Dai, Y.-T., Z.-G. Chen, C.-L. Hu, P.-F. Ning, S. Ouyang et al. 2024. Taxonomic reassessment of Scabies (Bivalvia: Unionidae) species in China based on multilocus and mitogenomic phylogenetic analyses. Invertebrate Systematics 38(6): 1-18. Available electronically.
(click the figure number to see the cladogram)
+----------------------------- Scabies_scobinatus
|
--+ +-------------------------- Unio_crassus
| |
| | +----- Aculamprotula_scripta
+--+ +-----------------+
| | | +-- Anodonta_anatina
| | +--+
+--+ +-- Lepidodesma_languilati
|
| +-------------------- Middendorffinaia_mongolica
| |
+--+ +----------------- Inversiunio_jokohamensis
| |
+--+ +-------------- Cuneopsis_celtiformis
| |
+--+ +----------- Nodularia_breviconcha
| |
+--+ +-- Nodularia_fusiformans
| +-----+
| | +-- Nodularia_huana
+--+
| +----- Nodularia_nuxpersicae
+--+
| +-- Nodularia_douglasiae
+--+
+-- Nodularia_nipponensis
+-- Margaritifera_margaritifera
+--------------------------+
| +-- Gibbosula_laosensis
|
| +----- Trapezidens_exolescens
--+ +--------------------+
| | | +-- Scabies_scobinatus
| | +--+
| | +-- Scabies_crispata
| |
+--+ +----------------------- Acuticosta_chinensis
| |
| +----------------------- Anemina_arcaeformis
| |
| +----------------------- Anodonta_anatina
+--+
| +-------------------- Lanceolaria_grayii
| |
| | +-- Aculamprotula_scripta
| +-----------------+
+--+ +-- Lepidodesma_languilati
|
| +----------------- Unio_crassus
| |
+--+ +-------------- Middendorffinaia_mongolica
| |
| | +----------- Cuneopsis_celtiformis
+--+ |
| | +-- Inversiunio_jokohamensis
| +--------+
| | +-- Schistodesmus_xinyuensis
+--+
| +-- Nodularia_huana
| +-----+
| | +-- Nodularia_breviconcha
| |
+--+ +-- Nodularia_nuxpersicae
| +--+
| | +-- Nodularia_nipponensis
+--+
| +-- Nodularia_douglasiae
+--+
+-- Nodularia_dualobtusa
+-- Margaritifera_margaritifera
+--------------------------------+
| +-- Gibbosula_laosensis
|
--+ +----- Trapezidens_exolescens
| +--------------------------+
| | | +-- Scabies_scobinatus
| | +--+
+--+ +-- Scabies_crispata
|
| +----------------------------- Anodonta_anatina
| |
+--+ +-- Acuticosta_chinensis
| +-----------------------+
| | +-- Anemina_arcaeformis
+--+
| +----------------------- Lepidodesma_languilati
| |
+--+ +-------------------- Lanceolaria_grayii
| |
+--+ +----------------- Aculamprotula_scripta
| |
+--+ +-------------- Unio_crassus
| |
| | +----------- Middendorffinaia_mongolica
+--+ |
| | +----- Cuneopsis_celtiformis
| | +--+
+--+ | | +-- Inversiunio_jokohamensis
| | +--+
| | +-- Schistodesmus_xinyuensis
| |
+--+ +-- Nodularia_nuxpersicae
| +--+
| | +-- Nodularia_nipponensis
| |
| | +-- Nodularia_huana
+--+--+
| +-- Nodularia_breviconcha
|
| +-- Nodularia_douglasiae
+--+
+-- Nodularia_dualobtusa
+-------- Quadrula_quadrula
+--------------------------+
| | +----- Venustaconcha_ellipsiformis
| +--+
| | +-- Lampsilis_powellii
| +--+
| +-- Lampsilis_siliquoidea
|
| +-- Chamberlainia_somsakpanhai
| +-----------+
| | +-- Sinohyriopsis_cumingii
| |
--+ | +-------- Potomida_littoralis
| +-----------------+ +--+
| | | | | +----- Pronodularia_japanensis
| | | | +--+
| | | | | +-- Lamprotula_caveata
| | +--+ +--+
| | | +-- Lamprotula_leaii
| | |
| | | +-------- Pilsbryoconcha_exilis
| | +--+
| | | +----- Parvasolenaia_rivularis
+--+ +--+
| | +-- Sinosolenaia_carinata
| +--+
| +-- Sinosolenaia_oleivora
|
| +----------------------------- Lepidodesma_languilati
| |
| | +----------- Lanceolaria_lanceolata
| | +--------------+
| | | | +-------- Acuticosta_chinensis
| | | +--+
+--+ | | +----- Sinanodonta_woodiana
| | +--+
| | | +-- Pyganodon_grandis
| | +--+
| | +-- Utterbackia_peninsularis
| |
+--+ +-------- Aculamprotula_tortuosa
| |
| +--------------+ +-- Aculamprotula_polysticta
| | | +--+
| | | | +-- Aculamprotula_scripta
| | +--+
| | | +-- Aculamprotula_coreana
| | +--+
| | +-- Aculamprotula_tientsinensis
| |
+--+ +-------------- Unio_tumidus
| +-----+
| | | +----------- Unio_nanus
| | +--+
| | | +-------- Unio_delphinus
| | +--+
| | | +----- Unio_pictorum
| | +--+
+--+ | +-- Unio_elongatulus
| +--+
| +-- Unio_mancus
|
| +----------------- Pseudocuneopsis_capitata
| |
| | +----------- Nodularia_breviconcha
| | +--+
+--+ | | +-------- Nodularia_fusiformans
| | +--+
| | | +----- Nodularia_douglasiae
| | +--+
| | | +-- Nodularia_nuxpersicae
+--+ +--+
| +-- Nodularia_nipponensis
|
| +-- Schistodesmus_lampreyanus
| +-----+
| | +-- Schistodesmus_xinyuensis
+-----+
| +----- Tchangsinaia_piscicula
+--+
| +-- Cuneopsis_heudei
+--+
+-- Cuneopsis_celtiformis