Publication Wu et al. (2026): cladograms

publication summary

Full Citation

Wu, R., X. Wang, X. Hu, F. Nan, X. Liu et al. 2026. Integrative taxonomic reappraisal and molecular phylogeny of freshwater mussels of the genus Lanceolaria (Bivalvia: Unionidae), with description of two new species. Zoologica Scripta: 1-24. https://doi.org/10.1111/zsc.70061.

Cladogram Figures

(click the figure number to see the cladogram)

Cladogram Figure 3

                    +-- Schistodesmus_spinosus   
  +-----------------+                            
  |                 +-- Schistodesmus_xinyuensis 
  |                                              
--+  +----------------- Lanceolaria_oxyrhyncha   
  |  |                                           
  |  |              +-- Lanceolaria_gladiola     
  +--+  +-----------+                            
     |  |           +-- Lanceolaria_bilirata     
     |  |                                        
     +--+        +----- Lanceolaria_acrorrhyncha 
        |  +-----+                               
        |  |     |  +-- Lanceolaria_grayii       
        |  |     +--+                            
        +--+        +-- Lanceolaria_danzhouensis 
           |                                     
           |  +-------- Lanceolaria_kihirai      
           +--+                                  
              |  +----- Lanceolaria_triformis    
              +--+                               
                 |  +-- Lanceolaria_lanceolata   
                 +--+                            
                    +-- Lanceolaria_helianthoides

Cladogram Figure 5

                             +-- Lamprotula_leaii               
  +--------------------------+                                  
  |                          +-- Potomida_littoralis            
  |                                                             
  |                          +-- Schistodesmus_xinyuensis       
  |                    +-----+                                  
--+                    |     +-- Schistodesmus_spinosus         
  |  +-----------------+                                        
  |  |                 |  +----- Nodularia_fusiformans          
  |  |                 +--+                                     
  |  |                    |  +-- Nodularia_douglasiae           
  |  |                    +--+                                  
  +--+                       +-- Nodularia_dualobtusa           
     |                                                          
     |                       +-- Scriptolamprotula_scripta      
     |  +--------------------+                                  
     |  |                    +-- Scriptolamprotula_tientsinensis
     |  |                                                       
     +--+  +-------------------- Lanceolaria_oxyrhyncha         
        |  |                                                    
        |  |                 +-- Lanceolaria_bilirata           
        +--+  +--------------+                                  
           |  |              +-- Lanceolaria_gladiola           
           |  |                                                 
           +--+              +-- Lanceolaria_grayii             
              |  +-----------+                                  
              |  |           +-- Lanceolaria_danzhouensis       
              +--+                                              
                 |  +----------- Lanceolaria_acrorrhyncha       
                 +--+                                           
                    |  +-------- Lanceolaria_kihirai            
                    +--+                                        
                       |  +----- Lanceolaria_triformis          
                       +--+                                     
                          |  +-- Lanceolaria_helianthoides      
                          +--+                                  
                             +-- Lanceolaria_lanceolata

Cladogram Figure 6B

                                         +-- Gibbosula_rochechouartii       
  +--------------------------------------+                                  
  |                                      +-- Margaritifera_margaritifera    
  |                                                                         
  |                                      +-- Lepidodesma_aligera            
  |  +-----------------------------------+                                  
  |  |                                   +-- Lepidodesma_languilati         
  |  |                                                                      
  |  |                             +-------- Scriptolamprotula_tortuosa     
  |  |     +-----------------------+                                        
  |  |     |                       |  +----- Scriptolamprotula_scripta      
  |  |     |                       +--+                                     
  |  |     |                          |  +-- Scriptolamprotula_coreana      
  |  |     |                          +--+                                  
--+  |     |                             +-- Scriptolamprotula_tientsinensis
  |  |     |                                                                
  |  |  +--+                       +-------- Unio_tumidus                   
  |  |  |  |  +--------------------+                                        
  |  |  |  |  |                    |  +----- Unio_elongatulus               
  |  |  |  |  |                    +--+                                     
  |  |  |  |  |                       |  +-- Unio_carneus                   
  |  |  |  |  |                       +--+                                  
  |  |  |  |  |                          +-- Unio_nanus                     
  |  |  |  +--+                                                             
  |  |  |     |  +-------------------------- Middendorffinaia_mongolica     
  |  |  |     |  |                                                          
  |  |  |     |  |                 +-------- Pseudocuneopsis_capitata       
  +--+  |     |  |  +--------------+                                        
     |  |     |  |  |              |  +----- Pseudocuneopsis_sichuanensis   
     |  |     +--+  |              +--+                                     
     |  |        |  |                 |  +-- Pseudocuneopsis_wuana          
     |  |        |  |                 +--+                                  
     |  |        |  |                    +-- Pseudocuneopsis_yangshuoensis  
     |  |        |  |                                                       
     |  |        +--+              +-------- Nodularia_hanensis             
     |  |           |  +-----------+                                        
     |  |           |  |           |  +----- Nodularia_fusiformans          
     |  |           |  |           +--+                                     
     |  |           |  |              |  +-- Nodularia_douglasiae           
     |  |           |  |              +--+                                  
     |  |           |  |                 +-- Nodularia_guiensis             
     |  |           +--+                                                    
     |  |              |        +----------- Schistodesmus_spinosus         
     |  |              |  +-----+                                           
     |  |              |  |     |  +-------- Schistodesmus_tongpenensis     
     |  |              |  |     +--+                                        
     |  |              |  |        |  +----- Schistodesmus_xinyuensis       
     |  |              |  |        +--+                                     
     |  |              +--+           |  +-- Schistodesmus_lampreyanus      
     +--+                 |           +--+                                  
        |                 |              +-- Schistodesmus_luqiaoensis      
        |                 |                                                 
        |                 |  +-------------- Tchangsinaia_piscicula         
        |                 |  |                                              
        |                 +--+  +----------- Cuneopsis_rufescens            
        |                    |  |                                           
        |                    +--+        +-- Cuneopsis_heudei               
        |                       |  +-----+                                  
        |                       |  |     +-- Cuneopsis_szechenyii           
        |                       +--+                                        
        |                          |  +----- Cuneopsis_celtiformis          
        |                          +--+                                     
        |                             |  +-- Cuneopsis_celtiosimilis        
        |                             +--+                                  
        |                                +-- Cuneopsis_kiangsiensis         
        |                                                                   
        |                                +-- Lanceolaria_bilirata           
        |                       +--------+                                  
        |                       |        +-- Lanceolaria_gladiola           
        |                       |                                           
        |     +-----------------+        +-- Lanceolaria_grayii             
        |     |                 |  +-----+                                  
        |     |                 |  |     +-- Lanceolaria_danzhouensis       
        |     |                 +--+                                        
        |     |                    |  +----- Lanceolaria_triformis          
        |     |                    +--+                                     
        |     |                       |  +-- Lanceolaria_helianthoides      
        |     |                       +--+                                  
        |     |                          +-- Lanceolaria_lanceolata         
        |     |                                                             
        |     |                       +----- Anodonta_anatina               
        |     |           +-----------+                                     
        |     |           |           |  +-- Anodonta_cygnea                
        +-----+           |           +--+                                  
              |           |              +-- Anodonta_exulcerata            
              |  +--------+                                                 
              |  |        |  +-------------- Pseudanodonta_complanata       
              |  |        |  |                                              
              |  |        |  |        +----- Pyganodon_grandis              
              |  |        +--+  +-----+                                     
              |  |           |  |     |  +-- Utterbackia_imbecillis         
              |  |           |  |     +--+                                  
              |  |           +--+        +-- Utterbackia_peninsularis       
              |  |              |                                           
              |  |              |  +-------- Pterosyna_complanata           
              |  |              +--+                                        
              +--+                 |  +----- Prolasmidonta_heterodon        
                 |                 +--+                                     
                 |                    |  +-- Alasmidonta_varicosa           
                 |                    +--+                                  
                 |                       +-- Platynaias_compressa           
                 |                                                          
                 |                    +----- Cristaria_liboi                
                 |  +-----------------+                                     
                 |  |                 |  +-- Cristaria_plicata              
                 |  |                 +--+                                  
                 |  |                    +-- Cristaria_truncata             
                 |  |                                                       
                 +--+                 +----- Amuranodonta_kijaensis         
                    |  +--------------+                                     
                    |  |              |  +-- Anemina_harlandi               
                    |  |              +--+                                  
                    |  |                 +-- Buldowskia_shadini             
                    +--+                                                    
                       |                 +-- Beringiana_fukuharai           
                       |  +--------------+                                  
                       |  |              +-- Pletholophus_guangzhouensis    
                       +--+                                                 
                          |  +-------------- Acudonta_baitiaoensis          
                          |  |                                              
                          +--+  +----------- Sinanodonta_lucida             
                             |  |                                           
                             +--+        +-- Sinanodonta_schrenkii          
                                |  +-----+                                  
                                |  |     +-- Sinanodonta_tumens             
                                +--+                                        
                                   |  +----- Sinanodonta_pacifica__2        
                                   +--+                                     
                                      |  +-- Sinanodonta_lauta              
                                      +--+                                  
                                         +-- Sinanodonta_pacifica__1