species Lens micropterus (Morelet, 1866): synonymy

Unio cambojensis Sowerby, 1866

Sowerby, 1866. Conch. Icon. 16: pl. 42, sp. 231.
Type(s): LECTOTYPE BMNH_1965148, Camboja.

Nodularia (s.s.) rustica — Simpson (1900)
Contradens rusticus — Haas (1914)
Nodularia (s.s.) rusticus — Simpson (1914)
Contradens cambojensis — Ortmann (1917)
Uniandra contradens rustica — Brandt (1974)
Lens micropterus — Pfeiffer et al. (2021)

Unio micropterus Morelet, 1866

Morelet, 1866. J. de Conch. 14: 63.
Morelet, 1875. Séries Conch. 4: 349, pl. 15, fig. 6 [7].
Type(s): HOLOTYPE BMNH_93-2-4-1596, Cambodia.

Physunio (s.s.) micropterus — Simpson (1900), Preston (1912), Simpson (1914), Haas (1919), Haas (1924), Haas (1969)
Physunio micropterus — Brandt (1974), Graf & Cummings (2007), Bogan (2010), IUCN (2012), Zieritz et al. (2018), Do et al. (2018), Konopleva et al. (2019), Ng et al. (2020), Graf & Cummings (2021)
Uniandra contradens — Bogan & Do (2014)
Lens micropterus — Pfeiffer et al. (2021), Bolotov et al. (2022), Zieritz et al. (2024), Campbell & Lydeard (2026), Zieritz et al. (2026)

Margaron (Unio) versus Lea, 1870

‘Lea’ Reeve, 1865. Conch. Icon. 16: pl. 29, sp. 147 [as ‘Unio inornatus’].
Lea, 1870. Synopsis Fam. Naiades: 46.

Nodularia (s.s.) asperula — Simpson (1900), Simpson (1914)
Uniandra contradens tumidula — Brandt (1974)
Lens micropterus — Pfeiffer et al. (2021)

Unio crossei Deshayes in Deshayes & Jullien, 1876

Deshayes in Deshayes & Jullien, 1876. Nouv. Arch. Mus. Hist. Nat. Paris 10: 124, pl. 6, figs. 5-7.
Type(s): SYNTYPES MNHN_IM-2000-1668, Cambodge.

Physunio (s.s.) crossei — Simpson (1900), Simpson (1914)
Contradens crossei — Haas (1914), Haas (1969)
Uniandra contradens crossei — Brandt (1974)
Contradens contradens — Do et al. (2018)
“Contradens crossei” ** — Jeratthitikul et al. (2019), Konopleva et al. (2019), Bolotov et al. (2020), Graf & Cummings (2021)
Lens micropterus — Pfeiffer et al. (2021)

Unio semialatus Deshayes in Deshayes & Jullien, 1876

Deshayes in Deshayes & Jullien, 1876. Nouv. Arch. Mus. Hist. Nat. Paris 10: 123, pl. 6, figs. 1-2.
Type(s): SYNTYPE MNHN_IM-2000-1780, Peam Chelang, Cambodge.

Physunio (s.s.) semialatus — Simpson (1900), Simpson (1914), Haas (1919), Haas (1969)
Physunio micropterus — Brandt (1974), IUCN (2012), Graf & Cummings (2021)
Physunio semialatus — Graf & Cummings (2007), Bogan (2010)
Lens micropterus — Pfeiffer et al. (2021)

Diplodon ludovicianum Rochebrune, 1882

Rochebrune, 1882. Bull. Soc. Philom. Paris 6: 43.
Type(s): SYNTYPES MNHN_IM-2000-35806, Cambodge.

Trapezoideus ludovicianus — Simpson (1914), Haas (1919), Haas (1969)
Trapezoideus exolescens comptus — Brandt (1974)
Trapezoideus exolescens — He & Zhuang (2013)
Contradens comptus — Konopleva et al. (2019)
Lens micropterus — Pfeiffer et al. (2021)

Unio inaequalis Rochebrune, 1882

Rochebrune, 1882. Bull. Soc. Philom. Paris 6: 44 [in ‘inæqualis’].

Ensidens (sec. Uniandra) inaequalis — Haas (1913), Thiele (1934)
Nodularia (s.s.) inaequalis — Simpson (1914)
Ensidens (Uniandra) inaequalis — Haas (1969), Haas (1969)
Contradens inaequalis — Graf & Cummings (2007), Zieritz et al. (2018), Konopleva et al. (2019), Graf & Cummings (2021)
Uniandra inaequalis — Bogan (2010)
Lens micropterus — Pfeiffer et al. (2021)

Unio fischerianus Morlet, 1883

Morlet, 1883. J. de Conch. 31: 109, pl. 4, fig. 6.
Type(s): SYNTYPE MNHN_IM-2000-1697, Cambodge.

Nodularia (s.s.) fischeriania [sic] — Simpson (1900), Simpson (1914)
Contradens dimotus fischerianus — Haas (1913)
Contradens ascia fischerianus — Haas (1924), Haas (1952)
Contradens ascia — Haas (1969)
Uniandra contradens fischeriana — Brandt (1974)
Contradens contradens — Jeratthitikul et al. (2019), Ng et al. (2020)
Lens micropterus — Pfeiffer et al. (2021)

Unio siamensis Morlet, 1889

(renamed Unio soboles Fischer, 1891; homonym of Unio siamensis Lea, 1866)
Morlet, 1889. J. de Conch. 37: 164, 194, pl. 7, fig. 2.
Morlet, 1904. Mission Pavie 3: 381, pl. 2, fig. 8.

Nodularia (s.s.) sobolus [sic] — Simpson (1900)
Contradens rusticus — Haas (1914), Haas (1969)
Nodularia (s.s.) soboles — Simpson (1914)
Uniandra contradens rustica — Brandt (1974)
Lens micropterus — Pfeiffer et al. (2021)

Unio micropterus ‘Morelet’ of Authors

(reference to Unio micropterus Morelet, 1866)
‘Morelet’ of Authors.
Fischer, 1891. Bull. Soc. Hist. Nat. Autun 4: 226.
Fischer & Dautzenberg, 1904. Mission Pavie 3: 439.

Physunio (s.s.) semialatus — Haas (1919)

Unio soboles Fischer, 1891

(new name for Unio siamensis Morlet, 1889)
Fischer, 1891. Bull. Soc. Hist. Nat. Autun 4: 227.

Nodularia (s.s.) sobolus [sic] — Simpson (1900)
Contradens rusticus — Haas (1914), Haas (1969)
Nodularia (s.s.) soboles — Simpson (1914)
Uniandra contradens rustica — Brandt (1974)
Lens micropterus — Pfeiffer et al. (2021)

Contradens contradens ‘Lea’ of Authors

(misidentified reference to Unio contradens Lea, 1838)
‘Lea’ of Authors.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Contradens aff. contradens sp. 1’].
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Contradens aff. contradens sp. 1’].
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Contradens aff. contradens sp. 1’].
Konopleva et al., 2019. Ecologica Montenegrina 24: Appendix 1 [as ‘Contradens aff. contradens sp. 1’].
Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘Contradens aff. contradens sp. 1’].
Huang et al., 2019. MPE 130: Table S4 [as ‘Contradens aff. contradens sp. 1’].
Bolotov et al., 2020. Sci. Repts. 10: Table S1 [as ‘Contradens aff. contradens sp. 1’].
Konopleva et al., 2023. Diversity 15(1), 10: Table S2 [as ‘Lens aff. contradens’].

Lens micropterus — Pfeiffer et al. (2021)

Unio inornatus Reeve, 1865

(misidentified reference to Unio inornatus Lea, 1856)
‘Lea’ Reeve, 1865. Conch. Icon. 16: pl. 29, sp. 147; errata.

Nodularia (s.s.) asperula — Simpson (1900), Simpson (1914)
Ensidens (Uniandra) asperulus — Haas (1969)
Uniandra contradens tumidula — Brandt (1974)
Lens micropterus — Pfeiffer et al. (2021)

Nodularia sobolus Simpson, 1900

(error for Unio soboles Fischer, 1891)
‘Fischer’ Simpson, 1900. Proc. U.S. Nat. Mus. 22: 819.

Contradens rusticus — Haas (1914), Haas (1969)
Nodularia (s.s.) soboles — Simpson (1914)

Nodularia (s.s.) fischeriania Simpson, 1900

(error for Unio fischerianus Morlet, 1883)
‘Morlet’ Simpson, 1900. Proc. U.S. Nat. Mus. 22: 818.
‘Morlet’ Simpson, 1914. Descr. Cat. Naiades: 1009.