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Taxonomy

Species & Genus Richness by
Taxonomy

Geography

Species Richness & Endemism by
Geography

Data last updated
Mon 1 Dec 2025

species Indonaia gratiosa (Philippi, 1843): synonymy

45 specimens + 1 Genbank voucher

Synonymy

Unio gratiosus Philippi, 1843

‘Parreyss’ Philippi, 1843. Abbild. Beschr. Conch. 1: 20, pl. 1, fig. 5.

Nodularia (s.s.) gratiosa — Simpson (1900), Simpson (1914), Preston (1915)
Ensidens (Uniandra) gratiosus — Haas (1969)
Contradens gratiosus — Graf & Cummings (2007), Zieritz et al. (2018), Konopleva et al. (2019), Graf & Cummings (2021)
Uniandra gratiosus — Bogan (2010), IUCN (2012)
Radiatula gratiosus — Pfeiffer et al. (2021)
Indonaia (cylindrica-group) gratiosa — Bolotov et al. (2022), Bolotov et al. (2024)
Indonaia gratiosa — Bolotov et al. (2022)

Unio occatus Lea, 1860

Lea, 1860. Proc. Acad. Nat. Sci. 12: 307.
Lea, 1863. Jour. Acad. Nat. Sci. 5 [O. 10]: 398, pl. 50, fig. 304.
Type(s): LECTOTYPE USNM_84077, Bengal, India.

Nodularia (s.s.) occata — Simpson (1900), Preston (1912), Simpson (1914), Preston (1915)
Indonaia occata — Annandale & Prashad (1921), Haas (1969), Graf & Cummings (2021)
Parreysia (Radiatula) occata — Subba Rao (1989), Graf & Cummings (2007), Ramakrishna & Dey (2007)
Radiatula (Indonaia) occata — Nesemann et al. (2003)
Radiatula occata — Nesemann et al. (2007), Zieritz et al. (2018)
Parreysia occata — Bogan (2010), IUCN (2010)
Indonaia (cylindrica-group) gratiosa — Bolotov et al. (2022)

Unio macilentus Benson, 1862

Benson, 1862. Ann. Mag. Nat. Hist. 10: 187.

Nodularia (s.s.) occata — Simpson (1900), Simpson (1914), Preston (1915)
Indonaia occata — Haas (1969)
Indonaia (cylindrica-group) gratiosa — Bolotov et al. (2022)

Unio siliguriensis Preston, 1908

Preston, 1908. Rec. Ind. Mus. 2: 47.
Preston, 1912. Rec. Ind. Mus. 7: 293.
Preston, 1915. Fauna Brit. India: 148.

Nodularia (Radiatula) lima var. siliguriensis — Preston (1912)
Nodularia (sec. Radiatula) lima var. siliguriensis — Simpson (1914), Preston (1915)
Radiatula lima — Haas (1969)
Parreysia (Radiatula) lima — Subba Rao (1989), Ramakrishna & Dey (2007)
Indonaia (cylindrica-group) gratiosa — Bolotov et al. (2022)

Unio rugosus Hanley & Theobald, 1876

specimen image from publication Hanley & Theobald, 1876. Conch. Indica (8): 62, pl. 154, fig. 3.
(misidentified reference to Unio rugosus Barnes, 1823)

Nodularia (s.s.) occata — Simpson (1900), Simpson (1914), Preston (1915)
Indonaia occata — Haas (1969)

Taxonomic History (since 1900)

Nodularia (s.s.) occata — Simpson (1900), Preston (1912), Simpson (1914), Preston (1915)
Nodularia (s.s.) gratiosa — Simpson (1900), Simpson (1914), Preston (1915)
Nodularia (Radiatula) lima var. siliguriensis — Preston (1912)
Nodularia (sec. Radiatula) lima var. siliguriensis — Simpson (1914), Preston (1915)
Indonaia occata — Annandale & Prashad (1921), Haas (1969), Graf & Cummings (2021)
Ensidens (Uniandra) gratiosus — Haas (1969)
Radiatula lima — Haas (1969)
Parreysia (Radiatula) occata — Subba Rao (1989), Graf & Cummings (2007), Ramakrishna & Dey (2007)
Parreysia (Radiatula) lima — Subba Rao (1989), Ramakrishna & Dey (2007)
Radiatula (Indonaia) occata — Nesemann et al. (2003)
Contradens gratiosus — Graf & Cummings (2007), Zieritz et al. (2018), Konopleva et al. (2019), Graf & Cummings (2021)
Radiatula occata — Nesemann et al. (2007), Zieritz et al. (2018)
Parreysia occata — Bogan (2010), IUCN (2010)
Uniandra gratiosus — Bogan (2010), IUCN (2012)
Radiatula gratiosus — Pfeiffer et al. (2021)
Indonaia (cylindrica-group) gratiosa — Bolotov et al. (2022), Bolotov et al. (2024)
Indonaia gratiosa — Bolotov et al. (2022)

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