MUSSELpdb
Taxonomy
Species & Genus Richness of
freshwater mussels
by Taxonomy
Species & Genus Richness of
freshwater bivalves
by Family
Geography
Species Richness & Endemism of
freshwater mussels
by Geography
Data last updated
Mon 2 March 2026
Bivalvia | ... | Unionidae | Unioninae | Anodontini | Cristariina | Anemina harlandi
nominal species Anodon euscaphys Heude, 1879
Anodon euscaphys Heude, 1879 = Anemina harlandi
Heude, 1879. Conch. Fluv. Nanking (5): pl. 35, fig. 68.
Type(s): TYPE USNM_126431, Tchen-kiang, Kiang-sou.
Taxonomic Opinions
+ Anemina harlandi (Baird & H. Adams, 1867) — Chen et al., 2026: 54.
= Anemina euscaphys — Xiang et al., 2025: Table 2.
= Anemina euscaphys (Heude, 1879) — Dai et al., 2025: Table 2.
+ Anemina arcaeformis — Wu et al., 2025: 11, fig. 5c, Table S1.
= Anodonta euscaphys — Perkins et al., 2025: Table S1.
+ Anemina arcaeformis — Hou et al., 2025: Table S3.
= Anemina euscaphys — Wu et al., 2024: 6.
= Anemina euscaphys — Wu et al., 2024: Table S3.
= Anemina euscaphys — Hou et al., 2024: Table 2.
= Anodonta euscaphys — Whelan et al., 2023: 13, Table S1.
= Anodonta euscaphys — Whelan et al., 2023: 13, Table S1.
= Anemina euscaphys — Wu et al., 2023: Table 2.
= Anemina euscaphys (Heude, 1879) — Wu et al., 2022: Table S2.
= Anemina euscaphys — Liu et al., 2022: Table 1.
= Anemina euscaphys (Heude, 1879) — Pavan-Kumar et al., 2022: Table S1.
= Anemina euscaphys (Heude, 1879) — Graf & Cummings, 2021: 16.
= Anemina euscaphys — Zieritz et al., 2021: 1092.
= Anemina euscaphys — Wu et al., 2021: attached Table 2.
= Anemina euscaphys — Lopes-Lima et al., 2020: 7.
= Anemina euscaphys — Riccardi et al., 2020: 750.
= Anemina euscaphys — Liu et al., 2020: [6].
= Anodonta euscaphys — Fukata & Iigo, 2020: 1216.
= Anemina euscaphys — Wu et al., 2020: 196.
= Anemina euscaphys — Sano et al., 2020: 101.
= Anodonta euscaphys — Chen et al., 2020: 2800.
= Anemina euscaphys — Huang et al., 2019: Tables S1, S5.
= Anemina euscaphys (Heude, 1879) — Wu et al., 2019: 29.
= Anemina euscaphys — Froufe et al., 2019: 185.
= Anemina euscaphys — Huang et al., 2019: Table S1.
= Anemina euscaphys (Heude, 1879) — Zieritz et al., 2018: online Table 1.
= Anemina euscaphys — Huang et al., 2018: 304.
= Anodonta euscaphys — Wang et al., 2018: 611.
= Anodonta euscaphys — Chase et al., 2018: Additional File 1.
= Anodonta euscaphys — Aunins et al., 2018: 1125.
= Anodonta euscaphys — Wang et al., 2018: 140.
= Anodonta euscaphys — Cha et al., 2018: 946.
= Anodonta euscaphys — Zhou et al., 2017: 235.
= Anemina euscaphys — Lopes-Lima et al., 2017: 250.
= Anodonta euscaphys — Soroka & Burzynski, 2017: 863.
= Anemina euscaphys (Heude, 1870) [sic] — Bogatov & Prozorova, 2017: 924.
= Anodonta euscaphys — Xue et al., 2016: 2698.
= Anemina euscaphys — Fonseca et al., 2016: 955.
= Anodonta euscaphys — Feng et al., 2016: 445.
= Anemina euscaphys (Heude) — Vinarski & Kantor, 2016: 45.
= Anemina euscaphys (Heude, 1879) — He & Zhuang, 2013: 31, figs. 62-63.
= Anodonta euscaphys — Xiong et al., 2012: 128, Table 2.
= Anemina euscaphys (Heude, 1879) — IUCN, 2011: [web].
= Anemina euscaphys Heude, 1879 — Bogan, 2010: [2].
= Anodonta euscaphys (Heude, 1879) — Graf, 2007: 87.
= Anemina euscaphys (Heude, 1879) — Graf & Cummings, 2007: 304.
= Anemina euscaphys (Heude, 1879) — Prozorova et al., 2005: 54.
= Anodonta enscaphys [sic] (Hende, 1879) [sic] — Bába, 2000: 133, fig. 2.
= Anodonta euscaphys (Heude, 1879) — Liu et al., 1979: 110, fig. 98.
+ — Haas, 1969.
= Anodonta (Haasiella) euscaphys (Heude, 1879) — Zhadin, 1965: 260, fig. 280.
= Anodonta (Haasiella) euscaphys (Heude, 1879) — Zhadin, 1938: 149, sp. 12, fig. 58.
= Anodonta euscaphys (Heude, 1879) — Simpson, 1914: 413.
= Anodonta euscaphys (Heude, 1879) — Simpson, 1900: 640.
= Anodon euscaphys Heude — Heude, 1879: pl. 35, fig. 68.
