Wu, R., L. Liu, L. Zhang, A.E. Bogan, G. Niu et al. 2024. Taxonomic revision of two species in the genus Ptychorhynchus Simpson, 1900 (Bivalvia: Unionidae: Gonideinae), with description of a new species. Invertebrate Systematics 38, IS24014: 1-17. Available electronically.
(click the figure number to see the cladogram)
+-- Cosmopseudodon_resupinatus
+--+
| +-- Ptychorhynchus_pfisteri
+-----+
| | +-- Postolata_guangxiensis
| +--+
--+ +-- Obovalis_omiensis
|
| +-------- Nodularia_breviconcha
| |
+--+ +-- Nodularia_douglasiae
| +--+
| | +-- Nodularia_dualobtusa
+--+
| +-- Nodularia_nipponensis
+--+
+-- Nodularia_nuxpersicae
+-- Margaritifera_dahurica
+--------------------------------+
| +-- Margaritifera_margaritifera
|
| +-------------------------------- Chamberlainia_somsakpanhai
| |
| | +-- Khairuloconcha_sahanae
| | +--------------+
--+ | | +-- Khairuloconcha_lunbawangorum
| | |
| | | +-------- Ensidens_sagittarius
| | +-----------+ +-----+
| | | | | | +----- Rectidens_sumatrensis
| | | | | +--+
| | | | | | +-- Hyriopsis_bialata
+--+ | | | +--+
| | +--+ +-- Hyriopsis_desowitzi
| | |
| | | +-- Physunio_superbus
| | | +--------+
| | | | +-- Trapezoideus_foliaceus
| | | |
| | +--+ +-- Lens_contradens
| | | +-----+
| | | | +-- Lens_eximius
| | +--+
+--+ | +----- Yaukthwa_elongatula
| +--+
| | +-- Yaukthwa_inlenensis
| +--+
| +-- Yaukthwa_paiensis
|
| +----- Potomida_littoralis
| +--------------------+
| | | +-- Lamprotula_leaii
| | +--+
| | +-- Lamprotula_caveata
| |
| | +-------------------- Sinosolenaia_carinata
| | |
| | +--+ +-- Gonidea_angulata
+--+ | | +--------------+
| | | | +-- Microcondylaea_bonellii
| | +--+
| | | +-------------- Koreosolenaia_sitgyensis
| | +--+
| | | +----------- Parvasolenaia_rivularis
| | +--+
| | | +-------- Postolata_guangxiensis
+--+ +--+
| | +----- Cosmopseudodon_resupinatus
| +--+
| | +-- Ptychorhynchus_pfisteri
| +--+
| +-- Obovalis_omiensis
|
| +----------------- Schepmania_nieuwenhuisi
| |
+-----+ +-- Indopseudodon_bogani
| +-----------+
| | +-- Indopseudodon_kayinensis
+--+
| +----------- Pilsbryoconcha_exilis
| |
+--+ +-- Pseudodon_mekongi
| +-----+
| | +-- Pseudodon_vondembuschianus
+--+
| +----- Thaiconcha_callifera
+--+
| +-- Bineurus_anodontinum
+--+
+-- Bineurus_loeiensis
+-- Margaritifera_dahurica
+--------------------------------+
| +-- Margaritifera_margaritifera
|
| +-- Quadrula_quadrula
| +-----------------+
| | +-- Uniomerus_tetralasmus
| |
| +-----------+ +-- Elliptio_complanata
| | | +--------------+
| | | | +-- Pleurobema_oviforme
| | +--+
| | | +-------------- Popenaias_popeii
| | +--+
--+ | | +----------- Toxolasma_parvum
| | +--+
| | | +-------- Potamilus_alatus
| | +--+
| | | +----- Leaunio_lienosus
| | +--+
| | | +-- Lampsilis_cardium
| | +--+
| | +-- Venustaconcha_ellipsiformis
| |
| | +-------------------------- Lamellidens_marginalis
| | |
| | | +----------------------- Lepidodesma_languilati
| | | |
+--+ +--+ | +-------------- Lanceolaria_lanceolata
| | | | |
| | | | | +----- Anodonta_cygnea
| | | | +-----+ +-----+
| | +--+ | | | | +-- Alasmidonta_varicosa
| | | | | | +--+
| | | | +--+ +-- Utterbackia_imbecillis
| | | | |
| | | | | +-------- Cristaria_plicata
| | | | +--+
| | +--+ | +----- Anemina_harlandi
| | | +--+
| | | | +-- Beringiana_fukuharai
| | | +--+
| | | +-- Sinanodonta_lucida
| | |
| | | +----------------- Aculamprotula_tortuosa
| | | |
| | +--+ +-------------- Unio_tumidus
| | | |
| | +--+ +----------- Pseudocuneopsis_sichuanensis
+--+ | |
| | | +----- Nodularia_breviconcha
| +--+ +--+
| | | | +-- Nodularia_douglasiae
| | | +--+
| +--+ +-- Nodularia_nipponensis
| |
| | +----- Tchangsinaia_piscicula
| +--+
| | +-- Cuneopsis_heudei
| +--+
| +-- Schistodesmus_xinyuensis
|
| +----- Chamberlainia_somsakpanhai
| +-----------------+
| | | +-- Sinohyriopsis_cumingii
| | +--+
| | +-- Sinohyriopsis_schlegelii
| |
| | +-- Hyriopsis_bialata
| | +--+
+-----+ | +-- Rectidens_sumatrensis
| +--------------+
| | | +-- Lens_contradens
| | +--+
| | +-- Physunio_superbus
| |
| | +----------- Potomida_littoralis
+--+ +-----+
| | | +-------- Pronodularia_japanensis
| | +--+
| | | +----- Lamprotula_leaii
| | +--+
| | | +-- Lamprotula_caveata
| | +--+
+--+ +-- Lamprotula_cornuumlunae
|
| +-- Pseudodon_vondembuschianus
| +-----------+
| | +-- Pilsbryoconcha_exilis
| |
| | +-------- Parvasolenaia_rivularis
+--+ +--+
| | | +----- Ptychorhynchus_pfisteri
| | +--+
| | | +-- Cosmopseudodon_resupinatus
+--+ +--+
| +-- Postolata_guangxiensis
|
| +-------- Microcondylaea_bonellii
+--+
| +----- Sinosolenaia_carinata
+--+
| +-- Sinosolenaia_iridinea
+--+
+-- Sinosolenaia_oleivora