Publication Wu et al. (2026): cladograms

publication summary

Complete Citation

Wu, R., X. Wang, F. Nan, L. Cheng & J. An. 2026. An integrative taxonomic reassessment of the Chinese endemic genus Acuticosta (Bivalvia: Unionidae), with the description of a new genus. Contributions to Zoology 95: 25-51. Available electronically.

Cladogram Figures

(click the figure number to see the cladogram)

Cladogram Figure 3
                       +-- Margaritifera_margaritifera
  +--------------------+                              
  |                    +-- Gibbosula_rochechouartii   
  |                                                   
--+  +-------------------- Amblema_plicata            
  |  |                                                
  |  |           +-------- Gonidea_angulata           
  |  |  +--------+                                    
  +--+  |        |  +----- Inversidens_trisulcata     
     |  |        +--+                                 
     |  |           |  +-- Inversidens_brandtii       
     |  |           +--+                              
     +--+              +-- Inversidens_rentianensis   
        |                                             
        |  +-------------- Unio_pictorum              
        |  |                                          
        |  |           +-- Lanceolaria_gladiola       
        +--+  +--------+                              
           |  |        +-- Lanceolaria_lanceolata     
           |  |                                       
           |  |     +----- Globunio_mirificus         
           +--+  +--+                                 
              |  |  |  +-- Pseudoacuticosta_retiaria  
              |  |  +--+                              
              |  |     +-- Huizhou_xietianxiangi      
              +--+                                    
                 |     +-- Acuticosta_ovata           
                 |  +--+                              
                 |  |  +-- Acuticosta_chinensis       
                 +--+                                 
                    |  +-- Brachyanodon_californiensis
                    +--+                              
                       +-- Brachyanodon_nuttalliana
Cladogram Figure 4
                                               +-- Gibbosula_rochechouartii       
  +--------------------------------------------+                                  
  |                                            +-- Margaritifera_margaritifera    
  |                                                                               
  |                                         +----- Chamberlainia_somsakpanhai     
  |           +-----------------------------+                                     
  |           |                             |  +-- Sinohyriopsis_cumingii         
  |           |                             +--+                                  
  |           |                                +-- Sinohyriopsis_schlegelii       
  |           |                                                                   
  |           |                                +-- Hyriopsis_bialata              
  |           |                             +--+                                  
  |           |                             |  +-- Rectidens_sumatrensis          
  |  +--------+  +--------------------------+                                     
  |  |        |  |                          |  +-- Lens_contradens                
  |  |        |  |                          +--+                                  
  |  |        |  |                             +-- Physunio_superbus              
  |  |        |  |                                                                
  |  |        |  |                          +----- Potomida_semirugata__2         
  |  |        |  |                    +-----+                                     
  |  |        |  |                    |     |  +-- Potomida_acarnanica            
  |  |        +--+                    |     +--+                                  
  |  |           |  +-----------------+        +-- Potomida_semirugata__1         
  |  |           |  |                 |                                           
  |  |           |  |                 |  +-------- Pronodularia_japanensis        
--+  |           |  |                 +--+                                        
  |  |           |  |                    |  +----- Lamprotula_leaii               
  |  |           |  |                    +--+                                     
  |  |           |  |                       |  +-- Lamprotula_caveata             
  |  |           +--+                       +--+                                  
  |  |              |                          +-- Lamprotula_cornuumlunae        
  |  |              |                                                             
  |  |              |                          +-- Pilsbryoconcha_exilis          
  |  |              |  +-----------------------+                                  
  |  |              |  |                       +-- Pseudodon_mekongi              
  |  |              |  |                                                          
  |  |              |  |                       +-- Inversidens_rentianensis       
  |  |              +--+  +--------------------+                                  
  |  |                 |  |                    +-- Inversidens_trisulcata         
  |  |                 |  |                                                       
  |  |                 |  |              +-------- Sinosolenaia_carinata          
  |  |                 +--+  +-----------+                                        
  |  |                    |  |           |  +----- Sinosolenaia_recognita         
  |  |                    |  |           +--+                                     
  |  |                    |  |              |  +-- Sinosolenaia_iridinea          
  |  |                    +--+              +--+                                  
  |  |                       |                 +-- Sinosolenaia_oleivora          
  |  |                       |                                                    
  |  |                       |  +----------------- Parvasolenaia_rivularis        
  +--+                       +--+                                                 
     |                          |  +-------------- Pseudopostolata_angula         
     |                          +--+                                              
     |                             |  +----------- Koreosolenaia_sitgyensis       
     |                             +--+                                           
     |                                |  +-------- Ptychorhynchus_pfisteri        
     |                                +--+                                        
     |                                   |  +----- Cosmopseudodon_resupinatus     
     |                                   +--+                                     
     |                                      |  +-- Postolata_guangxiensis         
     |                                      +--+                                  
     |                                         +-- Postolata_longjiangensis       
     |                                                                            
     |                                         +-- Lepidodesma_aligera            
     |  +--------------------------------------+                                  
     |  |                                      +-- Lepidodesma_languilati         
     |  |                                                                         
     |  |                                      +-- Acuticosta_chinensis           
     |  |                                +-----+                                  
     |  |                                |     +-- Acuticosta_ovata               
     |  |           +--------------------+                                        
     |  |           |                    |  +----- Lanceolaria_gladiola           
     |  |           |                    +--+                                     
     |  |           |                       |  +-- Lanceolaria_grayii             
     |  |           |                       +--+                                  
     |  |           |                          +-- Lanceolaria_lanceolata         
     |  |           |                                                             
     |  |           |                       +----- Anodonta_anatina               
     |  |           |           +-----------+                                     
     |  |           |           |           |  +-- Anodonta_cygnea                
     |  |           |           |           +--+                                  
     |  |  +--------+           |              +-- Anodonta_exulcerata            
     |  |  |        |  +--------+                                                 
     |  |  |        |  |        |  +-------------- Pseudanodonta_complanata       
     |  |  |        |  |        |  |                                              
     |  |  |        |  |        |  |        +----- Pyganodon_grandis              
     +--+  |        |  |        +--+  +-----+                                     
        |  |        |  |           |  |     |  +-- Utterbackia_imbecillis         
        |  |        |  |           |  |     +--+                                  
        |  |        |  |           +--+        +-- Utterbackia_peninsularis       
        |  |        |  |              |                                           
        |  |        |  |              |  +-------- Pterosyna_complanata           
        |  |        |  |              +--+                                        
        |  |        +--+                 |  +----- Prolasmidonta_heterodon        
        |  |           |                 +--+                                     
        |  |           |                    |  +-- Alasmidonta_varicosa           
        |  |           |                    +--+                                  
        |  |           |                       +-- Platynaias_compressa           
        |  |           |                                                          
        |  |           |                    +----- Cristaria_plicata__2           
        |  |           |  +-----------------+                                     
        |  |           |  |                 |  +-- Cristaria_plicata__1           
        |  |           |  |                 +--+                                  
        |  |           |  |                    +-- Cristaria_truncata             
        |  |           |  |                                                       
        |  |           +--+                 +----- Amuranodonta_kijaensis         
        |  |              |  +--------------+                                     
        |  |              |  |              |  +-- Anemina_harlandi               
        |  |              |  |              +--+                                  
        +--+              |  |                 +-- Buldowskia_shadini             
           |              +--+                                                    
           |                 |                 +-- Beringiana_fukuharai           
           |                 |  +--------------+                                  
           |                 |  |              +-- Pletholophus_guangzhouensis    
           |                 +--+                                                 
           |                    |  +-------------- Acudonta_baitiaoensis          
           |                    |  |                                              
           |                    +--+  +----------- Sinanodonta_lucida             
           |                       |  |                                           
           |                       +--+        +-- Sinanodonta_schrenkii          
           |                          |  +-----+                                  
           |                          |  |     +-- Sinanodonta_tumens             
           |                          +--+                                        
           |                             |  +----- Sinanodonta_woodiana           
           |                             +--+                                     
           |                                |  +-- Sinanodonta_lauta              
           |                                +--+                                  
           |                                   +-- Sinanodonta_pacifica           
           |                                                                      
           |                                +----- Pseudoacuticosta_retiaria      
           |  +-----------------------------+                                     
           |  |                             |  +-- Globunio_mirificus             
           |  |                             +--+                                  
           |  |                                +-- Huizhou_xietianxiangi          
           |  |                                                                   
           |  |                          +-------- Scriptolamprotula_tortuosa     
           |  |                          |                                        
           +--+  +-----------------------+     +-- Scriptolamprotula_coreana      
              |  |                       |  +--+                                  
              |  |                       |  |  +-- Scriptolamprotula_tientsinensis
              |  |                       +--+                                     
              |  |                          |  +-- Scriptolamprotula_polysticta   
              |  |                          +--+                                  
              |  |                             +-- Scriptolamprotula_scripta      
              +--+                                                                
                 |                       +-------- Unio_tumidus                   
                 |  +--------------------+                                        
                 |  |                    |  +----- Unio_elongatulus               
                 |  |                    +--+                                     
                 |  |                       |  +-- Unio_carneus                   
                 |  |                       +--+                                  
                 |  |                          +-- Unio_crassus                   
                 +--+                                                             
                    |  +-------------------------- Middendorffinaia_mongolica     
                    |  |                                                          
                    |  |                 +-------- Pseudocuneopsis_capitata       
                    |  |  +--------------+                                        
                    |  |  |              |  +----- Pseudocuneopsis_sichuanensis   
                    +--+  |              +--+                                     
                       |  |                 |  +-- Pseudocuneopsis_wuana          
                       |  |                 +--+                                  
                       |  |                    +-- Pseudocuneopsis_yangshuoensis  
                       |  |                                                       
                       +--+              +-------- Nodularia_hanensis             
                          |  +-----------+                                        
                          |  |           |  +----- Nodularia_fusiformans          
                          |  |           +--+                                     
                          |  |              |  +-- Nodularia_douglasiae           
                          |  |              +--+                                  
                          |  |                 +-- Nodularia_guiensis             
                          +--+                                                    
                             |        +----------- Schistodesmus_spinosus         
                             |  +-----+                                           
                             |  |     |  +-------- Schistodesmus_tongpenensis     
                             |  |     +--+                                        
                             |  |        |  +----- Schistodesmus_xinyuensis       
                             |  |        +--+                                     
                             +--+           |  +-- Schistodesmus_lampreyanus      
                                |           +--+                                  
                                |              +-- Schistodesmus_luqiaoensis      
                                |                                                 
                                |  +-------------- Tchangsinaia_piscicula         
                                |  |                                              
                                +--+  +----------- Cuneopsis_rufescens            
                                   |  |                                           
                                   +--+        +-- Cuneopsis_heudei               
                                      |  +-----+                                  
                                      |  |     +-- Cuneopsis_szechenyii           
                                      +--+                                        
                                         |  +----- Cuneopsis_celtiformis          
                                         +--+                                     
                                            |  +-- Cuneopsis_celtiosimilis        
                                            +--+                                  
                                               +-- Cuneopsis_kiangsiensis