MUSSELpdb
Taxonomy
Species & Genus Richness of
freshwater mussels
by Taxonomy
Species & Genus Richness of
freshwater bivalves
by Family
Geography
Species Richness & Endemism of
freshwater mussels
by Geography
Data last updated
Mon 2 March 2026
Bivalvia | Unionida | Hyrioidea | Hyriidae | Hyriinae | Rhipidodontini | Diplodon
species Diplodon gratus (Lea, 1860): synonymy
Synonymy
Unio gratus Lea, 1860
Lea, 1860. Proc. Acad. Nat. Sci. 12: 91.
Lea, 1863. Jour. Acad. Nat. Sci. 5 [O. 10]: 382 [18], pl. 43, fig. 290.
Type(s): HOLOTYPE USNM_85784, Uruguay R., S. Amercia.
Diplodon (Cyclomya s.s.) gratus — Simpson (1900)
Diplodon (Cyclomya s.s.) fontainianus var. gratus — Simpson (1914)
Diplodon (s.s.) gratus — Ortmann (1921), Morretes (1949)
Diplodon (Cyclomya) paranensis funebralis — Haas (1931)
Diplodon (Rhipidodonta) charruanus — Bonetto (1964), Bonetto (1965), Figueiras (1965), Bonetto & Mansur (1970)
Diplodon rotundus gratus — Parodiz (1968), Mansur (1970)
Diplodon (Rhipidodonta) paranensis funebralis — Haas (1969)
Rhipidodonta grata — Simone (2006), Graf & Cummings (2007), Bogan (2010)
Diplodon grata — ITIS (2013)
Diplodon gratus — Miyahira et al. (2019), Graf & Cummings (2021), Campbell & Lydeard (2026)
Unio rufofuscus Lea, 1865
Lea, 1865. Proc. Acad. Nat. Sci. 17: 76.
Lea, 1868. Jour. Acad. Nat. Sci. 6 [O. 12]: 282, pl. 39, fig. 96.
Type(s): HOLOTYPE USNM_84101, [no locality available - South America].
Diplodon (s.s.) rufofuscus — Simpson (1900), Simpson (1914)
Diplodon (Cyclomya) fontainianus — Haas (1931)
Diplodon (s.s.) granosus multistriatus — Bonetto (1964)
Diplodon rotundus gratus — Parodiz (1968), Mansur (1970)
Diplodon (Rhipidodonta) fontainianus — Haas (1969)
Diplodon rufofuscus — Miyahira et al. (2019)
Diplodon (Cyclomya s.s.) fontainianus var. deceptus Simpson, 1914
Simpson, 1914. Descr. Cat. Naiades: 1281.
(intro'd as intra-sp. of Unio fontainiana d’Orbigny, 1835)
Diplodon (Cyclomya s.s.) fontainianus var. deceptus — Simpson (1914)
Diplodon (s.s.) deceptus — Ortmann (1921), Morretes (1949)
Diplodon (Cyclomya) paranensis funebralis — Haas (1931)
Diplodon (s.s.) delodontus expansus — Bonetto (1964)
Diplodon (Rhipidodonta) funebralis — Figueiras (1965)
Diplodon rotundus gratus — Parodiz (1968), Mansur (1970)
Diplodon (Rhipidodonta) paranensis funebralis — Haas (1969)
Diplodon (s.s.) delodontus wimanii [sic] — Bonetto & Mansur (1970)
Diplodon deceptus — Mansur et al. (1988), Bogan & Hoeh (2000), Hoeh et al. (2001), Hoeh et al. (2002), Baker et al. (2003), Roe & Hoeh (2003), Baker et al. (2004), Walker et al. (2006), Hoeh et al. (2009), Marshall et al. (2014), Zieritz et al. (2015), Santos-Neto et al. (2016)
Diplodon rotundus — Graf & Cummings (2006)
Rhipidodonta grata — Simone (2006)
Diplodon (Rhipidodonta) deceptus — Mansur & Pereira (2006), Pereira et al. (2014)
Rhipidodonta decepta — Mansur et al. (2020)
Unio gratus Ihering, 1893
‘Lea’ Ihering, 1893. Arch. Naturg. 59(1): 92.
(reference to Unio gratus Lea, 1860)
Diplodon (s.s.) deceptus — Ortmann (1921)
Diplodon rotundus gratus — Parodiz (1968), Mansur (1970)
Taxonomic History (since 1900)
Diplodon (Cyclomya s.s.) gratus — Simpson (1900)
Diplodon (s.s.) rufofuscus — Simpson (1900), Simpson (1914)
Diplodon (Cyclomya s.s.) fontainianus var. gratus — Simpson (1914)
Diplodon (Cyclomya s.s.) fontainianus var. deceptus — Simpson (1914)
Diplodon (s.s.) gratus — Ortmann (1921), Morretes (1949)
Diplodon (s.s.) deceptus — Ortmann (1921), Morretes (1949)
Diplodon (Cyclomya) paranensis funebralis — Haas (1931)
Diplodon (Cyclomya) fontainianus — Haas (1931)
Diplodon (Rhipidodonta) charruanus — Bonetto (1964), Bonetto (1965), Figueiras (1965), Bonetto & Mansur (1970)
Diplodon (s.s.) granosus multistriatus — Bonetto (1964)
Diplodon (s.s.) delodontus expansus — Bonetto (1964)
Diplodon (Rhipidodonta) funebralis — Figueiras (1965)
Diplodon rotundus gratus — Parodiz (1968), Mansur (1970)
Diplodon (Rhipidodonta) paranensis funebralis — Haas (1969)
Diplodon (Rhipidodonta) fontainianus — Haas (1969)
Diplodon (s.s.) delodontus wimanii [sic] — Bonetto & Mansur (1970)
Diplodon deceptus — Mansur et al. (1988), Bogan & Hoeh (2000), Hoeh et al. (2001), Hoeh et al. (2002), Baker et al. (2003), Roe & Hoeh (2003), Baker et al. (2004), Walker et al. (2006), Hoeh et al. (2009), Marshall et al. (2014), Zieritz et al. (2015), Santos-Neto et al. (2016)
Rhipidodonta grata — Simone (2006), Graf & Cummings (2007), Bogan (2010)
Diplodon rotundus — Graf & Cummings (2006)
Diplodon (Rhipidodonta) deceptus — Mansur & Pereira (2006), Pereira et al. (2014)
Diplodon grata — ITIS (2013)
Diplodon gratus — Miyahira et al. (2019), Graf & Cummings (2021), Campbell & Lydeard (2026)
Diplodon rufofuscus — Miyahira et al. (2019)
Rhipidodonta decepta — Mansur et al. (2020)
