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Data last updated
Thu 18 Dec 2025
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Publication Hou et al. (2024): cladogramspublication summary
Complete CitationHou, K., L. Zhang, L. Liu, X. Liu, J. Jia & R. Wu. 2024. Reappraising the species validity of Cuneopsis szechenyii comb. nov. (Neumayr, 1899) and revising the taxonomy of Middendorffinaia mongolica (Middendorff, 1851). Invertebrate Systematics 38, IS24071: 1-14. Available electronically.
Cladogram Figures(click the figure number to see the cladogram) Cladogram Figure 1
+-- Lamprotula_caveata
+-----------------+
| +-- Lamprotula_leaii
|
| +----------- Tchangsinaia_piscicula
| |
--+ +-----+ +-- Cuneopsis_kiangsiensis
| | | +-----+
| | | | +-- Cuneopsis_celtiformis
| | +--+
| | | +----- Cuneopsis_szechenyii
| | +--+
+--+ | +-- Cuneopsis_heudei
| +--+
| +-- Cuneopsis_demangei
|
| +-------------- Cuneopsis_rufescens
| |
+--+ +----------- Middendorffinaia_mongolica
| |
+--+ +-------- Pseudocuneopsis_capitata
| |
+--+ +-- Pseudocuneopsis_sichuanensis
| +--+
| | +-- Pseudocuneopsis_yemaoi
+--+
| +-- Pseudocuneopsis_wuana
+--+
+-- Pseudocuneopsis_yangshuoensis
Cladogram Figure 4
+-- Margaritifera_dahurica
+-----------------------------------------------+
| +-- Margaritifera_margaritifera
|
| +-- Lampsilis_cardium
| +--------------------------------------------+
| | +-- Quadrula_quadrula
| |
--+ | +-- Lepidodesma_aligera
| | +-----------------------------------------+
| | | +-- Lepidodesma_languilati
| | |
| | | +----- Lanceolaria_gladiola
| | | +-----------------+
| | | | | +-- Lanceolaria_grayii
| | | | +--+
+--+ | | +-- Lanceolaria_lanceolata
| | |
| | | +-------------- Cristaria_plicata
| | | +-----+
| | +-----------------+ | | +----------- Anemina_arcaeformis
| | | | | +--+
| | | | | | +-------- Beringiana_fukuharai
| | | | | +--+
| | | | | | +----- Sinanodonta_lucida
| | | | | +--+
| | | | | | +-- Sinanodonta_tumens
+--+ | +--+ +--+
| | | +-- Sinanodonta_woodiana
| | |
| | | +-------- Pseudanodonta_complanata
| | | +--------+
| | | | | +----- Anodonta_anatina
| | | | +--+
| | | | | +-- Anodonta_cygnea
| | | | +--+
| | +--+ +-- Anodonta_exulcerata
| | |
| | | +-------------- Anodonta_nuttalliana
| | | |
| | | | +----- Pyganodon_grandis
| | +--+ +-----+
| | | | | +-- Utterbackia_imbecillis
| | | | +--+
| | +--+ +-- Utterbackia_peninsularis
| | |
+--+ | +-------- Pterosyna_complanata
| +--+
| | +----- Prolasmidonta_heterodon
| +--+
| | +-- Alasmidonta_varicosa
| +--+
| +-- Platynaias_compressa
|
| +-------- Aculamprotula_tortuosa
| |
| +-----------------------------+ +-- Aculamprotula_coreana
| | | +--+
| | | | +-- Aculamprotula_tientsinensis
| | +--+
| | | +-- Aculamprotula_polysticta
| | +--+
| | +-- Aculamprotula_scripta
| |
| | +-- Unio_gibbus
| | +-----------------------------+
| | | +-- Unio_tumidus
| | |
| | | +----------- Unio_terminalis
| | +--+ +-----------------+
| | | | | | +-------- Unio_elongatulus
| | | | | +--+
+--+ | | | | +----- Unio_pictorum
| | | | +--+
| | +--+ | +-- Unio_delphinus
| | | +--+
| | | +-- Unio_mancus
| | |
| | | +-------------------------- Unio_tumidiformis
| | | |
| | +--+ +----------------------- Unio_desectus
| | | |
| | +--+ +-------------------- Unio_sesirmensis
| | | |
| | +--+ +----------------- Unio_ionicus
| | | |
| | +--+ +-------------- Unio_damascensis
| | | |
| | | | +-- Unio_crassus
+--+ +--+ +--------+
| | | +-- Unio_gontierii
| | |
| +--+ +-- Unio_bruguierianus
| | +-----+
| | | +-- Unio_mardinensis
| +--+
| | +----- Unio_vicarius
| +--+
| | +-- Unio_carneus
| +--+
| +-- Unio_nanus
|
| +-------------------------- Middendorffinaia_mongolica
| |
| | +-------- Pseudocuneopsis_capitata
| | |
| | +--------------+ +-- Pseudocuneopsis_sichuanensis
| | | | +--+
| | | | | +-- Pseudocuneopsis_yemaoi
+--------+ | +--+
| | | +-- Pseudocuneopsis_wuana
| | +--+
| | +-- Pseudocuneopsis_yangshuoensis
| |
| | +-- Schistodesmus_xinyuensis
| | +--------------+
+--+ | +-- Schistodesmus_spinosus
| +--+
| | | +-------------- Tchangsinaia_piscicula
| | | |
| | +--+ +----------- Cuneopsis_rufescens
| | | |
| | +--+ +-- Cuneopsis_celtiformis
| | | +-----+
| | | | +-- Cuneopsis_kiangsiensis
| | +--+
+--+ | +----- Cuneopsis_szechenyii
| +--+
| | +-- Cuneopsis_demangei
| +--+
| +-- Cuneopsis_heudei
|
| +----- Nodularia_dorri
| +-----------+
| | | +-- Nodularia_hanensis
| | +--+
+--+ +-- Nodularia_micheloti
|
| +-------------- Nodularia_breviconcha
| |
+--+ +----------- Nodularia_huana
| |
+--+ +-------- Nodularia_fusiformans
| |
+--+ +-- Nodularia_douglasiae
| +--+
| | +-- Nodularia_dualobtusa
+--+
| +-- Nodularia_nipponensis
+--+
+-- Nodularia_nuxpersicae
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