Publication Konopleva et al. (2026): cladograms

publication summary

Full Citation

Konopleva, E.S., Y. Tenjing Singh, S.D. Sorokhaibam, M.M. Naorem, B.C. Loukrakpam et al. 2026. The freshwater mussel fauna of Manipur (North-East India) represents a tectonically derived mixture of endemic species and (sub)recent immigrants from adjacent basins. Biological Journal of the Linnean Society 148(3): ??. https://doi.org/10.1093/biolinnean/blag041.

Cladogram Figures

(click the figure number to see the cladogram)

Cladogram Figure 2

                             +-------------- Balwantia_soleniformis       
                             |                                            
                 +-----------+  +----------- Trapezidens_mogaungensis     
                 |           |  |                                         
                 |           +--+        +-- Trapezidens_exolescens       
                 |              |  +-----+                                
                 |              |  |     +-- Trapezidens_scutum           
                 |              +--+                                      
                 |                 |  +----- Trapezidens_yeti             
  +--------------+                 +--+                                   
  |              |                    |  +-- Trapezidens_angustior        
  |              |                    +--+                                
  |              |                       +-- Trapezidens_dolichorhynchus  
  |              |                                                        
  |              |  +----------------------- Lamellidens_corrianus        
  |              |  |                                                     
  |              +--+  +-------------------- Lamellidens_nongyangensis    
  |                 |  |                                                  
  |                 +--+                 +-- Lamellidens_savadiensis      
  |                    |  +--------------+                                
  |                    |  |              +-- Lamellidens_chindwinensis    
  |                    +--+                                               
  |                       |  +-------------- Lamellidens_ferrugineus      
  |                       |  |                                            
  |                       +--+  +----------- Lamellidens_marginalis       
  |                          |  |                                         
  |                          +--+  +-------- Lamellidens_generosus        
  |                             |  |                                      
  |                             +--+     +-- Lamellidens_mainwaringi      
  |                                |  +--+                                
  |                                |  |  +-- Lamellidens_phenchooganjensis
  |                                +--+                                   
  |                                   |  +-- Lamellidens_lamellatus       
--+                                   +--+                                
  |                                      +-- Lamellidens_jenkinsianus     
  |                                                                       
  |                                      +-- Leoparreysia_whitteni        
  |                                +-----+                                
  |                                |     +-- Leoparreysia_burmana         
  |                    +-----------+                                      
  |                    |           |  +----- Leoparreysia_olivacea        
  |                    |           +--+                                   
  |                    |              |  +-- Leoparreysia_tavoyensis      
  |  +-----------------+              +--+                                
  |  |                 |                 +-- Leoparreysia_canefrii        
  |  |                 |                                                  
  |  |                 |  +----------------- Assamnaia_involuta           
  |  |                 |  |                                               
  |  |                 +--+              +-- Parreysia_marcens            
  |  |                    |  +-----------+                                
  |  |                    |  |           +-- Parreysia_keralaensis        
  |  |                    +--+                                            
  |  |                       |  +----------- Parreysia_plagiosoma         
  |  |                       +--+                                         
  |  |                          |  +-------- Parreysia_corrugata          
  |  |                          +--+                                      
  |  |                             |  +----- Parreysia_nagpoorensis       
  |  |                             +--+                                   
  +--+                                |  +-- Parreysia_rakhinensis        
     |                                +--+                                
     |                                   +-- Parreysia_imphalensis        
     |                                                                    
     |                             +-------- Nitia_teretiuscula           
     |  +--------------------------+                                      
     |  |                          |  +----- Coelatura_aegyptiaca         
     |  |                          +--+                                   
     |  |                             |  +-- Nyassunio_nyassaensis        
     |  |                             +--+                                
     |  |                                +-- Coelatura_kunenensis         
     |  |                                                                 
     |  |              +-------------------- Pseudoscabies_scobina        
     |  |              |                                                  
     |  |              |                 +-- Indonaia_cylindrica          
     |  |  +-----------+  +--------------+                                
     +--+  |           |  |              +-- Indonaia_gratiosa            
        |  |           |  |                                               
        |  |           +--+  +-------------- Indonaia_corbis              
        |  |              |  |                                            
        |  |              |  |        +----- Indonaia_shurtleffiana       
        |  |              +--+  +-----+                                   
        |  |                 |  |     |  +-- Indonaia_bonneaudii          
        |  |                 |  |     +--+                                
        |  |                 +--+        +-- Indonaia_rugosa              
        |  |                    |                                         
        |  |                    |  +-------- Indonaia_andersoniana        
        +--+                    +--+                                      
           |                       |  +----- Indonaia_subclathrata        
           |                       +--+                                   
           |                          |  +-- Indonaia_caerulea            
           |                          +--+                                
           |                             +-- Indonaia_manipurensis        
           |                                                              
           |  +----------------------------- Indochinella_pugio           
           |  |                                                           
           |  |                          +-- Radiatula_crispisulcata      
           |  |                 +--------+                                
           |  |                 |        +-- Radiatula_theobaldi          
           |  |                 |                                         
           +--+  +--------------+     +----- Radiatula_kachinensis        
              |  |              |  +--+                                   
              |  |              |  |  |  +-- Radiatula_myitkyinae         
              |  |              |  |  +--+                                
              |  |              +--+     +-- Radiatula_chaudhurii         
              |  |                 |                                      
              |  |                 |  +----- Radiatula_mouhoti            
              +--+                 +--+                                   
                 |                    |  +-- Radiatula_myitthanensis      
                 |                    +--+                                
                 |                       +-- Radiatula_chindwinensis      
                 |                                                        
                 |  +----------------------- Scabiellus_songkramensis     
                 |  |                                                     
                 +--+                    +-- Unionetta_fabagina           
                    |  +-----------------+                                
                    |  |                 +-- Harmandia_somboriensis       
                    +--+                                                  
                       |  +----------------- Scabies_mandarinus           
                       |  |                                               
                       +--+----------------- Scabies_crispata             
                          |                                               
                          |  +-------------- Scabies_nucleus              
                          +--+                                            
                             |  +----------- Scabies_pilata               
                             +--+                                         
                                |  +-------- Scabies_humilis              
                                +--+                                      
                                   |  +----- Scabies_anceps               
                                   +--+                                   
                                      |  +-- Scabies_phaselus             
                                      +--+                                
                                         +-- Scabies_scobinatus

Cladogram Figure S1

                             +-------------- Balwantia_soleniformis       
                             |                                            
                 +-----------+  +----------- Trapezidens_mogaungensis     
                 |           |  |                                         
                 |           +--+  +-------- Trapezidens_exolescens       
                 |              |  |                                      
                 |              +--+  +----- Trapezidens_scutum           
                 |                 |  |                                   
                 |                 +--+  +-- Trapezidens_angustior        
  +--------------+                    |  |                                
  |              |                    +--+-- Trapezidens_dolichorhynchus  
  |              |                       |                                
  |              |                       +-- Trapezidens_yeti             
  |              |                                                        
  |              |  +----------------------- Lamellidens_corrianus        
  |              |  |                                                     
  |              +--+  +-------------------- Lamellidens_nongyangensis    
  |                 |  |                                                  
  |                 |  |                 +-- Lamellidens_chindwinensis    
  |                 +--+  +--------------+                                
  |                    |  |              +-- Lamellidens_savadiensis      
  |                    |  |                                               
  |                    +--+              +-- Lamellidens_ferrugineus      
  |                       |  +-----------+                                
  |                       |  |           +-- Lamellidens_marginalis       
  |                       +--+                                            
  |                          |  +----------- Lamellidens_generosus        
  |                          +--+                                         
  |                             |  +-------- Lamellidens_phenchooganjensis
  |                             +--+                                      
  |                                |  +----- Lamellidens_mainwaringi      
  |                                +--+                                   
  |                                   |  +-- Lamellidens_lamellatus       
--+                                   +--+                                
  |                                      +-- Lamellidens_jenkinsianus     
  |                                                                       
  |                                      +-- Leoparreysia_burmana         
  |                                +-----+                                
  |                                |     +-- Leoparreysia_whitteni        
  |                    +-----------+                                      
  |                    |           |  +----- Leoparreysia_olivacea        
  |                    |           +--+                                   
  |                    |              |  +-- Leoparreysia_canefrii        
  |  +-----------------+              +--+                                
  |  |                 |                 +-- Leoparreysia_tavoyensis      
  |  |                 |                                                  
  |  |                 |  +----------------- Assamnaia_involuta           
  |  |                 |  |                                               
  |  |                 +--+              +-- Parreysia_marcens            
  |  |                    |  +-----------+                                
  |  |                    |  |           +-- Parreysia_keralaensis        
  |  |                    +--+                                            
  |  |                       |  +----------- Parreysia_imphalensis        
  |  |                       +--+                                         
  |  |                          |  +-------- Parreysia_rakhinensis        
  |  |                          +--+                                      
  |  |                             |  +----- Parreysia_nagpoorensis       
  |  |                             +--+                                   
  +--+                                |  +-- Parreysia_corrugata          
     |                                +--+                                
     |                                   +-- Parreysia_plagiosoma         
     |                                                                    
     |                             +-------- Nitia_teretiuscula           
     |  +--------------------------+                                      
     |  |                          |  +----- Coelatura_aegyptiaca         
     |  |                          +--+                                   
     |  |                             |  +-- Coelatura_kunenensis         
     |  |                             +--+                                
     |  |                                +-- Nyassunio_nyassaensis        
     |  |                                                                 
     |  |           +----------------------- Pseudoscabies_scobina        
     |  |           |                                                     
     |  |  +--------+  +-------------------- Indonaia_cylindrica          
     |  |  |        |  |                                                  
     +--+  |        +--+  +----------------- Indonaia_gratiosa            
        |  |           |  |                                               
        |  |           +--+  +-------------- Indonaia_corbis              
        |  |              |  |                                            
        |  |              |  |        +----- Indonaia_rugosa              
        |  |              +--+  +-----+                                   
        |  |                 |  |     |  +-- Indonaia_bonneaudii          
        |  |                 |  |     +--+                                
        |  |                 +--+        +-- Indonaia_shurtleffiana       
        |  |                    |                                         
        |  |                    |  +-------- Indonaia_andersoniana        
        +--+                    +--+                                      
           |                       |  +----- Indonaia_subclathrata        
           |                       +--+                                   
           |                          |  +-- Indonaia_caerulea            
           |                          +--+                                
           |                             +-- Indonaia_manipurensis        
           |                                                              
           |  +----------------------------- Indochinella_pugio           
           |  |                                                           
           |  |                          +-- Radiatula_crispisulcata      
           |  |                 +--------+                                
           |  |                 |        +-- Radiatula_theobaldi          
           |  |                 |                                         
           +--+  +--------------+     +----- Radiatula_kachinensis        
              |  |              |  +--+                                   
              |  |              |  |  |  +-- Radiatula_chaudhurii         
              |  |              |  |  +--+                                
              |  |              +--+     +-- Radiatula_myitkyinae         
              |  |                 |                                      
              |  |                 |  +----- Radiatula_mouhoti            
              +--+                 +--+                                   
                 |                    |  +-- Radiatula_chindwinensis      
                 |                    +--+                                
                 |                       +-- Radiatula_myitthanensis      
                 |                                                        
                 |  +----------------------- Scabiellus_songkramensis     
                 |  |                                                     
                 +--+  +-------------------- Harmandia_somboriensis       
                    |  |                                                  
                    +--+  +----------------- Unionetta_fabagina           
                       |  |                                               
                       +--+              +-- Scabies_crispata             
                          |  +-----------+                                
                          |  |           +-- Scabies_mandarinus           
                          +--+                                            
                             |  +----------- Scabies_nucleus              
                             |  |                                         
                             +--+        +-- Scabies_humilis              
                                |  +-----+                                
                                |  |     +-- Scabies_pilata               
                                +--+                                      
                                   |  +----- Scabies_phaselus             
                                   +--+                                   
                                      |  +-- Scabies_anceps               
                                      +--+                                
                                         +-- Scabies_scobinatus

Cladogram Figure S2

  +----------------------------------------------------- Indopseudodon_kayinensis     
  |                                                                                   
  |                                                  +-- Parreysia_marcens            
  |                                      +-----------+                                
  |                                      |           +-- Parreysia_keralaensis        
  |  +-----------------------------------+                                            
  |  |                                   |  +----------- Parreysia_corrugata          
  |  |                                   +--+                                         
  |  |                                      |  +-------- Parreysia_nagpoorensis       
  |  |                                      +--+                                      
  |  |                                         |  +----- Parreysia_plagiosoma         
  |  |                                         +--+                                   
  |  |                                            |  +-- Parreysia_rakhinensis        
--+  |                                            +--+                                
  |  |                                               +-- Parreysia_imphalensis        
  |  |                                                                                
  |  |                                               +-- Leoparreysia_whitteni        
  |  |                                         +-----+                                
  |  |                                         |     +-- Leoparreysia_burmana         
  |  |                    +--------------------+                                      
  |  |                    |                    |  +----- Leoparreysia_olivacea        
  |  |                    |                    +--+                                   
  |  |                    |                       |  +-- Leoparreysia_canefrii        
  |  |                    |                       +--+                                
  |  |                    |                          +-- Leoparreysia_tavoyensis      
  |  |                    |                                                           
  +--+                    |           +----------------- Balwantia_soleniformis       
     |  +-----------------+  +--------+                                               
     |  |                 |  |        |  +-------------- Trapezidens_mogaungensis     
     |  |                 |  |        +--+                                            
     |  |                 |  |           |  +----------- Trapezidens_exolescens       
     |  |                 |  |           +--+                                         
     |  |                 |  |              |  +-------- Trapezidens_scutum           
     |  |                 |  |              +--+                                      
     |  |                 |  |                 |  +----- Trapezidens_yeti             
     |  |                 +--+                 +--+                                   
     |  |                    |                    |  +-- Trapezidens_angustior        
     |  |                    |                    +--+                                
     |  |                    |                       +-- Trapezidens_dolichorhynchus  
     |  |                    |                                                        
     |  |                    |                       +-- Lamellidens_corrianus        
     |  |                    |  +--------------------+                                
     |  |                    |  |                    +-- Lamellidens_nongyangensis    
     |  |                    +--+                                                     
     |  |                       |  +-------------------- Lamellidens_savadiensis      
     |  |                       |  |                                                  
     |  |                       +--+  +----------------- Lamellidens_chindwinensis    
     |  |                          |  |                                               
     +--+                          +--+              +-- Lamellidens_marginalis       
        |                             |  +-----------+                                
        |                             |  |           +-- Lamellidens_ferrugineus      
        |                             +--+                                            
        |                                |  +----------- Lamellidens_generosus        
        |                                +--+                                         
        |                                   |  +-------- Lamellidens_phenchooganjensis
        |                                   +--+                                      
        |                                      |  +----- Lamellidens_mainwaringi      
        |                                      +--+                                   
        |                                         |  +-- Lamellidens_jenkinsianus     
        |                                         +--+                                
        |                                            +-- Lamellidens_lamellatus       
        |                                                                             
        |  +-------------------------------------------- Assamnaia_involuta           
        |  |                                                                          
        |  |                                   +-------- Nitia_teretiuscula           
        |  |  +--------------------------------+                                      
        |  |  |                                |  +----- Coelatura_aegyptiaca         
        |  |  |                                +--+                                   
        |  |  |                                   |  +-- Nyassunio_nyassaensis        
        +--+  |                                   +--+                                
           |  |                                      +-- Coelatura_kunenensis         
           |  |                                                                       
           |  |                    +-------------------- Pseudoscabies_scobina        
           |  |                    |                                                  
           |  |                    |                 +-- Indonaia_cylindrica          
           |  |  +-----------------+  +--------------+                                
           +--+  |                 |  |              +-- Indonaia_gratiosa            
              |  |                 |  |                                               
              |  |                 +--+  +-------------- Indonaia_corbis              
              |  |                    |  |                                            
              |  |                    |  |        +----- Indonaia_shurtleffiana       
              |  |                    +--+  +-----+                                   
              |  |                       |  |     |  +-- Indonaia_rugosa              
              |  |                       |  |     +--+                                
              |  |                       +--+        +-- Indonaia_bonneaudii          
              |  |                          |                                         
              |  |                          |  +-------- Indonaia_andersoniana        
              +--+                          +--+                                      
                 |                             |  +----- Indonaia_subclathrata        
                 |                             +--+                                   
                 |                                |  +-- Indonaia_caerulea            
                 |                                +--+                                
                 |                                   +-- Indonaia_manipurensis        
                 |                                                                    
                 |  +----------------------------------- Indochinella_pugio           
                 |  |                                                                 
                 |  |                                +-- Radiatula_theobaldi          
                 |  |                    +-----------+                                
                 |  |                    |           +-- Radiatula_crispisulcata      
                 |  |                    |                                            
                 +--+  +-----------------+           +-- Radiatula_myitkyinae         
                    |  |                 |  +--------+                                
                    |  |                 |  |        +-- Radiatula_kachinensis        
                    |  |                 +--+                                         
                    |  |                    |  +-------- Radiatula_chaudhurii         
                    |  |                    +--+                                      
                    +--+                       |  +----- Radiatula_chindwinensis      
                       |                       +--+                                   
                       |                          |  +-- Radiatula_mouhoti            
                       |                          +--+                                
                       |                             +-- Radiatula_myitthanensis      
                       |                                                              
                       |  +----------------------------- Scabiellus_songkramensis     
                       +--+                                                           
                          |  +-------------------------- Harmandia_somboriensis       
                          +--+                                                        
                             |  +----------------------- Unionetta_fabagina           
                             +--+                                                     
                                |  +-------------------- Scabies_crispata             
                                +--+                                                  
                                   |  +----------------- Scabies_mandarinus           
                                   +--+                                               
                                      |  +-------------- Scabies_nucleus              
                                      +--+                                            
                                         |  +----------- Scabies_pilata               
                                         +--+                                         
                                            |  +-------- Scabies_humilis              
                                            +--+                                      
                                               |  +----- Scabies_anceps               
                                               +--+                                   
                                                  |  +-- Scabies_phaselus             
                                                  +--+                                
                                                     +-- Scabies_scobinatus

Cladogram Figure S3

  +-------------------------------------------- Indopseudodon_kayinensis     
  |                                                                          
  |                             +-------------- Parreysia_keralaensis        
  |                             |                                            
  |  +--------------------------+  +----------- Parreysia_marcens            
  |  |                          |  |                                         
  |  |                          +--+  +-------- Parreysia_plagiosoma         
  |  |                             |  |                                      
  |  |                             +--+  +----- Parreysia_corrugata          
  |  |                                |  |                                   
  |  |                                +--+----- Parreysia_nagpoorensis       
  |  |                                   |                                   
--+  |                                   |  +-- Parreysia_rakhinensis        
  |  |                                   +--+                                
  |  |                                      +-- Parreysia_imphalensis        
  |  |                                                                       
  |  |  +-------------------------------------- Assamnaia_involuta           
  |  |  |                                                                    
  |  |  |                             +-------- Leoparreysia_whitteni        
  |  |  |                             |                                      
  |  |  |              +--------------+-------- Leoparreysia_burmana         
  |  |  |              |              |                                      
  |  |  |              |              |  +----- Leoparreysia_olivacea        
  |  |  |              |              +--+                                   
  |  |  |              |                 |  +-- Leoparreysia_canefrii        
  +--+  |              |                 +--+                                
     |  |              |                    +-- Leoparreysia_tavoyensis      
     |  |              |                                                     
     |  +--------------+     +----------------- Balwantia_soleniformis       
     |  |              |  +--+                                               
     |  |              |  |  |  +-------------- Trapezidens_mogaungensis     
     |  |              |  |  +--+                                            
     |  |              |  |     |  +----------- Trapezidens_exolescens       
     |  |              |  |     +--+                                         
     |  |              |  |        |  +-------- Trapezidens_scutum           
     |  |              |  |        +--+                                      
     |  |              +--+           |  +----- Trapezidens_yeti             
     |  |                 |           +--+                                   
     |  |                 |              |  +-- Trapezidens_angustior        
     |  |                 |              +--+                                
     |  |                 |                 +-- Trapezidens_dolichorhynchus  
     |  |                 |                                                  
     |  |                 |  +----------------- Lamellidens_nongyangensis    
     |  |                 |  |                                               
     |  |                 +--+  +-------------- Lamellidens_corrianus        
     |  |                    |  |                                            
     +--+                    +--+  +----------- Lamellidens_chindwinensis    
        |                       |  |                                         
        |                       |  +----------- Lamellidens_savadiensis      
        |                       +--+                                         
        |                          |  +-------- Lamellidens_marginalis       
        |                          |  |                                      
        |                          |  +-------- Lamellidens_ferrugineus      
        |                          +--+                                      
        |                             |  +----- Lamellidens_generosus        
        |                             |  |                                   
        |                             |  +----- Lamellidens_mainwaringi      
        |                             +--+                                   
        |                                +----- Lamellidens_phenchooganjensis
        |                                |                                   
        |                                |  +-- Lamellidens_jenkinsianus     
        |                                +--+                                
        |                                   +-- Lamellidens_lamellatus       
        |                                                                    
        |                             +-------- Nitia_teretiuscula           
        |  +--------------------------+                                      
        |  |                          |  +----- Coelatura_aegyptiaca         
        |  |                          +--+                                   
        |  |                             |  +-- Nyassunio_nyassaensis        
        |  |                             +--+                                
        |  |                                +-- Coelatura_kunenensis         
        |  |                                                                 
        |  |                 +----------------- Pseudoscabies_scobina        
        |  |                 |                                               
        |  |                 |              +-- Indonaia_cylindrica          
        |  |  +--------------+  +-----------+                                
        +--+  |              |  |           +-- Indonaia_gratiosa            
           |  |              |  |                                            
           |  |              +--+  +----------- Indonaia_corbis              
           |  |                 |  |                                         
           |  |                 |  |  +-------- Indonaia_andersoniana        
           |  |                 +--+  |                                      
           |  |                    |  |  +----- Indonaia_subclathrata        
           |  |                    |  +--+                                   
           |  |                    +--+  |  +-- Indonaia_caerulea            
           |  |                       |  +--+                                
           |  |                       |     +-- Indonaia_manipurensis        
           +--+                       |                                      
              |                       |  +----- Indonaia_shurtleffiana       
              |                       +--+                                   
              |                          |  +-- Indonaia_bonneaudii          
              |                          +--+                                
              |                             +-- Indonaia_rugosa              
              |                                                              
              |  +----------------------------- Indochinella_pugio           
              |  |                                                           
              |  |                          +-- Radiatula_theobaldi          
              |  |                 +--------+                                
              |  |                 |        +-- Radiatula_crispisulcata      
              |  |                 |                                         
              +--+  +--------------+     +----- Radiatula_mouhoti            
                 |  |              |  +--+                                   
                 |  |              |  |  |  +-- Radiatula_chindwinensis      
                 |  |              |  |  +--+                                
                 |  |              +--+     +-- Radiatula_myitthanensis      
                 |  |                 |                                      
                 |  |                 |  +----- Radiatula_chaudhurii         
                 +--+                 +--+                                   
                    |                    |  +-- Radiatula_myitkyinae         
                    |                    +--+                                
                    |                       +-- Radiatula_kachinensis        
                    |                                                        
                    |  +----------------------- Scabiellus_songkramensis     
                    |  |                                                     
                    +--+  +-------------------- Harmandia_somboriensis       
                       |  |                                                  
                       +--+  +----------------- Unionetta_fabagina           
                          |  |                                               
                          +--+  +-------------- Scabies_crispata             
                             |  |                                            
                             +--+  +----------- Scabies_mandarinus           
                                |  |                                         
                                +--+  +-------- Scabies_humilis              
                                   |  |                                      
                                   |  +-------- Scabies_nucleus              
                                   +--+                                      
                                      +-------- Scabies_pilata               
                                      |                                      
                                      |  +----- Scabies_anceps               
                                      +--+                                   
                                         |  +-- Scabies_phaselus             
                                         +--+                                
                                            +-- Scabies_scobinatus