MUSSELpdb
Taxonomy
Species & Genus Richness of
freshwater mussels
by Taxonomy
Species & Genus Richness of
freshwater bivalves
by Family
Geography
Species Richness & Endemism of
freshwater mussels
by Geography
Data last updated
Thu 19 March 2026
Publication Wang et al. (2025): cladograms
Complete Citation
Wang, X., F. Nan, X. Liu & R. Wu. 2025. Pseudocuneopsis heqing sp. nov. (Bivalvia, Unionidae) from karst rivers of Guizhou, China: integrative species delimitation and mitogenomic phylogeny. Zoosystematics and Evolution 101(4): 2181-2190. Available electronically.
Cladogram Figures
(click the figure number to see the cladogram)
Cladogram Figure 3B
+-- Gibbosula_rochechouartii
+--------------------------------------+
| +-- Margaritifera_margaritifera
|
| +-- Lepidodesma_aligera
| +-----------------------------------+
| | +-- Lepidodesma_languilati
| |
| | +-- Acuticosta_chinensis
| | +--------------------------+
| | | +-- Acuticosta_ovata
| | |
| | | +----- Anodonta_anatina
--+ | | +-----------+
| | | | | +-- Anodonta_exulcerata
| | | | +--+
| | | | +-- Anodonta_cygnea
| | | +--------+
| | +-----+ | | +-------------- Pseudanodonta_complanata
| | | | | | |
| | | | | | | +----- Pyganodon_grandis
| | | | | +--+ +-----+
| | | | | | | | +-- Utterbackia_imbecillis
| | | | | | | +--+
| | | | | +--+ +-- Utterbackia_peninsularis
+--+ | | | |
| | | | | +-------- Pterosyna_complanata
| | | | +--+
| | +--+ | +----- Prolasmidonta_heterodon
| | | +--+
| | | | +-- Alasmidonta_varicosa
| | | +--+
| | | +-- Platynaias_compressa
| | |
| | | +----- Cristaria_liboi
| | | +-----------------+
| | | | | +-- Cristaria_plicata
| | | | +--+
| | | | +-- Cristaria_truncata
| | | |
| | +--+ +----- Amuranodonta_kijaensis
| | | +--------------+
| | | | | +-- Anemina_harlandi
| | | | +--+
| | | | +-- Buldowskia_shadini
+--+ +--+
| | +-- Beringiana_fukuharai
| | +--------------+
| | | +-- Pletholophus_guangzhouensis
| +--+
| | +-------------- Acudonta_baitiaoensis
| | |
| +--+ +----------- Sinanodonta_lucida
| | |
| +--+ +-- Sinanodonta_schrenkii
| | +-----+
| | | +-- Sinanodonta_tumens
| +--+
| | +----- Sinanodonta_woodiana
| +--+
| | +-- Sinanodonta_lauta
| +--+
| +-- Sinanodonta_pacifica
|
| +-------- Scriptolamprotula_tortuosa
| |
| +-----------------------+ +-- Scriptolamprotula_coreana
| | | +--+
| | | | +-- Scriptolamprotula_tientsinensis
| | +--+
| | | +-- Scriptolamprotula_polysticta
| | +--+
| | +-- Scriptolamprotula_scripta
+--+
| +-------- Unio_tumidus
| +--------------------+
| | | +----- Unio_elongatulus
| | +--+
| | | +-- Unio_carneus
| | +--+
| | +-- Unio_nanus
+--+
| +-------------------------- Middendorffinaia_mongolica
| |
| | +-------------- Pseudocuneopsis_capitata
| | |
| | +--------+ +----------- Pseudocuneopsis_perflora
| | | | |
+--+ | +--+ +-------- Pseudocuneopsis_liuovatus
| | | |
| | +--+ +-- Pseudocuneopsis_sichuanensis
| | | +--+
| | | | +-- Pseudocuneopsis_yemaoi
| | +--+
| | | +-- Pseudocuneopsis_wuana
+--+ +--+
| +-- Pseudocuneopsis_yangshuoensis
|
| +-------- Nodularia_hanensis
| +-----------+
| | | +----- Nodularia_fusiformans
| | +--+
| | | +-- Nodularia_douglasiae
| | +--+
| | +-- Nodularia_guiensis
+--+
| +----------- Schistodesmus_spinosus
| +-----+
| | | +-------- Schistodesmus_tongpenensis
| | +--+
| | | +----- Schistodesmus_xinyuensis
| | +--+
+--+ | +-- Schistodesmus_lampreyanus
| +--+
| +-- Schistodesmus_luqiaoensis
|
| +-------------- Tchangsinaia_piscicula
| |
+--+ +----------- Cuneopsis_rufescens
| |
+--+ +-- Cuneopsis_heudei
| +-----+
| | +-- Cuneopsis_szechenyii
+--+
| +----- Cuneopsis_celtiformis
+--+
| +-- Cuneopsis_celtiosimilis
+--+
+-- Cuneopsis_kiangsiensis
