (Heude, 1877) — Dai et al., 2024: 1194, Table 1.
= Anemina arcaeformis — Dai et al., 2024: Table 2.
= Anemina arcaeformis — Wu et al., 2024: Table S3.
= Anemina arcaeformis — Dai et al., 2024: Table 2.
= Anemina arcaeformis — Hou et al., 2024: Table 2.
= Anodonta arcaeformis — Galal-Khallaf et al., 2024: 22.
= Anemina arcaeformis (Heude, 1877) — Konopleva et al., 2023: Table S1.
= Anodonta arcaeformis — Whelan et al., 2023: 13, Table S1.
= Anemina arcaeformis (Heude, 1877) — Dai et al., 2023: 157, Table 2.
= Anemina arcaeformis — Bogan et al., 2023: Table 1.
= Anodonta arcaeformis — Whelan et al., 2023: Table S1.
= Anemina arcaeformis (Heude, 1877) — Chen et al., 2023: 133, Table 1.
= Anemina arcaeformis (Heude, 1877) — Wu et al., 2023: Tables 2-3.
= Anemina arcaeformis (Heude, 1877) — Wu et al., 2022: Table S2.
= Anemina arcaeformis — Liu et al., 2022: Table 1.
= Anodonta arcaeformis — Pavan-Kumar et al., 2022: Figure S5.
= Anemina arcaeformis — Hata et al., 2022: Table S1.
= Anemina arcaeformis (Heude, 1877) — Graf & Cummings, 2021: 16.
= Anemina arcaeformis — Zieritz et al., 2021: 1092.
= Anodonta arcaeformis — Bauer et al., 2021: 6.
= Anemina arcaeformis (Heude, 1877) — Wu et al., 2021: 1799, fig. 7E, attached Tables 1-2.
= Anemina arcaeformis (Heude, 1877) — Lopes-Lima et al., 2020: 7, Tables 1, 3, S3.
= Anemina arcaeformis — Riccardi et al., 2020: 750.
= Anemina arcaeformis — Liu et al., 2020: [6].
= Anodonta arcaeformis — Fukata & Iigo, 2020: 1216.
= Anemina arcaeformis — Wu et al., 2020: 196.
= Anemina arcaeformis — Sano et al., 2020: 101.
= Anemina arcaeformis (Heude, 1877) — Wu et al., 2020: 54.
= Anemina arcaeformis (Heude, 1877) — Bolotov et al., 2020: Table S1.
= Anemina arcaeformis — Sano et al., 2020: 101.
= Anemina arcaeformis (Heude, 1877) — Huang et al., 2019: Table S4.
= Anemina arcaeformis (Heude, 1877) — Wu et al., 2019: 29.
= Anemina arcaeformis — Froufe et al., 2019: 185.
= Anemina arcaeformis (Heude, 1877) — Gomes-dos-Santos et al., 2019: 424.
= Anemina arcaeformis (Heude, 1877) — Zieritz et al., 2018: online Table 1.
= Anemina arcaeformis — Huang et al., 2018: 304.
= Anemina arcaeformis (Heude, 1877) — Wu et al., 2018: 4.
= Anodonta arcaeformis — Wang et al., 2018: 611.
= Anodonta arcaeformis — Chase et al., 2018: Additional File 1.
= Anodonta arcaeformis — Aunins et al., 2018: 1125.
= Anodonta arcaeformis — Wang et al., 2018: 140.
= Anemina arcaeformis (Heude, 1877) — Bespalaya et al., 2018: 97.
= Anodonta arcaeformis — Cha et al., 2018: 946.
= Anemina arcaeformis — Wu et al., 2018: 732.
= Anemina cf. arcaeformis (Heude, 1877) — Kondakov et al., 2018: fig. 5.
= Anemina arcaeformis (Heude, 1877) — Lopes-Lima et al., 2017: 177.
= Anemina arcaeformis (Heude, 1877) — Sayenko et al., 2017: 267.
= Anodonta arcaeformis — Zhou et al., 2017: 235.
= Anemina arcaeformis — Lopes-Lima et al., 2017: 250.
= Anemina arcaeformis Heude, 1977 [sic] — Sano et al., 2017: 494.
= Anemina arcaeformis — Froufe et al., 2017: 3265.
= Anemina arcaeformis (Heude, 1877) — Vikhrev et al., 2017: 5, Table 1.
= Anodonta arcaeformis — Soroka & Burzynski, 2017: 863.
= Anemina arcaeformis (Heude, 1877) — Bogatov & Prozorova, 2017: 924.
= Anemina arcaeformis (Heude, 1877) — Liu et al., 2017: Table S4.
= Anodonta arcaeformis (Heude) — An et al., 2016: 1552.
= Anodonta arcaeformis — Xue et al., 2016: 2699.
= Anodonta arcaeformis — Wu et al., 2016: 4521.
= Anemina arcaeformis — Fonseca et al., 2016: 955.
= Anodonta arcaeformis — Feng et al., 2016: 445.
= Anemina (s.s.) arcaeformis (Heude, 1877) — Vinarski & Kantor, 2016: 45.
= Anodonta arcaeformis — Soliman et al., 2016: 122.
= Anodonta arcaeformis — Ouyang et al., 2015: Table 1.
= Anemina arcaeformis (Heude, 1877) — He & Zhuang, 2013: 29, figs. 56-61.
= Anodonta arcaeformis — Zhang et al., 2013: 376.
= Anemina arcaeformis (Heude, 1877) — IUCN, 2012: [web].
= Anodonta arcaeformis — Xiong et al., 2012: 128, Table 2.
= Anemina arcaeformis (Heude, 1877) — Zieritz et al., 2012: 742, 745.
= Anodonta arcaeformis — Ouyang et al., 2011: 544.
= Anemina arcaeformis Heude, 1877 — Bogan, 2010: [2].
= Anodonta arcaeformis — Soroka, 2010: 178.
= Anemina arcaeformis (Heude, 1877) — Kondo, 2008: 42, pl. 5, fig. d.
= Anodonta arcaeformis (Heude, 1877) — Graf, 2007: 87.
= Anemina arcaeformis (Heude, 1877) — Graf & Cummings, 2007: 304.
= Buldowskia arcaeformis (Heude, 1877) — Prozorova et al., 2005: 55.
= Anemina arcaeformis — Sayenko et al., 2005: 121.
= Anodonta araeformis [sic] (Hende, 1877) [sic] — Bába, 2000: 133, fig. 2.
= Anodonta arcaeformis — Wu et al., 1999: 140.
= Anemina arcaeformis (Heude, 1877) — Korniushin, 1998: 78.
= Anemina arcaeformis (Heude, 1877) — Habe, 1991: 5.
= Anodonta arcaeformis (Heude) — Liu et al., 1980: 366.
= Anodonta arcaeformis arcaeformis (Heude, 1877) — Liu et al., 1979: 112, fig. 101.
= Anodonta (Anemina) arcaeformis Heude, 1877 — Habe, 1977: 120.
= Anodonta (Anemina) arcaeformis (Heude, 1877) — Haas, 1969: 359.
= Anodonta (Haasiella) arcaeformis (Heude, 1877) — Haas, 1969: N444, fig. D38:1.
= Anodonta (Haasiella) arcaeformis (Heude, 1877) — Zhadin, 1965: 260, fig. 279.
= Anodonta (Haasiella) arcaeformis (Heude, 1877) — Zhadin, 1938: 147, sp. 11, fig. 56.
= Anodonta arcaeformis (Heude) — Thiele, 1934: 832, [1267].
= Anodonta arcaeformis (Heude, 1877) — Simpson, 1914: 417.
= Anodonta arcaeformis Heude — Martens, 1905: 63.
= Anodonta arcaeformis (Heude, 1877) — Simpson, 1900: 641.
= Anodonta arcaeformis Heude — Neumayr, 1899: 647.
= Anodon arcaeformis Heude — Heude, 1877: pl. 19, sp. 40.
